Ummidia gertschi, Godwin & Bond, 2021

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia gertschi sp. nov. Figs 30 , 31 , Map 4

Type material.

HOLOTYPE: 1 ♂ (UMM165) and PARATYPE: 1 ♀ (UMM572) from 5 miles west of Portal, Southwestern Research Station, Cochise County, Arizona, 31.8842 -109.206 5, 1645 m a.s.l., 10-20.vii.1955, coll. WJ Gertsch, AMNH.

Etymology.

The specific epithet is a patronym named for Dr. Willis J. Gertsch, famous arachnologist who put decades of work into a revision of Ummidia and the then Ctenizidae but was unable to finish before his passing; and whose notes, stored in the archives of the American Museum of Natural History, proved invaluable in the completion of this revision.

Diagnosis.

Ummidia gertschi can be differentiated from U. rosillos by the lack of a pale dorsal heart patch and brush on the retrolateral face of tarsus IV. Males can be differentiated from U. rosillos , U. modesta , U. timcotai , U. gabrieli , and U. rodeo by having an embolus with an even curve and from U. mercedesburnsae by having relatively more spines on the retrolateral face of tibia I. Males can also be differentiated from all other geographically proximal species except U. paulacushingae by having a distinct comb on the retrolateral face of tarsus IV. Females can be distinguished from all other geographically proximal species by possessing simple, straight spermathecae. Males disperse from July to August.

Description of male holotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Specimen in poor condition. General coloration. Carapace and chelicerae very dusky red 10R 2.5/2, legs dark reddish brown 2.5YR 2.5/4. Abdomen dark reddish brown 5YR 2.5/2. Cephalothorax. Carapace 6.04 long, 5.9 wide. Pars cephalica 3.91 long. Foveal groove procurved, 0.49 long, 1.22 wide. All eyes on moderate tubercle. AER procurved. PER procurved. Eye group 0.88 long, 1.42 wide, AME 0.38, PME 0.25, ALE 0.39, PLE 0.22. Sternum sparsely setose around outer 1/3, thicker anteriorly, STRl 3.48, STRw 3.69. Chelicerae with anterior tooth row comprising seven teeth, posterior margin with five teeth. Palpal endites with 27 long, thin cuspules over proximal half of endite face, lacking distal endite cuspules, ENDw 1.39, ENDl 2.51. Labium with six long, thin cuspules, LBw 1.44, LBl 0.99. Rastellum with many teeth on process. Abdomen setose. Legs. F1 5.73; F1w 1.42; P1 2.79; Ti1 3.86; Mt1 2.96; Tr1 1.23; F3 4.1; F3w 1.63; P3 2.17; Ti3 2.45; Sd3 1.32; Mt3 2.4; Tr3 1.62; F4 5.25; F4w 1.58; P4 2.5; Ti4 3.46; Mt4 3.87; Tr4 1.75. Retrolateral face of tarsus IV with proximal paired setae and with comb of spinules over distal half. Leg I spination pattern: TSp 4, TSpv 8, TSrd 0, TSr 0, TSrv 21, MtSp 4, MtSr 9, TrSp 4, TrSr 9. Pedipalps. PTl 3.17, PTw 1.25, Bl 2.78. Embolus evenly curved.

Varia tion, males

(n = 17). CL 4.04-7.65, 5.92 ± 0.26; CW 4.1-7.5, 5.83 ± 0.23; Cap 2.76-5.11, 3.93 ± 0.17; ENDl 0.59-1.07, 0.8 ± 0.04; ENDw 1.09-1.84, 1.39 ± 0.05; STRl 2.56-4.41, 3.44 ± 0.14; STRw 2.46-4.51, 3.47 ± 0.14; LBl 0.67-1.2, 0.95 ± 0.04; LBw 0.92-1.56, 1.29 ± 0.05; F1 4.18-6.85, 5.53 ± 0.18; F1w 0.84-1.87, 1.41 ± 0.07; P1 1.97-3.46, 2.73 ± 0.1; Ti1 2.79-4.51, 3.7 ± 0.12; Mt1 1.94-2.99, 2.59 ± 0.09; Tr1 0.99-3.8, 1.36 ± 0.16; F3 2.94-4.92, 3.98 ± 0.14; F3w 1.18-2.22, 1.63 ± 0.07; P3 1.48-2.69, 2.1 ± 0.09; Ti3 1.74-3.12, 2.41 ± 0.1; Mt3 1.77-2.69, 2.27 ± 0.07; Tr3 1.23-2.15, 1.73 ± 0.07; F4 3.75-6.23, 5.16 ± 0.19; F4w 1.1-2.64, 1.63 ± 0.09; P4 1.72-3.12, 2.44 ± 0.09; Ti4 2.53-4.18, 3.48 ± 0.12; Mt4 1.76-4.37, 3.58 ± 0.16; Tr4 1.19-2.49, 1.87 ± 0.08; TSp 0-10, 4.76 ± 0.66; TSpv 0-17, 7.59 ± 0.88; TSr 0-7, 0.65 ± 0.41; TSrv 15-39, 24.06 ± 1.32; PTl 2.53-3.93, 3.24 ± 0.1; PTw 0.95-1.55, 1.25 ± 0.04; BL 1.97-3.18, 2.63 ± 0.08.

