<document id="720C906793F8C0565CB0BE9BB663B7D6" ID-DOI="10.1080/00222933.2025.2498047" ID-ISSN="1464-5262" ID-Zenodo-Dep="17006654" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1756538573921" checkinUser="plazi" docAuthor="Shafaie, Sepideh, Pekár, Stano &amp; Ortiz, David" docDate="2025" docId="1533B37EFFB6E15BD867FBC880A3FB67" docLanguage="en" docName="JNatHist.59.25-28.1825-1875.pdf" docOrigin="Journal of Natural History 59 (25 - 28)" docSource="https://doi.org/10.1080/00222933.2025.2498047" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Zodarion sanchezi Shafaie and Pekar 2025, sp. n." docType="treatment" docVersion="2" lastPageNumber="1842" masterDocId="E90ACB06FFA6E14AD91EFFEE822AFFAC" masterDocTitle="Zodarion spiders (Araneae: Zodariidae): morphology, molecular characterisation and reassessment of the elegans, italicum, and pusio groups from Iberia and the Canary Islands" masterLastPageNumber="1875" masterPageNumber="1825" pageNumber="1841" updateTime="1756543730871" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AC703BFB2A024C4EF052F577DA2371E0">Zodarion spiders (Araneae: Zodariidae): morphology, molecular characterisation and reassessment of the elegans, italicum, and pusio groups from Iberia and the Canary Islands</mods:title>
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<mods:namePart id="0ED9A54C6E56F2D281255ED138AA15DA">Shafaie, Sepideh</mods:namePart>
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<mods:namePart id="7C025A19FB641D1E228D81D8F9789397">Pekár, Stano</mods:namePart>
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<mods:namePart id="93BB2CA04DA613D846EDE7D867C4AD2B">Ortiz, David</mods:namePart>
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<mods:title id="C22A140660C5D785F9D6C096C52DC26D">Journal of Natural History</mods:title>
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<mods:part id="14C95E85A5381A55392F62DC7258C9D7">
<mods:date id="B7A5C1464015EFAEAFF69C6F57FB9B33">2025</mods:date>
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<mods:number id="3FF42DA5B81E8BE871546E3E185457F3">2025-06-05</mods:number>
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<mods:number id="3AF38F00C2C0529AA29437403637ED14">59</mods:number>
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<mods:number id="13792B53D0439852B03E4E4AF484A30E">25 - 28</mods:number>
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<mods:start id="C4736F28C9A3A3AAE3B852680BD8BE11">1825</mods:start>
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<emphasis id="AFEEDE7AFFB6E15AD867FBC88181FBED" bold="true" box="[377,939,1062,1089]" pageId="16" pageNumber="1841">
<taxonomicName id="5A9A79EBFFB6E15AD867FBC88171FBED" authority="Shafaie and Pekar" authorityName="Shafaie and Pekar" authorityYear="2025" box="[377,859,1062,1089]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1841" phylum="Arthropoda" rank="species" species="sanchezi" status="sp. nov.">
<emphasis id="AFEEDE7AFFB6E15AD867FBC8804EFBED" bold="true" box="[377,612,1062,1089]" italics="true" pageId="16" pageNumber="1841">Zodarion sanchezi</emphasis>
Shafaie and Pekár
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<taxonomicNameLabel id="B4DD6301FFB6E15ADA7DFBC88181FBED" box="[867,939,1062,1089]" pageId="16" pageNumber="1841" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D58051E3FFB6E15ADB2DFBAB80DAFBCC" box="[563,752,1093,1120]" pageId="16" pageNumber="1841" type="description">
<paragraph id="9D250268FFB6E15ADB2DFBAB80DAFBCC" blockId="16.[377,939,1062,1120]" box="[563,752,1093,1120]" pageId="16" pageNumber="1841">
(
<figureCitation id="05A11EEDFFB6E15ADB25FBAB8094FBCC" box="[571,702,1093,1120]" captionStart="Figure 12" captionStartId="26.[160,225,679,702]" captionTargetBox="[165,1150,144,645]" captionTargetId="figure-196@26.[163,1153,142,648]" captionTargetPageId="26" captionText="Figure 12. Habitus of Z. sanchezi Shafaie and Pekár sp. n. A–B, holotype, male (NMP). A, dorsal view. B, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17006695" httpUri="https://zenodo.org/record/17006695/files/figure.png" pageId="16" pageNumber="1841">Figures 12</figureCitation>
