Trochomorpha hermia Hutton, 1883

Brook, Fred J., Kennedy, Martyn, King, Tania M., Ridden, Johnathon, Shaw, Matthew D. & Spencer, Hamish G., 2020, Catalogue of New Zealand land, freshwater and estuarine molluscan taxa named by Frederick Wollaston Hutton between 1879 and 1904, Zootaxa 4865 (1), pp. 1-73 : 57

Trochomorpha hermia Hutton, 1883

Pl. 7, fig. D

Hutton, 1883. The New Zealand Journal of Science, 1: 477.

Type material. Lectotype, CMNZ M353 , and possible paralectotype (1), NMNZ M.180049 (dry shells) . Lectotype fixed by inference of holotype ( ICZN Art. 74.6) by Freeman et al. (1997: 32) .

CMNZ molluscan catalogue details. M353—‘ Trochomorpha ? hermia Hutton, Manawatu , Wellington, 1 specimen (135)’.

Label details. NMNZ M.180049—‘ Trochomorpha ? hermia Hutt., Manawatu, T.W. Kirk Coll., Pres. Tawera a Kapiti Lodge’, in R.K. Dell’s handwriting.

Type locality. Stated by Hutton (1883g: 477, 1884b: 183) to be ‘Manawatu’, but this is incorrect (see below).

Remarks. Type material of Trochomorpha hermia is illustrated here for the first time in pl. 7 fig. D. Hutton submitted a description of Trochomorpha (?) hermia to the Transactions and Proceedings of the New Zealand Institute issue for 1883, but publication was delayed until May 1884 ( Hutton 1884b: 183), and was pre-empted by a brief description of Trachomorpha [sic] (?) hermia in an account of a meeting of the Philosophical Institute of Canterbury ( Hutton 1883g: 477). These descriptions were based on specimens that T.W. Kirk sent to Hutton, and which reportedly had been collected in Manawatu, north of Wellington. Suter (1892c: 282) noted that hermia “corresponds quite well” with Helix (Helicigona) inversicolor Férussac [family Helicarionidae ], from Mauritius, and inferred that the type material was mislocalised, and not from New Zealand. Suter (1913: 684) stated that hermia is not a New Zealand species, and treated it as a synonym of Pachystyla inversicolor (Férussac, 1821) , as did Freeman et al. (1997: 32), but this name is a nomen nudum, as pointed out by Kennard (1943: 123), and the currently recognised name of the Mauritian taxon is Pachystyla bicolor (Lamarck, 1822) (e.g., Griffiths & Florens 2006: 127– 128, pl. 32. figs. C–F). Subadult shells of P. bicolor are similar to Trochomorpha hermia , but the two taxa have very different teleoconch microsculpture, and are not conspecific (Owen Griffiths pers. comm. 2019). During the present study Michael Shea and Isabel Hyman (pers. comm. 2019) pointed out that, rather than being from Mauritius, Trochomorpha hermia is in fact conspecific with Helix campbellii Gray, 1834 , which is endemic to Norfolk and Philip islands, Australia. The latter is the type species of Advena Gude, 1913 , and belongs in Microcystinae Thiele, 1931 (Isabel Hyman unpub. data). Advena campbelli was described from material collected on Philip Island ( Gray 1834: 65); a syntype (NHMUK 1982239) is illustrated here for the first time in pl. 7, fig. E. Advena campbellii charon Preston, 1913 , described from material collected on Norfolk Island ( Preston 1913: 526, Iredale 1945: 66, Smith 1992: 226), is a synonym, according to Hyman (2005) and Stanisic et al. (2010: 346, 564).

Taxonomy. Treated here as a subjective junior synonym of Advena campbellii (Gray, 1843) N. syn.

Family RHYTIDIDAE Pilsbry, 1893