Timbellus radulfiensis, Merle & Pacaud & Ledon & Goret, 2024

Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel & Goret, Bernard, 2024, New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe, Geodiversitas 46 (15), pp. 495-551 : 520-521

Timbellus radulfiensis n. sp.

( Figs 15 ; 26 N-P)

urn:lsid:zoobank.org:act:ADEDF3CF-00EE-4FBA-8D11-3435CD75F8A5

TYPE MATERIAL. — Holotype. France • Normandy, Manche, Rauville-la-Place ; Argiles à Corbules Formation ; late Eocene (Priabonian); MNHN.F.A77754 ( MNHN coll.), H: 13.5 mm ( Figs 15A, B ; 26N ). Paratypes. France • 1 spm; same as for the holotype; MNHN.F.A77755 ( MNHN coll.) ( Figs 15C ; 26O ) • 3 spm; idem; MNHN.F.A77756 ( MNHN coll.) • 3 spm; idem; MNHN.F.A24118 (Vasseur coll.) • 1 spm; idem; MNHN.F.A90559 (Vasseur coll.) ( Figs 15D ; 26P ).

ETYMOLOGY. — From the ancient name of the type locality Radulfi villa around 1083.

TYPE HORIZON. — Argiles à Corbules Formation, late Eocene (Priabonian), stratigraphic data given byCavelier (1979) and Chateauneuf (1980).

TYPE LOCALITY. — France, Normandy, Manche, Rauville-la-Place (Cauvin), Cavelier (1979).

DISTRIBUTION. — Only know from the type locality.

DESCRIPTION

Bulbous protoconch of one whorl.Teleoconch up to 20.5 mm in height, up to 7.5 mm in width, subfusiform in profile, composed of six whorls. Moderately high spire. Last whorl up to 74% of total length. Apical angle 61°. Spiral sculpture with moderately marked primary cords. First whorl: no cord. Second whorl: appearance of P2. Third whorl: appearance of P1 (weaker than P2) and s2. Fourth and fifth whorl: no change. Sixth whorl: P1 to P6, s1; P6 not atrophied; ADP, MP and ABP on siphonal canal. From first whorl to second whorl: ten protovarices. On third whorl: seven protovarices. Fourth whorl: appearance of varices; three varices and one intervarix intercalated. From fifth to sixth whorl: no change. Higher relief of intervarices on P2. Varices sublamellose. Appearance of short P1 spine on third whorl. Abapically spinelets from P1 to P6 and ADP. Ovate aperture up to 34% of diameter and up to 74% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, poorly erect anteriorly, forming curved inductura at base. Parietal lip slightly adherent. Outer lip with developed denticles including ID, D1 to D6. Outer lip not crenulate. Siphonal canal open, straight, up to 47% of aperture length. Pseudoumbilicus narrow.

COMPARISONS

This species can be compared to other Priabonian and Rupelian species from Northern Europe ( Germany) and Eastern Europe ( Ukraine) with which it shares a narrow shape. It differs from T detritus (von Koenen, 1890) , T. arenarius ( Steuer, 1912) and T. danapris Pacaud (2018) in having finer spiral cords. The spiral sculpture resembles more that of T. priabonicus Pacaud, 2018 (nom. n. for Murex trialatus von Koenen, 1889 , non G. B. Sowerby (II), 1834), but T. priabonicus bears strongly developed P1 and P2 spines, whereas only P1 spine is developed in T. radulfiensis n. sp.

Figures

FIG. 15. — Spiral cords and internal denticles of the outer lip in Timbellus radulfiensis n. sp.: A, B, holotype, MNHN.F.A77754 (MNHN. coll.), Rauville-la-Place, Manche, Normandy, France, late Eocene (Priabonian); C, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; D, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: 5 mm. Credits: A-C, P. Loubry (MNHN/CNRS); D, L. Cazes (MNHN/CNRS).

FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm.