Paraneseuthia tanimbariana, Jałoszyński, 2019

Jałoszyński, Paweł, 2019, Four new Australo-Oriental species of Paraneseuthia (Coleoptera: Staphylinidae: Scydmaeninae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (2), pp. 491-498 : 495

Paraneseuthia tanimbariana sp. nov.

( Figs 3 , 9–10 )

Type material. HOLOTYPE: ♂, INDONESIA: MALUKU: TANIMBAR ISLANDS: two labels: ʻINDO:MALUC / Tanimbar: Yamdena Isls . / 22 km N Saumlaki, litter / Agosti 11.9.91 (9)’ [white, printed], ʻ PARANESEUTHIA / tanimbariana / det. P. Jałoszyński, 2019 / HOLOTYPUS’ [red, printed] ( MHNG). PARATYPE: ♀, same data as for holotype ( MHNG).

Diagnosis. Body moderately convex; in both sexes frons between eyes with pair of distinct pits with diffuse margins separated by space subequal to their diameters; pronotum with narrow transverse groove connecting lateral antebasal pits; punctures on elytra superficial, inconspicuous; aedeagus with median lobe conspicuously slender, broadest in sub-basal region, with pair of long and slender lateral apical projections, each with its apical portion strongly curved laterad and with hyaline, sharply delimited area; distomedian portion of median lobe narrow and strongly elongate, in lateral view visible as curved projection with broadened apex; endophallus lacking dark sclerites; parameres slender, each with one apical and one subapical seta, one long seta is also inserted near middle of each lateral apical projection.

Description. Male. Body ( Fig. 3 ) suboval, moderately convex; BL 0.78 mm; cuticle glossy, brown, appendages indistinctly lighter, setae light brown.

Head broadest at eyes, HL 0.10 mm, HW 0.18 mm; vertex and frons confluent; frons between eyes with pair of distinct pits with diffuse margins, each pit closer to the other one than to eye, distance between pits subequal to their diameters; vertex transverse, flattened. Supraantennal tubercles weakly elevated. Punctures on frons and vertex fine, inconspicuous, cuticle glossy; setae very short, suberect, sparse. Eyes large, only slightly bean-shaped, strongly convex. Antennae short, AnL 0.30 mm; antennomeres 1–2 each elongate, 3–5 each about as long as broad, 6–7 each slightly transverse, 8 strongly transverse, almost ring-like, 9 and 10 each distinctly, but not strongly transverse, 11 much broader than 10, indistinctly longer than broad.

Pronotum in dorsal view inversely subtrapezoidal, rounded, broadest slightly behind middle, PL 0.23 mm, PW 0.28 mm; anterior margin broadly rounded and demarcated from lateral margins by blunt and obtuse-angled but well-marked anterior corners; lateral margins weakly and almost evenly rounded, in posterior third weakly convergent posterad; posterior pronotal corners obtuse-angled and blunt; posterior margin in front of scutellar base straight, at each side indistinctly concave and slightly bent anterad; lateral antebasal impressions distinct, subtriangular, each accompanied by small and shallow but distinct pit closer to lateral margin than to middle, pits connected by narrow but distinct transverse groove. Punctures on pronotal disc very fine, inconspicuous; setae moderately long, sparse, suberect.

Elytra together oval, broadest slightly in front of middle, EL 0.45 mm, EW 0.33 mm, EI 1.38; humeral calli small, elongate, elytral apices separately, broadly rounded. Punctures much larger than those on pronotum but superficial and with diffuse margins, inconspicuous; setae sparse, suberect, slightly longer than those on pronotum.

Legs moderately long and slender; unmodified, protibiae nearly straight.

Aedeagus ( Figs 9–10 ) slender, approximately drop-shaped, AeL 0.23 mm, median lobe in ventral view broadest in sub-basal region, strongly narrowing distad, with pair of long and slender lateral apical projections, each with distal region strongly curved laterad and with well-defined hyaline apical portion, each projection with long seta near middle of external margin; median apical portion of median lobe forms elongate, dorso-ventrally curved projection with thickened apex; endophallus lacking darkly sclerotized components; parameres slender, in lateral view each bent ventrad at an obtuse angle, each with one apical and one subapical seta, the latter far from apex.

Female externally very similar to male, slightly larger and with stouter elytra. BL 0.84 mm; HL 0.11 mm, HW 0.18 mm, AnL 0.30 mm; PL 0.25 mm, PW 0.30 mm; EL 0.48 mm, PW 0.38 mm, EI 1.27.

Etymology. After the Tanimbar Islands; adjective.

Distribution. Indonesia, Maluku Province, Tanimbar Islands (= Timur Laut), southern region of Yamdena Island.

Remarks. Adults of this species are unremarkable and identification is possible only by the conspicuously slender and complicated aedeagus with a bifurcate apex and a pair of thick lateral setae in the subapical region. No otherAsian species has a similar aedeagus. Similarities in the male genitalic structures can be found only in some congeners that occur in Australia. However, the Australian species have much more slender and more flattened body than the only known male of P. tanimbariana .

Figures

Figs 1–4. Dorsal habitus of male holotypes. 1 – Paraneseuthia luzonica sp. nov.; 2 – P. kaibesariana sp. nov.; 3 – P. tanimbariana sp. nov.; 4 – P. morobensis sp. nov.

Figs 5–12. Aedeagus in ventral (5, 7, 9, 11) and lateral (6, 8, 10, 12) views. 5–6 – Paraneseuthia luzonica sp. nov.; 7–8 – P. kaibesariana sp. nov.; 9–10 – P. tanimbariana sp. nov.; 11–12 – P. morobensis sp. nov.