Amphineurus (Rhamphoneurus) chiloeanus Alexander, 1969

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

3.3.8. Amphineurus (Rhamphoneurus) chiloeanus Alexander, 1969

Amphineurus (Rhamphoneurus) chiloeanus Alexander, 1969: 374 (description, fig. 37: wing, fig. 38: male terminalia).

Material examined.

Holotype: ♂, Chile, Chiloé Is., Dalcahue [42°22′S 73°39′W], 1/ 4-IV-1954, Peña, Holotype 12553 (USNM) [part of specimen pinned and part in slide prepared by C.P. Alexander]. Paratype: 1 ♂, Chile, Chiloé Is., Dalcahue [42°22′S 73°39′W], 1/ 4-IV-1954, Peña (USNM). Additional material: CHILE. 1 [sex unknown], Chile , Chiloé Is. Ancud [41°52′S 73°48′W], 23-1-1952, L. Guzman (USNM) ; 1 ♂, Chiloé Is., Dalcahue [42°22′S 73°39′W], 1/ 4-IV-1954, Peña (USNM) *.

Diagnosis.

The species is distinguished by having m-cu strongly sinuous, straight anal vein and beginning of M1+2 retains sclerotized section. Furthermore, this species has a characteristic sheath of aedeagus with a long basal spur, mesal lobe is very reduced and male tergite IX is remarkable by its rounded posterolateral lobes.

Redescription.

Wing length 7.37 mm, width 2.14 mm. - Coloration: General coloration light brown. Rostrum and palpus dark brown. First segment of antenna light yellow, remaining segments dark yellow. Head dark brown. Eyes black. Thorax light brown, pleura dark yellow. Scutum with three brown stripes. Halter brownish yellow with knob ochreous. Coxae brownish yellow and legs yellowish-brown. - Head (dorsal view Fig. 15B , ventral view 15A): terminal segments of antennae missing, as indicated by asterisks. Rostrum bifurcated with short appendices; first palpal segment longer than terminal segment; scape as thick as pedicel, pedicel constricted at base. - Thorax (lateral view Fig. 15E , dorsal view Fig. 15F ): Division line of laterotergite very sinuous; Anatergite shorter than katatergite. Halter with dilated knob. Wing (Fig. 15D ) with some markings, including faint markings on A1 and Rs; markings highlighting m-cu, r-m, fork of Rs and R2+3+4; distal quarter of wing slightly darkened, with large, strong marking covering R2 and last segment of R1; veins near fork of bM (Fig. 15C ) faded; R2+3+4 longer than R2+3. - Male terminalia (Fig. 15G ): Male tergite IX faintly darkened at posterior margin, with wide, shallow U-shaped median notch; posterolateral lobes rounded. Dorsal branch of gonocoxite poorly developed, shorter than half-length of ventral branch. Lobe of gonostylus with elongate lobule, longer than wide, longer than half-length of lobe of gonostylus, but narrower than stem. Clasper with differently shaped branches: lateral branch straight and long; medial branch curved, wide club-shaped. Mesal lobes atrophied, similar in shape to dorsal branch of gonocoxite. Sheath of aedeagus long and sinuose, darkened almost entire length, with pointed perpendicular spur projecting at end of straight section.

Remarks.

Some specimens of this species were misidentified by C.P. Alexander as A. (R.) nothofagetorum Alexander. This species resembles A. (R.) glabristylatus Alexander but differs mainly in attachment of m-cu, length of cercus, shape of male tergite IX, sheath of aedeagus and mesal lobe.

Figures

Figure 15. Amphineurus (Rhamphoneurus) chiloeanus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX.