Indratilla Lelej, 1993

Lelej, Arkady S. & Williams, Kevin A., 2023, Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae), Zootaxa 5231 (2), pp. 101-140 : 115

10. Indratilla Lelej, 1993

( Figs 28–29 )

Indratilla Lelej, 1993: 233 , ♁ non ♀; 2005: 42, 179, ♁, ♀; Lelej & Brothers 2008: 28; Brothers & Lelej 2017: 95, ♁, ♀; Pagliano et al. 2020: 174.

Type species. Indratilla gynandromorpha Lelej, 1993 , ♁, by original designation.

Diagnosis. MALE. Apterous. Head 1.3 × wider than pronotum. Mandible with well developed preapical tooth on inner border and obtuse basal lobe outside beneath. Ocelli small, ratio POD:OOD 0.6. Clypeus weakly modified, with weakly elevated medial part and slightly emarginated anterior margin. Genal carina developed and ends anterad by obtuse tubercle, ratio of eye height to gena height (head in lateral aspect) 0.75 ×. Eyes weakly sinuate within, not reniform. Preoccipital carina dorsally well developed. Antenna unmodified, two apical flagellomeres dorsoventally compressed, ratio F1–3 2.4:2.4:2.3. Mesosoma elongated with rounded humeral angle, 2.5 × longer than minimal width. Tegula large, not touching propodeal spiracle. Scuto-scutellar suture visible, posterolateral scutal angle produced into small tooth, metanotum well visible, mesopleuron weakly convex. Metacoxa mesally carinate. T1 with well developed dorsal and anterior surfaces. T2 with slightly flattened disc and lateral felt lines. S1 with well-developed carina; S2 with basal weak medial, longitudinal carina, without any lateral felt lines. T7 slightly convex. S8 (hypopygium) semitransparent apically, with weakly rounded posterior margin. Genitalia with curved down gonostylus and short broad volsella, penial valves symmetrical, gonostylus ventro-basally with long setae, volsella with dense setae. FEMALE. Head width slightly larger than pronotal width. Eyes moderately large. Genal carina developed and joint with hypostomal carina, latter widened at this point. Prementun not tuberculate. F1 1.5 × F2. Mandible acuminates with two very weak preapical teeth, without inner subbasal tooth. Mesosoma elongated, 1.5 × longer than width between propodeal spiracles. Scutellar scale distinct. T1 short with distinct dorsal surface. Disc of T2 flattened even weakly invaginated, with lateral felt line. T6 with pygidial plate glabrous shiny, widened posterad and weakly carinated laterally in apical half and microsculptured in basal half.

Diversity and distribution. Only type species, known from both sexes, from the dry zone of Sri Lanka ( Lelej 2005) and South India (Tamil Nadu, new record) is recognized .

Remarks. The female Indratilla ceylonica Lelej, 1993 , which was originally assigned to this genus was subsequently transferred to the trogaspidiine genus Radoszkowskitilla Lelej, 2005 by Lelej (2005).

Figures

FIGURES 26–34. 26. Guineomutilla mateui Suárez, 1977, ♀, dorsal habitus; 27. Gynandrotilla ferruginea Arnold, 1946, ♁, dorsal habitus; 28–29. Indratilla gynandromorpha Lelej, 1993: 28. ♀, dorsal habitus, 29. ♁, holotype, dorsal habitus; 30–34. Karunaratnea palatupanae Lelej, 2005: 30–31. Paratype, ♀ (30. Dorsal habitus, 31. Lateral habitus); 32–34. Holotype, ♁: 32. Dorsal habitus, 33. Head, face view, 34. Genitalia, ventral view. (Photographs by D. Brothers (26, 27), V. Loktionov (28, 29), and A. Lelej (30–34)).