Tomoglossa ornatella ( Casey, 1910 ) Gusarov, 2002

Gusarov, Vladimir I., 2002, A revision of Nearctic species of the genus Tomoglossa Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae), Zootaxa 30, pp. 1-19 : 10-13

2. Tomoglossa ornatella ( Casey, 1910) , comb. nov. ( Figs. 29 , 34­37 )

Noverota ornatella Casey, 1910: 90 .

Type material: lectotype (here designated),, UNITED STATES: “N[ew].J[ersey].”, “ Noverota ornatella Csy.”, “Casey bequest 1925”, “ TYPE USNM 39099” (red label), “ Lectotype Noverota ornatella Casey. V.I.Gusarov des. 2002” (red label) (NMNH). While describing N. ornatella , Casey did not specifically state how many specimens were available.

Diagnosis. In comparison to T. decora , T. ornatella has elongate antennal articles 4­5 ( Figs. 29, 31 ), thicker spermatheca with different shape of umbiculus ( Figs. 36 , 26 ) and larger female accessory sclerite ( Figs. 37 , 28 ). Tomoglossa ornatella is very similar to T. improvisa but has deeper puncturation of the forebody. Compared to T. arizonica and T. floridana , T. ornatella has elongate antennal articles 4­5, less transverse articles 6­10 and longer body ( Figs. 29, 32­33 ). Tomoglossa ornatella also differs from T. floridana in having larger spermatheca with different shape of umbiculus ( Figs. 36 , 45 ) and larger female accessory sclerite ( Figs. 37 , 46 ).

Description. Length 2.0 mm. Head and abdominal tergum VI brown; pronotum, elytra, abdominal segments III­V, VII­VIII and antennal articles 2­11 reddish yellow; apical fifth of elytra, antennal article 1, mouthparts and legs yellow.

Head transverse, surface on disk glossy, without visible (at x100) microsculpture, with fine and dense punctures, distance between them equal to ¼­½ their diameter. Antennal article 2 longer than 3, 4­5 elongate, 6­10 subquadrate, last article as long as 9 and 10 combined ( Fig. 29 ).

Pronotum transverse, width to length ratio 0.46 mm: 0.36 mm, wider than head (pronotum width to head width ratio 0.46 mm: 0.37 mm); surface glossy, without visible microsculpture, puncturation finer than on head, distance between punctures equals ½ their diameter. Elytra measured from humeral angle shorter than wide (0.44 mm: 0.54 mm); surface glossy, without microsculpture and with puncturation as on pronotum.

Abdominal terga without microsculpture, and with fine and sparse puncturation, puncturation becoming finer towards abdominal apex, on terga III­V distance between punctures within rows equal to their diameter and between rows 2­3 times their diameter, on tergum VI distance between punctures within rows equal to 1­2 times their diameter and between rows 3­4 times their diameter, on tergum VII distance between punctures within rows equal to 2­3 times their diameter and between rows 3­5 times their diameter. Tergum VII with white edge.

Spermatheca with wide umbilicus ( Fig. 36 ). Female accessory sclerite large ( Fig. 37 ).

Distribution. Known only from New Jersey ( Fig. 47 ).

Natural History. Casey (1910) did not provide habitat information.

Figures

FIGURE 47. Geographical distribution of Nearctic Tomoglossa. T. ornatella (Casey) (c), T. improvisa (Casey) (⌂), T. decora (Casey) (z), T. arizonica Gusarov, sp. nov. (S), T. floridana Gusarov, sp. nov. („).

FIGURES 29 ­ 33. Antennae of Nearctic Tomoglossa. 29, T. ornatella (Casey) (lectotype); 30, T. improvisa (Casey) (lectotype); 31, T. decora (Casey) (Pennsylvania); 32, T. arizonica Gusarov, sp. nov. (holotype); 33, T. floridana Gusarov, sp. nov. (holotype). Scale bar 0.1 mm.

FIGURES 34 ­ 37. Details of Tomoglossa ornatella (Casey) (lectotype). 34, female tergum VIII; 35, female sternum VIII; 36, spermatheca; 37, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (34 ­ 35), 0.1 mm (36 ­ 37).

FIGURES 21 ­ 28. Genitalia and terminalia of Tomoglossa decora (Casey) (Vermont (21 ­ 25), Pennsylvania (26 ­ 28 )). 21, median lobe of aedeagus, lateral view; 22, median lobe of aedeagus, ventral view; 23, internal sac of aedeagus, in packed state; 24, deeply concave sclerite of internal sac; 25, copulatory piece; 26, spermatheca; 27, left portion of female sternum IX; 28, female accessory sclerite. U ­ umbilicus. Scale bar 0.1 mm.

FIGURES 43 ­ 46. Details of Tomoglossa floridana Gusarov, sp. nov. (holotype). 43, female tergum VIII; 44, female sternum VIII; 45, spermatheca; 46, female accessory sclerite. U ­ umbilicus. Scale bar 0.2 mm (43 ­ 44), 0.1 mm (45 ­ 46).