, 
<figureCitation id="05A11EEDFFB6E15ADBD4FBAB80C2FBCC" box="[714,744,1093,1120]" captionStart="Figure 23" captionStartId="38.[160,225,1317,1340]" captionTargetBox="[165,1150,144,1283]" captionTargetId="figure-50@38.[163,1153,142,1286]" captionTargetPageId="38" captionText="Figure 23. Male palp of Z. sanchezi Shafaie and Pekár sp. n., holotype (NMP). A, D, prolateral view.B, E, ventral view. C, F, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; Tm = mesal tubercle of the tibial apophysis; Tr = retrolateral tubercle of the tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.17006731" httpUri="https://zenodo.org/record/17006731/files/figure.png" pageId="16" pageNumber="1841">23</figureCitation>
)
</paragraph>
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<paragraph id="9D250268FFB6E15AD9DEFB958128FB38" blockId="16.[192,770,1147,1172]" box="[192,770,1147,1172]" pageId="16" pageNumber="1841">
<treatmentCitationGroup id="BD8A2546FFB6E15AD9DEFB958128FB38" box="[192,770,1147,1172]" pageId="16" pageNumber="1841">
<taxonomicName id="5A9A79EBFFB6E15AD9DEFB95834BFB38" box="[192,353,1147,1172]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1841" phylum="Arthropoda" rank="species" species="undefined-3">
<emphasis id="AFEEDE7AFFB6E15AD9DEFB958309FB38" box="[192,291,1147,1172]" italics="true" pageId="16" pageNumber="1841">Zodarion</emphasis>
sp. 3
</taxonomicName>
. 
<treatmentCitation id="1C3B2479FFB6E15AD875FB95807CFB38" author="Ortiz D &amp; Pekar S &amp; Bilat J &amp; Shafaie S &amp; Alvarez N &amp; Gauthier J" box="[363,598,1147,1172]" page="62" pageId="16" pageNumber="1841" year="2023">
<bibRefCitation id="F90B7F99FFB6E15AD875FB95807CFB38" author="Ortiz D &amp; Pekar S &amp; Bilat J &amp; Shafaie S &amp; Alvarez N &amp; Gauthier J" box="[363,598,1147,1172]" pageId="16" pageNumber="1841" pagination="20230797" refId="ref16185" refString="Ortiz D, Pekar S, Bilat J, Shafaie S, Alvarez N, Gauthier J. 2023 b. Dynamic evolution of size and colour in the highly specialized Zodarion ant-eating spiders. Proc R Soc B. 290: 20230797. doi: 10.1098 / rspb. 2023.0797." type="journal article" year="2023">
Ortiz 
<emphasis id="AFEEDE7AFFB6E15AD8B6FB9583F2FB38" box="[424,472,1147,1172]" italics="true" pageId="16" pageNumber="1841">et al</emphasis>
. 2023b: 62
</bibRefCitation>
</treatmentCitation>
; figs 1, 3, 4 (
<specimenCount id="8B9CC9E1FFB6E15ADBFFFB9580DFFB3E" box="[737,757,1147,1170]" count="1" pageId="16" pageNumber="1841" type="male">♂</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D58051E3FFB6E15AD9BEFB2E8676FA2B" pageId="16" pageNumber="1841" type="materials_examined">
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<emphasis id="AFEEDE7AFFB6E15AD9BEFB2E836EFB76" bold="true" box="[160,324,1216,1242]" italics="true" pageId="16" pageNumber="1841">Type material</emphasis>
</paragraph>
<paragraph id="9D250268FFB6E15AD9BEFB0D839EFAB3" blockId="16.[160,1155,1216,1311]" pageId="16" pageNumber="1841">
<materialsCitation id="2DF20835FFB6E15AD9BEFB0D839EFAB3" collectingDate="2012-05-28" collectionCode="NMP" collectorName="S. Pekar" country="Spain" county="Nijar" location="Almeria" municipality="Punta de Las Majadas" pageId="16" pageNumber="1841" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="4221BCCAFFB6E15AD9BEFB0D832DFB50" box="[160,263,1251,1276]" pageId="16" pageNumber="1841" type="holotype">Holotype</typeStatus>
(
<specimenCount id="8B9CC9E1FFB6E15AD809FB0A8301FB57" box="[279,299,1252,1275]" count="1" pageId="16" pageNumber="1841" type="male">♂</specimenCount>
) and 
<typeStatus id="4221BCCAFFB6E15AD873FB0D83F1FB50" box="[365,475,1251,1276]" pageId="16" pageNumber="1841" type="paratype">paratypes</typeStatus>
(
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): 
