Aceclostria Vuillot, 1893

Laurent, Ryan A. St & Kawahara, Akito Y., 2019, Reclassification of the Sack-bearer Moths (Lepidoptera, Mimallonoidea, Mimallonidae), ZooKeys 815, pp. 1-114 : 30-32

Aceclostria Vuillot, 1893 Figs 33, 69, 146, 147

Type species.

Aceclostria mus Vuillot, 1893.

Diagnosis.

Silvery-gray to gray brown ground color, sinuate, diffuse postmedial lines, falcate forewings, and broad darker gray to dark gray brown discal marks that vary from fully scaled to fully hyaline within species (often from the same series); maculation overall is dark and diffuse giving the wings a soiled appearance.

Apomorphy.

Asymmetric, complex genitalia with phallus situated on left side (when viewed ventrally) of vinculum (Fig. 33a) with single vinculum tusk reaching out above phallus.

Remarks.

St Laurent et al. (2018a) did not include this genus in their phylogeny and instead placed this genus in Cicinninae: Cicinnini based on morphology, namely the presence of all apomorphies of Cicinninae and Cicinnini in what was at the time the single Aceclostria species, A. mus . Our ongoing molecular work, which includes broadened taxon coverage, and Aceclostria , fully supports this placement (St Laurent et al. in prep.). The morphological phylogenetic analyses carried out here support this placement as well, with Aceclostria always recovered within Cicinnini , sister to (unconstrained ML and parsimony analysis, Suppl. materials 5 and 7 respectively) or nested within (constrained ML analysis, Fig. 1) the broader, so far poorly resolved Cicinnus sensu lato clade.

See annotations in Section 4 for information pertaining to the novel inclusion of A. cordubensis comb. n. and A. nigrescens comb. n. et stat. rev. in Aceclostria . Aceclostria was long considered a monotypic genus, but morphological examinations, including the genitalia of A. cordubensis , conclusively support these additional poorly known taxa in this genus. This genus is apparently closely associated with Anacardiaceae (see Annotations).

The Cicinnus s.l. clade is discussed in more detail below as it is the most poorly resolved set of taxa in the family.