Description of female paratype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. Left leg IV Mt-Tr and abdomen removed, in vial with specimen. General coloration. Carapace, chelicerae, and legs dark reddish brown 5YR 3/4. Abdomen very dark brown 7.5YR 2.5/3, spinnerets dark yellowish brown 10YR 4/6. Cephalothorax. Carapace 8.08 long, 7.93 wide. Pars cephalica 5.55 long. Foveal groove procurved, 0.65 long, 1.7 wide. Eye tubercle low under ME. AER procurved PER straight. Eye group 1.0 long, 1.87 wide, AME 0.44, PME 0.33, ALE 0.56, PLE 0.34. Sternum with posterior fringe, sparsely setose anteriorly STRl 5.01, STRw 5.28. Chelicerae with anterior row comprising eight teeth, posterior margin with six teeth. Palpal endites with 40 cuspules spread across proximal half of endite and 40 cuspules distally, equal in size to proximal cuspules anteriorly, smaller posteriorly, ENDw 2.04, ENDl 3.22. Labium with five cuspules, LBw 1.99, LBl 1.35. Rastellum with many strong spines on process and up cheliceral face. Abdomen. Evenly setose with pale speckles creating light chevrons at apodemes. Legs. F1 5.32; F1w 1.78; P1 3.31; Ti1 3.01; Mt1 2.27; Tr1 1.04; F3 4.37; F3w 2.25; P3 2.88; Ti3 2.33, Sd3 1.55; Mt3 2.2; Tr3 1.86; F4 5.5; F4w 2.31; P4 3.11; Ti4 3.15; Mt4 3.39; Tr4 1.89. Retrolateral face tarsus IV with defined comb over length of tarsus with proximal cluster of spinules. Pedipalps. PF 4.59, PP 2.62, PTi 2.93, PTr 2.78. Spermathecae simple with medial tilt, bulbs facing medioventrally.

Variation, females

(n = 10) CL 6.23-9.72, 7.9 ± 0.32; CW 6.23-9.06, 7.51 ± 0.31; Cap 4.18-6.78, 5.54 ± 0.24; ENDl 0.92-1.2, 1.05 ± 0.03; ENDw 1.55-2.2, 1.83 ± 0.06; STRl 3.83-5.7, 4.77 ± 0.2; STRw 4.02-6.12, 5 ± 0.24; LBl 1-1.62, 1.32 ± 0.06; LBw 1.39-2.36, 1.83 ± 0.09; F1 4.14-6.27, 5.11 ± 0.21; F1w 1.55-2.27, 1.89 ± 0.08; P1 2.71-4, 3.28 ± 0.15; Ti1 2.49-3.98, 3.12 ± 0.15; Mt1 1.91-2.92, 2.34 ± 0.12; Tr1 0.98-1.63, 1.21 ± 0.07; F3 3.61-5.25, 4.36 ± 0.17; F3w 1.88-2.78, 2.28 ± 0.1; P3 2.31-3.54, 2.84 ± 0.13; Ti3 2.09-3.17, 2.54 ± 0.12; Mt3 1.75-2.62, 2.05 ± 0.09; Tr3 1.52-2.18, 1.78 ± 0.08; F4 4.58-6.66, 5.46 ± 0.22; F4w 1.93-2.83, 2.31 ± 0.1; P4 2.55-3.79, 3.1 ± 0.13; Ti4 2.67-3.64, 3.11 ± 0.12; Mt4 2.75-3.87, 3.22 ± 0.13; Tr4 1.41-2.27, 1.8 ± 0.09; PF 4.06-5.59, 4.66 ± 0.17; PP 2.29-3.27, 2.73 ± 0.1; PTi 2.34-3.65, 2.92 ± 0.14; PTr 2.15-3.26, 2.6 ± 0.11.