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<collectingCountry id="E58D42F8FFB6E15ADB0AFB0D804AFB50" box="[532,608,1251,1276]" name="Spain" pageId="16" pageNumber="1841">SPAIN</collectingCountry>
: 
<location id="984554B3FFB6E15ADB73FB0D80E7FB50" LSID="urn:lsid:plazi:treatment:1533B37EFFB6E15BD867FBC880A3FB67:984554B3FFB6E15ADB73FB0D80E7FB50" box="[621,717,1251,1276]" country="Spain" county="Nijar" municipality="Punta de Las Majadas" name="Almeria" pageId="16" pageNumber="1841">Almería</location>
, 
<collectingMunicipality id="7D419812FFB6E15ADBC7FB0D81CBFB50" box="[729,993,1251,1276]" pageId="16" pageNumber="1841">Punta de Las Majadas</collectingMunicipality>
</emphasis>
, 
<collectingCounty id="74447AE4FFB6E15ADAF1FB0D860FFB50" box="[1007,1061,1251,1276]" pageId="16" pageNumber="1841">Níjar</collectingCounty>
; 
<date id="E92424A8FFB6E15ADD2FFB0D82F3FAB3" pageId="16" pageNumber="1841" value="2012-05-28">
<collectingDate id="F960DD40FFB6E15ADD2FFB0D82F3FAB3" pageId="16" pageNumber="1841" value="2012-05-28">28 May 2012</collectingDate>
</date>
; 
<collectorName id="306F67BEFFB6E15AD9FAFAE88316FAB3" box="[228,316,1286,1311]" pageId="16" pageNumber="1841">S. Pekár</collectorName>
leg.; 
<collectionCode id="FB8B9AADFFB6E15AD866FAE8839AFAB3" box="[376,432,1286,1311]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="16" pageNumber="1841" type="">NMP</collectionCode>
.
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</paragraph>
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<emphasis id="AFEEDE7AFFB6E15AD9BEFAA583E0FAC9" bold="true" box="[160,458,1355,1381]" italics="true" pageId="16" pageNumber="1841">Other material examined</emphasis>
</paragraph>
<paragraph id="9D250268FFB6E15AD9BEFA808676FA2B" blockId="16.[160,1116,1355,1415]" box="[160,1116,1390,1415]" pageId="16" pageNumber="1841">
<materialsCitation id="2DF20835FFB6E15AD9BEFA8083B4FA2B" box="[160,414,1390,1415]" country="Spain" location="Almeria" pageId="16" pageNumber="1841" specimenCount="1" stateProvince="Almeria">
<emphasis id="AFEEDE7AFFB6E15AD9BEFA80837FFA2B" bold="true" box="[160,341,1390,1415]" pageId="16" pageNumber="1841">
<collectingCountry id="E58D42F8FFB6E15AD9BEFA8082C6FA2B" box="[160,236,1390,1415]" name="Spain" pageId="16" pageNumber="1841">SPAIN</collectingCountry>
: 
<collectingRegion id="5F5ECC8AFFB6E15AD9E6FA80837FFA2B" box="[248,341,1390,1415]" pageId="16" pageNumber="1841">Almería</collectingRegion>
</emphasis>
, 
<specimenCount id="8B9CC9E1FFB6E15AD87CFA8083B4FA2B" box="[354,414,1390,1415]" count="3" pageId="16" pageNumber="1841" type="generic">3 juv</specimenCount>
</materialsCitation>
<materialsCitation id="2DF20835FFB6E15AD880FA8081B1FA2B" box="[414,923,1390,1415]" collectingDate="2012-05-28" collectorName="Nijar" country="Spain" location="Punta de Las Majadas" pageId="16" pageNumber="1841" specimenCount="1" stateProvince="Almeria">
.; 
<location id="984554B3FFB6E15AD8B2FA808089FA2B" LSID="urn:lsid:plazi:treatment:1533B37EFFB6E15BD867FBC880A3FB67:984554B3FFB6E15AD8B2FA808089FA2B" box="[428,675,1390,1415]" country="Spain" name="Punta de Las Majadas" pageId="16" pageNumber="1841" stateProvince="Almeria">Punta de Las Majadas</location>
, 
<collectorName id="306F67BEFFB6E15ADBB3FA8080C9FA2B" box="[685,739,1390,1415]" pageId="16" pageNumber="1841">Níjar</collectorName>
; 
<date id="E92424A8FFB6E15ADBF0FA8081AAFA2B" box="[750,896,1390,1415]" pageId="16" pageNumber="1841" value="2012-05-28">
<collectingDate id="F960DD40FFB6E15ADBF0FA8081AAFA2B" box="[750,896,1390,1415]" pageId="16" pageNumber="1841" value="2012-05-28">28 May 2012</collectingDate>
</date>
; S
</materialsCitation>
<materialsCitation id="2DF20835FFB6E15ADA84FA80863BFA2B" box="[922,1041,1390,1415]" collectingDate="2012-05-28" collectorName="Pekar" country="Spain" isTailEnumeration="true" location="Punta de Las Majadas" pageId="16" pageNumber="1841" specimenCount="1" stateProvince="Almeria">
. 
