Omoedus, Thorell, 1881

Prószyński, Jerzy, 2017, Pragmatic classification of the World's Salticidae (Araneae), Ecologica Montenegrina 12, pp. 1-133 : 75-76

Omoedus , Pystira , Zenodorus

Striking color pattern and body shape differences between the above genera are demonstrated on Figs 39 M- P, not less convincing differences in genitalic characters are documented on Figs 1B, 1C , 38 G-I. Merging of these genera by Zhang J. & Maddison (2012b: 21), done in search for molecular diagnostic data, went astray, taxonomically untested molecular experiments cannot overthrow obvious morphological differences. Their supporting arguments on similarities of palps are taken out of the context of general similarities of palps in about thousand species of EUOPHRYINES and related groups, more suitable internal structures of epigyne were analyzed superficially ( Figs 1B [A-B], 38G-I), the authors have overlooked significant distinct differences. To asses correctly significance of diversity of these characters in research of this kind, the revision should be carried out on ALL relevant species (like those demonstrated in Figs 1B [D1-8, E1-8, G1- 8, H1-8 and F9-13]), single representatives, even type species, do not suffice in a case of composite, never revised genera. Incidentally authors committed formal mistake, basing definition of the genus Omoedus on non type species Omoedus piceus Simon, 1902 instead of Omoedus niger Thorell, 1881 ) (compare Figs. 1B [F1] with 1B[A]). Extensive documentation of these genera, showing ALL data known, can be seen at: http://www.peckhamia.com/salticidae/q10-Omo.html, http://www.peckhamia.com/salticidae/q10- Pystira .html, http://www.peckhamia.com/salticidae/q10-Zen.html, preliminary account at http://www.peckhamia.com/salticidae/ZZZ-Omoedus-synonymy_clas.html).

Therefore merging of the above genera is rejected, correct combinations of all names are listed below: Omoedus ephippigerus (Simon, 1885) = Pystira ephippigera (Simon, 1885) (TYPE SPECIES, correction of erroneous combination, see Figs 1B [B], 38G-I, 38H for extensive survey of diversity see also: http://www.peckhamia.com/salticidae/q10- Pystira .html).

[Species inquirenda pending revision of the type specimens kept in Museum in Genoa, transfer to Omoedus without providing their documentation unjustified: Omoedus cyanothorax (Thorell, 1881) = Pystira cyanothorax (Thorell, 1881) , Omoedus karschi (Thorell, 1881) = Pystira karschi (Thorell, 1881) , Omoedus nigripalpis (Thorell, 1877) = Pystira nigripalpis (Thorell, 1877) ], Omoedus versicolor Dyal, 1935 = Pystira [dubia] versicolor Dyal, 1935 (species unrecognizable from the original description, type specimes lost, any conclusions without examination of specimens from terra typica [formerly, in 1935, "Government college grounds" in Lahur, Pakistan] are unjustified), Omoedus durvillei (Walckenaer, 1837) = Zenodorus durvillei (Walckenaer, 1837) (TYPE SPECIES, correction of erroneous combination, see Figs 1B , 38I, for extensive survey of diversity see http://www.peckhamia.com/salticidae/q10-Zen.html), [likewise: Omoedus albertisi (Thorell, 1881) = Zenodorus albertisi (Thorell, 1881) , Omoedus arcipluvii (Peckham, Peckham, 1901) = Zenodorus arcipluvii (Peckham, Peckham, 1901) , Omoedus bernsteini (Thorell, 1881) = Zenodorus bernsteini (Thorell, 1881) , Omoedus brevis Zhang J., Maddison, 2012 = Zenodorus brevis (Zhang J., Maddison, 2012) (see Fig. 1B, 1C ), Omoedus danae (Hogg, 1915) = Zenodorus danae Hogg, 1915 , Omoedus darleyorum Zhang J., Maddison, 2012 = Zenodorus darleyorum (Zhang J., Maddison, 2012) (see Fig. 1B, 1C ), Omoedus lepidus (Guerin, 1834) = Zenodorus lepidus (Guerin, 1834) , Omoedus metallescens (Koch L., 1879) = Zenodorus metallescens (Koch L., 1879) , Omoedus meyeri Zhang J., Maddison, 2012 = Zenodorus meyeri (Zhang J., Maddison, 2012) (see Fig. 1B, 1C ), Omoedus microphthalmus (Koch L., 1881) = Zenodorus microphthalmus (Koch L., 1881) , Omoedus obscurofemoratus (Keyserling, 1883) = Zenodorus obscurofemoratus (Keyserling, 1883) , Omoedus omundseni Zhang J., Maddison, 2012 = Zenodorus omundseni (Zhang J., Maddison, 2012) (see Fig. 38), Omoedus orbiculatus (Keyserling, 1881) = Zenodorus orbiculatus (Keyserling, 1881) , Omoedus papuanus Zhang J., Maddison, 2012 = Zenodorus papuanus (Zhang J., Maddison, 2012) (see Fig. 1B, 1C ), Omoedus ponapensis (Berry, Beatty, Prószyński, 1996) = Zenodorus ponapensis Berry, Beatty, Prószynski, 1996 , Omoedus semirasus (Keyserling, 1882) = Zenodorus semirasus (Keyserling, 1882) , Omoedus swiftorum Zhang J., Maddison, 2012 = Zenodorus swiftorum (Zhang J., Maddison, 2012) (see Fig. 38), Omoedus tortuosus Zhang J., Maddison, 2012 = Zenodorus tortuosus (Zhang J., Maddison, 2012) (see Fig. 1B, 1C ).

Unrecognizable species of the above genera should be restituted to their original combination as to avoid increasing chaos, until positive definitions could be worked out: Omoedus jucundus (Rainbow, 1912) = Zenodorus jucundus (Rainbow, 1912) , Omoedus juliae (Thorell, 1881) = Zenodorus juliae (Thorell, 1881) , Omoedus marginatus (Simon, 1902) = Zenodorus marginatus (Simon, 1902) , Omoedus niger (Karsch, 1878) = Zenodorus niger (Karsch, 1878) [no replacement name needed], Omoedus pupulus (Thorell, 1881) = Zenodorus pupulus (Thorell, 1881) , Omoedus pusillus (Strand, 1913) = Zenodorus pusillus (Strand, 1913) , Omoedus rhodopae (Hogg, 1915) = Zenodorus rhodopae (Hogg, 1915) , Omoedus syrinx (Hogg, 1915) = Zenodorus syrinx Hogg, 1915 , Omoedus variatus (Pocock, 1899) = Zenodorus variatus (Pocock, 1899) , Omoedus varicans (Thorell, 1881) = Zenodorus varicans Thorell, 1881 , Omoedus wangillus (Strand, 1911) = Zenodorus wangillus Strand, 1911 .

Figures

Figure 1A. Convenient and most reliable morphological diagnostic characters in Salticidae. A, C, E, I-L - Psenuc sp., D - Psenuc courti Żabka, 1993, F - Afraflacilla reiskindi (Prószyński, 1992), G - Afraflacilla philippinensis (Prószyński, 1992), H - Psenuc vesporum (Proszynski 1992), A-D. Genitalic charaters: A-B - palp in standard position, C - epigyne, external view, D - internal structure of epigyne: note copulatory openings, ducts, spermathecae (with pheromone exuding pore, developed here duct like, pockets), E-H - morphological characters of this particular genus, E – lateral subocular row of stridulatory bristles on tubercles, F - modified leg I, G - variation in shape and spination of tibia I, left and right in the same specimens, upper male, lower - female, H - male and female variation in tibia I in another species, I-K - body shape and color pattern in male, L - the same in female.