Can a rainbow consist of a single colour? A new comprehensive generic arrangement of the ‘ Iris sensu latissimo’ clade (Iridaceae), congruent with morphology and molecular data Author Crespo, Manuel B. dCARN & CIBIO (Instituto de la Biodiversidad), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain; e-mail: crespo @ ua. es crespo@ua.es Author Martínez-Azorín, Mario dCARN & CIBIO (Instituto de la Biodiversidad), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain; e-mail: crespo @ ua. es crespo@ua.es Author Mavrodiev, Evgeny V. dCARN & CIBIO (Instituto de la Biodiversidad), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain; e-mail: crespo @ ua. es & Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA. * author for correspondence. & dCARN & CIBIO (Instituto de la Biodiversidad), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain; e-mail: crespo @ ua. es crespo@ua.es text Phytotaxa 2015 2015-10-28 232 1 448 450 http://dx.doi.org/10.11646/phytotaxa.232.1.1 journal article 10.11646/phytotaxa.232.1.1 1179-3163 13632077 25. Siphonostylis Wern.Schulze, Oesterr. Bot. Z. 112: 331 (1965) [≡ Iris subg. Siphonostylis (Wern.Schulze) C.A.Wilson , Taxon 60: 33 (2011) ]. Type :— S. unguicularis (Poir.) Wern.Schulze [≡ Iris unguicularis Poir., Voy. Barbarie 2: 86 (1789) ] Description :—Rhizomatous herbs, with persistent aerial structures. Rhizome stout, branched, clothed with dark remains of leaves; roots fibrous, wiry, somewhat fleshy. Stems hypogeal, inconspicuous, simple, solid, rounded in cross section, elongating in fruit. Leaves isobilateral, equitant, ensiform to linear, with prominent ribs and margins, not fetid. Flowers 1–2, terminal, subsessile. Spathe valves 2, scarious or herbaceous, ± keeled. Perigone in 2 rows, differing in size and shape, fused in a very long, scapiform tube; falls erect-patent, obovate-cuneate to subpanduriform, reflexed in the upper part, gradually tapering into a short claw, smooth, with nectar drops at the base; standards erect to erect-patent, oblong to oblong-spatulate, about equalling the length of falls, narrowed into a long haft. Stamen filaments fused into a tube, adnate to the fall bases. Ovary trilocular, with axile placentation. Style filiform, with 3 petaloid branches, bearing yellow glands on the upper surface, about half longer than falls, each one concealing a stamen; crests long, straight, triangular-lanceolate; stigma triangular to linguiform, with crenate margin. Capsule oblong to ellipsoid, subtrigonous, dehiscing from apex to about middle, not hidden into spathes, shortly beaked; pericarp coriaceous. Seeds numerous, globose to angulose, without aril; testa surface wrinkled, covered with glistening sessile glands. p = 8 or 10. Figs. 2I , 3A , 6T , 7A . 66 • Phytotaxa 232 (1) © 2015 Magnolia Press CRESPO ET AL. It includes 4 species, occurring in the central and eastern parts of the Mediterranean basin, to the southern shore of the Black Sea ( Fig. 8A ). Needed new combinations :— Siphonostylis carica (Wern.Schulze) M.B.Crespo, Mart. -Azorín & Mavrodiev, comb. nov. Basionym: Siphonostylis cretensis subsp. carica Wern.Schulze, Oesterr. Bot. Z. 112: 338 (1965) [ Iris unguicularis subsp. carica (Wern.Schulze) A.P.Davis & Jury, Bot. J. Linn. Soc. 103: 295 (1990) ]. Siphonostylis carica var. angustifolia (Boiss. & Heldr.) M.B.Crespo, Mart. -Azorín & Mavrodiev, comb. nov. Basionym: Iris stylosa var. angustifolia Boiss. & Heldr., Diagn. Pl. Orient . 1(13): 15 (1853) [ Iris unguicularis var. angustifolia (Boiss. & Heldr.) A.P.Davis & Jury, Bot. J. Linn. Soc. 103: 296 (1990) ]. Siphonostylis carica var. syriaca (Wern.Schulze) M.B.Crespo, Mart. -Azorín & Mavrodiev, comb. nov. Basionym: Siphonostylis cretensis subsp. syriaca Wern.Schulze, Oesterr. Bot. Z. 112: 339 (1965) [ Iris unguicularis var. syriaca (Wern.Schulze) A.P.Davis & Jury, Bot. J. Linn. Soc. 103: 297 (1990) ]. Typification of other generic names Neubeckia Alef., Bot. Zeitung (Berlin) 21: 297 (1863) . Type ( lectotype here designated): — N. scorpioides (Desf.) Alef., Bot. Zeitung (Berlin) 21: 297 (1863) [≡ Iris scorpioides Desf., Fl. Atlant. 1: 40, t. 6 (1798). = Juno planifolia (Mill.) Asch., Bot. Zeitung (Berlin) 22: 114 (1864) ]. Note : — In the protologue, Alefeld (1863) included eight species apparently only sharing long perigone tubes, but strongly differing in many vegetative and floral traits. According to recent classifications, they fall into several subgenera, sections and series of Iris s.l. ( Mathew 1989b ), or into seven of the genera here accepted (namely, Juno , Siphonostylis , Phaeiris , Lophiris , Iridodictyum , Cryptobasis and Chamaeiris ). However, Alefeld included two species of Juno , J. planifolia ( Miller 1768 : without pagination) Ascherson (1864: 114) and J. caucasica ( Hoffmann 1808: 40 ) Kotschy in Ascherson (1864: 114), and he explicitly mentioned that the fruit characteristics in the generic description were taken only from J. planifolia [= Neubeckia scorpioides ( Desfontaines 1798: 40 ) Alefeld 1863: 297 ]. Therefore, to avoid further threats to any of those genera, we typify here Neubeckia on the basis of J. planifolia , a solution which perhaps fits more properly Alefeld’s original concept of that genus. Identification key to the genera 1. - 2. - 3. - 4. - 5. - 6. - 7. -