<collectorName id="306F67BEFFB6E15ADABBFA8081C9FA2B" box="[933,995,1390,1415]" pageId="16" pageNumber="1841">Pekár</collectorName>
leg
</materialsCitation>
.; 
<collectionCode id="FB8B9AADFFB6E15ADD01FA808672FA2B" box="[1055,1112,1390,1415]" pageId="16" pageNumber="1841">CMU</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="D58051E3FFB6E15AD9BEFA5D8337F9BF" pageId="16" pageNumber="1841" type="etymology">
<paragraph id="9D250268FFB6E15AD9BEFA5D8337FA61" blockId="16.[160,1156,1459,1555]" box="[160,285,1459,1485]" pageId="16" pageNumber="1841">
<emphasis id="AFEEDE7AFFB6E15AD9BEFA5D8337FA61" bold="true" box="[160,285,1459,1485]" italics="true" pageId="16" pageNumber="1841">Etymology</emphasis>
</paragraph>
<paragraph id="9D250268FFB6E15AD9BEFA398337F9BF" blockId="16.[160,1156,1459,1555]" pageId="16" pageNumber="1841">The specific epithet is in honour of Íñigo Sánchez, who provided us with rare Spanish specimens.</paragraph>
</subSubSection>
<subSubSection id="D58051E3FFB6E15BD9BEF9D086A9FE9E" lastPageId="17" lastPageNumber="1842" pageId="16" pageNumber="1841" type="diagnosis">
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<emphasis id="AFEEDE7AFFB6E15AD9BEF9D0833EF9F4" bold="true" box="[160,276,1598,1624]" italics="true" pageId="16" pageNumber="1841">Diagnosis</emphasis>
</paragraph>
<paragraph id="9D250268FFB6E15BD9BEF98C86A9FE9E" blockId="16.[160,1155,1598,1728]" lastBlockId="17.[160,1155,142,306]" lastPageId="17" lastPageNumber="1842" pageId="16" pageNumber="1841">
The male of the new species differs from that of 
<taxonomicName id="5A9A79EBFFB6E15ADBCCF98C8170F9D7" authorityName="Denis" authorityYear="1935" box="[722,858,1634,1659]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1841" phylum="Arthropoda" rank="species" species="diatretum">
<emphasis id="AFEEDE7AFFB6E15ADBCCF98C8170F9D7" box="[722,858,1634,1659]" italics="true" pageId="16" pageNumber="1841">Z. diatretum</emphasis>
</taxonomicName>
by several characters: the tibial apophysis gently curved, medially with a jagged mesal tubercle, terminally ending in a claw-shaped tip (vs strongly curved, medially humped, and terminally ending in a recurved, rounded tip; compare 
<figureCitation id="05A11EEDFFB7E15BDB12FF6080AFFF0B" box="[524,645,142,167]" captionStart="Figure 23" captionStartId="38.[160,225,1317,1340]" captionTargetBox="[165,1150,144,1283]" captionTargetId="figure-50@38.[163,1153,142,1286]" captionTargetPageId="38" captionText="Figure 23. Male palp of Z. sanchezi Shafaie and Pekár sp. n., holotype (NMP). A, D, prolateral view.B, E, ventral view. C, F, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; Tm = mesal tubercle of the tibial apophysis; Tr = retrolateral tubercle of the tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.17006731" httpUri="https://zenodo.org/record/17006731/files/figure.png" pageId="17" pageNumber="1842">Figure 23C</figureCitation>
and 
<figureCitation id="05A11EEDFFB7E15BDBA0FF60811FFF0B" box="[702,821,142,167]" captionStart="Figure 21" captionStartId="36.[160,225,907,930]" captionTargetBox="[165,1150,145,872]" captionTargetId="figure-119@36.[163,1153,142,875]" captionTargetPageId="36" captionText="Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.17006725" httpUri="https://zenodo.org/record/17006725/files/figure.png" pageId="17" pageNumber="1842">Figure 21F</figureCitation>
); the median apophysis more developed in basal part (vs reduced and shaped as a spur; compare 