Material examined.

United States: Arizona: Cochise Co: 5 mi W of Portal, 31.9142 -109.226 5, 1907 m a.s.l. (UMM0165, 10-20.vii.1955, 1♂, WJ Gertsch, AMNH); 5 mi W Portal, Southwestern Research Station, 31.8842 -109.206 5, 1645 m a.s.l. (UMM0348, 1.viii.1955, 1♂, WJ Gertsch, AMNH); (UMM0380, 10.ix.1970, 1♂, WJ Gertsch, AMNH); (UMM0351, 10.viii.1956, 1♂, E Ordway, AMNH); (UMM0356, 10-20.vii.1955, 1♂, WJ Gertsch, AMNH); (UMM0572, 1♀, AMNH); (UMM0254, 12.vii.1992, 1♂, S Lingafelter, AMNH); (UMM0350, 15-20.viii.1968, 1♂, V Roth, AMNH); (UMM0358, 19.viii.1965, 1♂, Mrs Anderson, AMNH); (UMM0266, 19.viii.1988, 1♂, 1♀, J Rozen, E Brewster, AMNH); (UMM0360, 20.vii.1962, 1♂, WJ Gertsch, AMNH); (UMM0470, 20.vii.1976, 1♂, S Johnson, AMNH); (UMM0344, 21.viii.1966, 1♂, B Rozen, AMNH); (UMM0324, 23.viii.1974, 1♂, AMNH); (UMM0359, 25.viii.1966, 1♂, Rozens, AMNH); (UMM0395, 26.viii.1955, 1♂, WJ Gertsch, AMNH); (UMM0550, 29.vii.1976, 1♂, S Johnson, AMNH); (UMM0385, 30.viii.2003, 1♂, JG Rozch, AMNH); (UMM0386, 31.vii.1956, 1♂, E Ordway, AMNH); (UMM0612, 31.vii.1968, 1♀, SE Roth, AMNH); (UMM0618, 7.vii.1966, 1♂, B Nockumas, AMNH); (UMM0595, 7.vii.1972, 1juv, Riechart, AMNH); (UMM0361, 7.viii.1963, 1♂, K Muma, AMNH); (UMM0394, iv.1956, 1♀, WJ Gertsch, E Ordway, AMNH); 5 mi W Portal, Southwestern Research Station, 31.8842 -109.206 5, 1645 m a.s.l. (UMM0661, 1♂, AMNH); Chiricahua Mountains, 31.8831 -109.2829 6, 2751 m a.s.l. (UMM0387, 15.vii.1961, 1♂, DJ Knull, JN Knull, AMNH); (UMM0355, vii-viii.1972, 1♂, JAL Cooke, AMNH); Chiricahua Mountains, Southwestern Research Station, 31.8842 -109.206 5, 1645 m a.s.l. (UMM0336, 10.ix.1979, 1♂, V Roth, AMNH); (UMM0276, 14.ix.1967, 1♂, AMNH); (UMM0375, 14.viii.1969, 1♂, AMNH); (UMM0337, 17.viii.1981, 1♂, AMNH); (UMM0376, 18.viii.1982, 1♂, AMNH); (UMM0357, 2.viii.1971, 1♂, AMNH); (UMM0372, 21.vii.1981, 1♂, AMNH); (UMM0378, 25.viii.1974, 1♂, AMNH); (UMM0352, 28.vii.1970, 1♂, AMNH); (UMM0342, 28.viii.1979, 1♂, AMNH); (UMM0379, 31.viii.1968, 1♂, AMNH); (UMM0374, 4.ix.1967, 1♂, AMNH); (UMM0373, 8.vii.1984, 1♂, AMNH); (UMM0338, 22.vii.1971, 1♂, T Haley, AMNH); (UMM0393, 22-24.viii.1980, 1♂, D Hawks, AMNH); (UMM0343, 23.vii.1964, 1♂, SE Roth, AMNH); (UMM0368, 5.ix.1978, 1♀, D Hawks, AMNH); (UMM0370, 7.viii.1967, 1♂, D Rich, AMNH); Chiricahua Mountains, Southwestern Research Station, 31.8842 -109.206 5, 1645 m a.s.l. (UMM0391, 30.vii.1968, 1♂, AMNH); Huachuca Mountains, Miller Canyon, 31.4177 -110.2775 6, 2059 m a.s.l. (UMM0185, 27.ii.1972, 1♀, DB Richman, MS Richman, UFMNH); Huachucha Mountains, Ramsey Canyon, 31.4486 -110.3087 4, 1710 m a.s.l. (UMM0346, 