Tokyosoma fanfan, Mikhaljova, Elena V., Golovatch, Sergei I. & Chang, Hseuh-Wen, 2010

Mikhaljova, Elena V., Golovatch, Sergei I. & Chang, Hseuh-Wen, 2010, The millipede family Diplomaragnidae in Taiwan, with descriptions of nine new species (Diplopoda, Chordeumatida), Zootaxa 2615, pp. 23-46 : 31-32

Tokyosoma fanfan sp. nov.

Figs 16–19 .

Material examined: Holotype: 1 male ( NMNS –6345–009), Taiwan, Ilan County, Datong Township, Chilan, SE foot of Mt Fanfan , 24°36.840’ N, 121°29.101’ E, 1031 m, mixed forest with streams, 18 October 2009, leg. L. Dányi & E. Lazányi.

Diagnosis: Differs from congeners mainly by the shape of the posterior gonopod lateral coxal branch with a strong unciform process laterally and a serrate apex, as well as in the shape of the posterior gonopod colpocoxite supplied with a long subconical outgrowth distally.

Description: Male. Length about 23 mm, width with paraterga about 2.0 mm. Coloration in alcohol marbled brown with a pattern of transverse pale and dark brown bands on dorsum, increasingly intensely coloured towards telson. Legs with marbled light brown distal parts. Venter, proximal parts of legs and lower portions of pleura pale. Head light marbled brown with a pale anterior portion. Ocellaria black. Antennae brown with pale proximal parts.

Body with 32 segments. Head covered with sparse, relatively long and short setae. Each eye patch composed of at least 30 ocelli. Collum semi-circular. Body width gradually increasing until somite 7, body parallel-sided on somites 8–14(15), thereafter gradually tapering. Beginning from somite 4, paraterga normally developed, growing increasingly less distinct towards hind part of body, strongly reduced on somites 26 and 27, absent from somites 28–31. Paraterga of somites 2 and 3 small. Metazonital macrochaetae in a transverse row on somites 30 and 31, like an elongate (to different degrees) triangle on preceding somites. Nearly all macrochaetae broken off, remaining ones in middle part of body short and pointed apically, but not very sharply so. In anterior and posterior parts of body posterolateral macrochaetae longest, pointed apically, anterolateral and medial ones subequal in length, pointed apically, but not very sharply so. Metazonites with two tiny knobs placed near axial suture on each side.

Leg pairs 3–7 somewhat enlarged. Leg pairs 6 and 7 with a small group of funnel-shaped tarsal papillae apically near claw. Claw of leg pairs 6 and 7 at base with a long setoid filament ventrally, but without additional claw dorsally. Postgonopodal legs (including leg pairs 10 and 11) without tarsal papillae. Claw of legs 10 and 11 at base with one small additional claw dorsally and a long setoid filament ventrally. Claw of midbody legs at base with one small additional claw dorsally and a long setoid filament ventrally. Claw of hindmost legs at base with a tiny additional claw dorsally, but without filament ventrally.

Legs 10 and 11 with coxal glands. Coxa 10 with a caudoventral, oblong, somewhat concave, shagreened process with an anteriad curved rounded apex covered with papillae, and with a small, frontal, subapical knob with a seta ( Fig. 16 ). Trochanter 10 with a tiny ventral outgrowth setose apically. Coxa 11 covered with low papillae mainly caudally, without processes ( Fig. 17 ). Trochanter 11 with a caudal process papillate mesally and rounded apically.

Anterior gonopod telopodite 1-segmented, flagelliform, beset with cuticular spinules, its distal part positioned inside sheaths with strongly elevated and tightly closed edges (e) ( Fig. 18 ). These sheath edges like high plates in lateral view. Telopodite base and distal part of coxosternum tightly attached to adjacent mesal portion of posterior gonopod. Posterior gonopod colpocoxites fused sub-basally. Colpocoxite with a distal, slender, subconical process (c) somewhat curved posteriorly, and with a lateral rounded blade (b). Mesal sheath processes of posterior gonopod colpocoxites fused medially into a single cup-shaped structure (ms) carrying pointed spinules. Lateral sheath processes of colpocoxites (lp) cup-shaped, covered with short pointed spinules caudally.

Posterior gonopod angiocoxite with a globule, but without process in posterior view. Posterior gonopod coxal part with a long, lateral, flat branch (lb) curved mesally and then, laterally, supplied with a strong lateral unciform (p) and a dentiform subapical process (sp). Apex of the branch serrate. Basal part of this branch fused with both colpocoxite and anterior angiocoxite.

Angiocoxite depressed centrally in anterior view ( Fig. 19 ), supplied with a long process (ap); distal portion of this process penetrating colpocoxite and visible outside in anterolateral view. Posterior gonopod telopodite 2-segmented; trochanteroprefemur hollow for accommodation of lateral branch (lb) mesocaudally; femur short and thin.

Female unknown.

Name: The specific epithet refers to the type locality, a noun in apposition.

Figures

FIGURES 16 – 19. Tokyosoma fanfan sp. nov., male holotype. 16, coxa and trochanter 10, front view; 17, coxa and trochanter 11, caudal view; 18, gonopods, caudal view; 19, gonopods, front view; e, edge of posterior gonopod colpocoxite sheath; c, distal process of posterior gonopod colpocoxite; b, posterior gonopod colpocoxite lateral blade; ms, mesal sheath processes of posterior gonopod colpocoxites; lp, lateral sheath process of posterior gonopod colpocoxite; lb, posterior gonopod lateral coxal branch; p, lateral process of posterior gonopod lateral coxal branch; sp, aubapical process of posterior gonopod lateral coxal branch; ap, anterior process of posterior gonopod angiocoxite. Scale in mm.