<figureCitation id="05A11EEDFFB7E15BDA91FF5F862FFF66" box="[911,1029,177,202]" captionStart="Figure 23" captionStartId="38.[160,225,1317,1340]" captionTargetBox="[165,1150,144,1283]" captionTargetId="figure-50@38.[163,1153,142,1286]" captionTargetPageId="38" captionText="Figure 23. Male palp of Z. sanchezi Shafaie and Pekár sp. n., holotype (NMP). A, D, prolateral view.B, E, ventral view. C, F, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; Tm = mesal tubercle of the tibial apophysis; Tr = retrolateral tubercle of the tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.17006731" httpUri="https://zenodo.org/record/17006731/files/figure.png" pageId="17" pageNumber="1842">Figure 23B</figureCitation>
and 
<figureCitation id="05A11EEDFFB7E15BDD22FF5F82E2FF41" captionStart="Figure 21" captionStartId="36.[160,225,907,930]" captionTargetBox="[165,1150,145,872]" captionTargetId="figure-119@36.[163,1153,142,875]" captionTargetPageId="36" captionText="Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.17006725" httpUri="https://zenodo.org/record/17006725/files/figure.png" pageId="17" pageNumber="1842">Figure 21E</figureCitation>
); the embolus with a stout subterminal tooth that slightly diverged from the embolus (vs acutely pointed, with the subterminal tooth converging towards the embolus; compare 
<figureCitation id="05A11EEDFFB7E15BD9C7FEF7837AFE9E" box="[217,336,281,306]" captionStart="Figure 23" captionStartId="38.[160,225,1317,1340]" captionTargetBox="[165,1150,144,1283]" captionTargetId="figure-50@38.[163,1153,142,1286]" captionTargetPageId="38" captionText="Figure 23. Male palp of Z. sanchezi Shafaie and Pekár sp. n., holotype (NMP). A, D, prolateral view.B, E, ventral view. C, F, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; Tm = mesal tubercle of the tibial apophysis; Tr = retrolateral tubercle of the tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.17006731" httpUri="https://zenodo.org/record/17006731/files/figure.png" pageId="17" pageNumber="1842">Figure 23B</figureCitation>
and 
<figureCitation id="05A11EEDFFB7E15BD894FEF7802AFE9E" box="[394,512,281,306]" captionStart="Figure 21" captionStartId="36.[160,225,907,930]" captionTargetBox="[165,1150,145,872]" captionTargetId="figure-119@36.[163,1153,142,875]" captionTargetPageId="36" captionText="Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.17006725" httpUri="https://zenodo.org/record/17006725/files/figure.png" pageId="17" pageNumber="1842">Figure 21E</figureCitation>
); and distinct prosoma pattern (
<figureCitation id="05A11EEDFFB7E15BDA75FEF781CEFE9E" box="[875,996,281,306]" captionStart="Figure 12" captionStartId="26.[160,225,679,702]" captionTargetBox="[165,1150,144,645]" captionTargetId="figure-196@26.[163,1153,142,648]" captionTargetPageId="26" captionText="Figure 12. Habitus of Z. sanchezi Shafaie and Pekár sp. n. A–B, holotype, male (NMP). A, dorsal view. B, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17006695" httpUri="https://zenodo.org/record/17006695/files/figure.png" pageId="17" pageNumber="1842">Figure 12A</figureCitation>
vs 
<figureCitation id="05A11EEDFFB7E15BDD14FEF7865CFE9E" box="[1034,1142,281,306]" captionStart="Figure 4" captionStartId="10.[160,225,1500,1523]" captionTargetBox="[165,1150,145,1466]" captionTargetId="figure-47@10.[163,1153,142,1469]" captionTargetPageId="10" captionText="Figure 4. Habitus of Z. diatretum Denis, 1935. A–B, lectotype, male (MNHN-AR1513). C–D, paralectotype, female (MNHN-AR1513). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17006669" httpUri="https://zenodo.org/record/17006669/files/figure.png" pageId="17" pageNumber="1842">Figure 4A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D58051E3FFB7E15BD9BEFE8C83CBFBF2" pageId="17" pageNumber="1842" type="description">
<paragraph id="9D250268FFB7E15BD9BEFE8C830CFED0" blockId="17.[160,1156,354,900]" box="[160,294,354,380]" pageId="17" pageNumber="1842">
<emphasis id="AFEEDE7AFFB7E15BD9BEFE8C830CFED0" bold="true" box="[160,294,354,380]" italics="true" pageId="17" pageNumber="1842">Description</emphasis>
</paragraph>
<paragraph id="9D250268FFB7E15BD9BEFE6B81C5FC28" blockId="17.[160,1156,354,900]" pageId="17" pageNumber="1842">
<emphasis id="AFEEDE7AFFB7E15BD9BEFE6B8346FE33" bold="true" box="[160,364,389,415]" italics="true" pageId="17" pageNumber="1842">
Male. (
<typeStatus id="4221BCCAFFB7E15BD9E8FE6B8374FE33" box="[246,350,389,415]" pageId="17" pageNumber="1842" type="holotype">holotype</typeStatus>
).
</emphasis>
Total length 3.85, carapace 1.9 long, 1.25 wide. 
<emphasis id="AFEEDE7AFFB7E15BDAAEFE688633FE33" bold="true" box="[944,1049,390,415]" pageId="17" pageNumber="1842">Prosoma</emphasis>
. (
<figureCitation id="05A11EEDFFB7E15BDD2CFE6882CDFE6E" captionStart="Figure 12" captionStartId="26.[160,225,679,702]" captionTargetBox="[165,1150,144,645]" captionTargetId="figure-196@26.[163,1153,142,648]" captionTargetPageId="26" captionText="Figure 12. Habitus of Z. sanchezi Shafaie and Pekár sp. n. A–B, holotype, male (NMP). A, dorsal view. B, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17006695" httpUri="https://zenodo.org/record/17006695/files/figure.png" pageId="17" pageNumber="1842">Figures 12A–B</figureCitation>
) Carapace dark brown with an elongated heart-shaped median band, dorsally followed by a postero-median, smaller area; clypeus dark brown; chelicerae light; mouthparts pale; sternum light grey. 
<emphasis id="AFEEDE7AFFB7E15BDB1EFE00801CFDAB" bold="true" box="[512,566,494,519]" pageId="17" pageNumber="1842">Eyes</emphasis>
. AME 0.12, ALE 0.09, PME 0.08, PLE 0.08, AME–AME 0.05, ALE–ALE 0.7, PME–PME 0.35, PLE–PLE 0.22, ALE–PLE 0.04. 
<emphasis id="AFEEDE7AFFB7E15BDA4BFDFF81E0FD86" bold="true" box="[853,970,529,554]" pageId="17" pageNumber="1842">Abdomen</emphasis>
. (
<figureCitation id="05A11EEDFFB7E15BDAC3FDFF8651FD86" box="[989,1147,529,554]" captionStart="Figure 12" captionStartId="26.[160,225,679,702]" captionTargetBox="[165,1150,144,645]" captionTargetId="figure-196@26.[163,1153,142,648]" captionTargetPageId="26" captionText="Figure 12. Habitus of Z. sanchezi Shafaie and Pekár sp. n. A–B, holotype, male (NMP). A, dorsal view. B, ventral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.17006695" httpUri="https://zenodo.org/record/17006695/files/figure.png" pageId="17" pageNumber="1842">Figures 12A–B</figureCitation>
) Abdomen dorsum uniformly black, venter yellow and brown. Spinnerets pale apically and light brown basally. 
<emphasis id="AFEEDE7AFFB7E15BD89EFDB8839DFDC3" bold="true" box="[384,439,598,623]" pageId="17" pageNumber="1842">Legs</emphasis>
. Leg segments yellow except for dark femur I, femur II distal half and femora III–IV distal tip, and yellow tibiae I–IV. Legs: I 5.75 (1.6, 0.6, 1.3, 1.5, 0.75), II 5.45 (1.45, 0.55, 1.1, 1.45, 0.9), III 5.34 (1.34, 0.6, 1.05, 1.45, 0.9), IV 7.9 (2.15, 0.7, 1.85, 2.1, 1.1). 