20-25.viii.1932, 1♂, WS Creighton, AMNH); 1mi S Portal, 31.899 -109.1492 4, 1833m (UMM0620, 23.viii.1966, 1♂, AMNH); Portal, 31.9125 -109.1421 5, 1457 m a.s.l. (UMM0690, 15.vii.1965, 1♂, M Muma, AMNH); 31.9125 -109.1421 6, 1457 m a.s.l. (UMM0688, 28.ix.1983, 1♂, M Muma, AMNH); (UMM0390, 15.vii.1965, 1♂, WJ Gertsch, AMNH); (UMM0349, 19.viii.1967, 1♂, AMNH); (UMM0389, 25.vii.1970, 1♂, AMNH); (UMM0354, vii.1968, 1♂, AMNH); (UMM0371, 20.vii.1980, 1♂, V Roth, H Joboff, AMNH); (UMM0549, 22.viii.1970, 1♂, L Anderson, AMNH); (UMM0317, 24.viii.1964, 1♂, J Rozen, AMNH); (UMM0353, 30.vi.1967, 1♂, T Smell, AMNH); Portal, Jensen’s Store, 31.9137 -109.1414 6, 1453 m a.s.l. (UMM0347, 1.viii.1971, 1♂, S Roth, AMNH); 5mi W Portal, Southwestern Research Station, 31.8842 -109.206 5, 1645 m a.s.l. (UMM0206, 13-22.viii.1972, 1♂, N Platnick, MCZ); Coconino Co: Jct I-40 and Meteor Crater Road, 35.1105 -111.0316 4, 1639 m a.s.l. (UMM849, 1♂, E Riley, M Yoder, TAMU); Graham Co: Galiura Mountains, 10 mi W Sunset, 32.6011 -110.3533 6, 1753 m a.s.l. (UMM0181, 18.vii.1956, 1juv, MJ Westfall, UFMNH); Pinaleno Mountains, 32.641 -109.8436 6, 2824 m a.s.l. (UMM0153, 15.vii.1917, 1♀, CUNHC); Maricopa Co: Phoenix, South Mountain, 33.3304 -112.092 5, 560 m a.s.l. (UMM0417, 20.viii.1966, 1♂, SC Williams, AMNH); Pima Co: Santa Catalina Mountains, Bear Canyon Picnic Area, 32.3154 -110.791 4, 934 m a.s.l. (UMM0455, 3.vii.1961, 1♂, FG Werner, AMNH); Santa Catalina Mountains, Molina Canyon, 31.3022 -110.7178 5, 871 m a.s.l. (UMM0168, 25.vii.1970, 1♂, K Stephens, AMNH); Tucson, 32.222 -110.9262 6, 759 m a.s.l. (UMM0157, 15.vii.1953, 1♂, GM Bradt, AMNH); (UMM0616, 15.vii.1953, 1♂, GM Bradt, AMNH); (UMM0438, 9.v.1909, 1♀, G Bradt, AMNH); (UMM0468, vii.1950, 1♂, RE Crandall, AMNH); Pinal Co: San Tan Valley, 33.1788 -111.4942 1, 474 m a.s.l. (UMM0662, 7.ii.2016, 1♀, TA Cota, BME); (UMM0664, 1♀, BME); (UMM0665, 1♀, BME); New Mexico: Grant Co: Fort Bayard, 32.7961 -108.1502 5, 1867 m a.s.l. (UMM0689, 20.viii.1976, 1♂, M Muma, AMNH); Hidalgo Co: off Hwy # 9, 7 mi W Aimes, 31.9406 -108.7077 7, 1372 m a.s.l. (UMM0508, 1♂, B Tomberlin, T Snell, AMNH).

Figures

Map 4. Locality records for western United States and northern Mexico species.

Figure 30. Ummidia gertschi sp. nov. from Cochise Co., Arizona. A male habitus illustration UMM 165 B female habitus illustration UMM 664. Scale bars: 4.0 mm.

Figure 31. Ummidia gertschi sp. nov. from Cochise Co., Arizona A-D male holotype (UMM 165) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female exemplar (UMM 664) F cleared spermathecae female paratype (UMM 572). Scale bars: 1.0 mm.