<emphasis id="AFEEDE7AFFB7E15BD9BEFD5082FFFD7B" bold="true" box="[160,213,702,727]" pageId="17" pageNumber="1842">Palp</emphasis>
. (
<figureCitation id="05A11EEDFFB7E15BD9F4FD5083A3FD7B" box="[234,393,702,727]" captionStart="Figure 23" captionStartId="38.[160,225,1317,1340]" captionTargetBox="[165,1150,144,1283]" captionTargetId="figure-50@38.[163,1153,142,1286]" captionTargetPageId="38" captionText="Figure 23. Male palp of Z. sanchezi Shafaie and Pekár sp. n., holotype (NMP). A, D, prolateral view.B, E, ventral view. C, F, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; Tm = mesal tubercle of the tibial apophysis; Tr = retrolateral tubercle of the tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.17006731" httpUri="https://zenodo.org/record/17006731/files/figure.png" pageId="17" pageNumber="1842">Figures 23A–F</figureCitation>
) Palp segments entirely yellow. Tibial apophysis elongated, ending in a diamond-shaped arm in ventral view, gradually recurved in retrolateral view, ending in a triangular tip, pointed at 3 o’clock position, medially accompanied by a jagged mesal tubercle, partially covering a triangular retrolateral tubercle. Median apophysis developed in basal part. Embolus oblong and stout, accompanied by a similar subterminal tooth, the two structures slightly converging. Tegulum slightly protruding posteriorly.
</paragraph>
<paragraph id="9D250268FFB7E15BD9BEFC5B82DCFC63" blockId="17.[160,275,949,1009]" box="[160,246,949,975]" pageId="17" pageNumber="1842">
<emphasis id="AFEEDE7AFFB7E15BD9BEFC5B82DCFC63" bold="true" box="[160,246,949,975]" italics="true" pageId="17" pageNumber="1842">Female</emphasis>
</paragraph>
<paragraph id="9D250268FFB7E15BD9BEFC368339FC5D" blockId="17.[160,275,949,1009]" box="[160,275,984,1009]" pageId="17" pageNumber="1842">Unknown.</paragraph>
<paragraph id="9D250268FFB7E15BD9BEFBCC837EFB90" blockId="17.[160,481,1058,1118]" box="[160,340,1058,1084]" pageId="17" pageNumber="1842">
<emphasis id="AFEEDE7AFFB7E15BD9BEFBCC837EFB90" bold="true" box="[160,340,1058,1084]" italics="true" pageId="17" pageNumber="1842">Molecular data</emphasis>
</paragraph>
<paragraph id="9D250268FFB7E15BD9BEFBAB83CBFBF2" blockId="17.[160,481,1058,1118]" box="[160,481,1093,1118]" pageId="17" pageNumber="1842">No molecular data available.</paragraph>
</subSubSection>
<subSubSection id="D58051E3FFB7E15BD9BEFB6180A3FB67" box="[160,649,1167,1227]" pageId="17" pageNumber="1842" type="distribution">
<paragraph id="9D250268FFB7E15BD9BEFB618306FB05" blockId="17.[160,649,1167,1227]" box="[160,300,1167,1193]" pageId="17" pageNumber="1842">
<emphasis id="AFEEDE7AFFB7E15BD9BEFB618306FB05" bold="true" box="[160,300,1167,1193]" italics="true" pageId="17" pageNumber="1842">Distribution</emphasis>
</paragraph>
<paragraph id="9D250268FFB7E15BD9BEFB5C80A3FB67" blockId="17.[160,649,1167,1227]" box="[160,649,1202,1227]" pageId="17" pageNumber="1842">
<materialsCitation id="2DF20835FFB7E15BD9BEFB5C80AFFB67" box="[160,645,1202,1227]" country="Spain" location="Spain" pageId="17" pageNumber="1842" specimenCount="1">
Known only from the 
<typeStatus id="4221BCCAFFB7E15BD889FB5C83E3FB67" box="[407,457,1202,1227]" pageId="17" pageNumber="1842">type</typeStatus>
locality in 
<collectingCountry id="E58D42F8FFB7E15BDB5AFB5C80AFFB67" box="[580,645,1202,1227]" name="Spain" pageId="17" pageNumber="1842">Spain</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>