Thyropygus enghoffi ( Demange, 1989 )

Pimvichai, Piyatida, Enghoff, Henrik & Panha, Somsak, 2009, A revision of the Thyropygus allevatus group. Part 2: the T. bifurcus subgroup (Diplopoda, Spirostreptida, Harpagophoridae), Zootaxa 2165, pp. 1-15 : 5-6

Thyropygus enghoffi ( Demange, 1989)

( Figs. 2 A–D, 7B)

Thyropisthus enghoffi Demange, 1989: 779 Thyropygus enghoffi: Enghoff 2005: 94 Cornugonus enghoffi: Jeekel 2006: 19

Material: HOLOTYPE male THAILAND, Phang Nga Province, Poung-chang cave, 8° 31ˏ 40˝ N, 98° 29ˏ 58˝ E. 19 August 1987. L. Deharveng leg., ( MNHN), 2 males THAILAND, Phang Nga Province, Khao Lak. 28 August–13 September 2008. P. Decker leg., ( ZMUC), 1 male THAILAND, Phang Nga Province, Tai Muang district, Khao Lak Lam-Ru National Park, 8° 37ˏ 34˝ N, 98° 14ˏ 23˝ E. 17 October 2008. P. Prasankok leg., ( CUMZ), 1 male, 1 female THAILAND, Phang Nga Province, Mueang district, Nam Pud cave, 8° 27ˏ 47˝ N, 98° 32ˏ 30˝ E. 17 January 2009. P. Pimvichai and C. Sutcharit leg., ( CUMZ), 1 male THAILAND, Phuket Province, Phuket Island, Thalang district, Tonesai waterfall, 8° 3ˏ 47˝ N, 98° 21ˏ 7˝ E. 17 November 1984. P. Beron & S. Andreev leg., ( NMNH).

Diagnosis: A species of the bifurcus subgroup. Lateral margin of lateral process of anterior coxal fold (alp) without a small crest or spine. Similar in this respect to T. casjeekeli and T. richardhoffmani . Differs from all other species of T. bifurcus subgroup by having the mesal process of anterior coxal fold (amp) much shorter than alp.

Description: Adult males with 61–63 podous rings, no apodous rings. Length ca. 13–14 cm (broken), width ca. 8.0– 9.4 mm. Adult female with 63 podous rings, no apodous rings. Length ca. 10 cm, width ca. 7.5 mm. Overall color of living animal ( Fig. 7 B) brown. Legs brownish, metazona dark brown.

Gonopods ( Figs. 2 A–D): Anterior coxal fold (ac) ( Fig. 2 A): lateral process (alp) long, slender, regularly curved, sickle-shaped; mesal process (amp) much shorter than lateral process, curved, pointed. Posterior coxal fold (pc) ( Fig. 2 B) basally with moderately high lateral paracoxites (px); mesal process (pmp) distally rounded, visible in anterior view between alp and amp (In Demange’s fig. 11 pmp looks like an additional process between alp and amp); lateral process (plp) erect, digitiform; shelf (sh) for accommodation of telopodite between pmp and plp. Telopodite ( Figs. 2 C–D) leaving coxite over shelf of posterior coxal fold; double femoral spine (fe), the first fe (fe 1) longer than fe 2, slender, straight, terminally slightly curved, its tip in situ resting close to tip of amp, the second fe (fe 2) situated under fe 1, very short, slender, curved rearwards; tibial spine (ti) long, slender, curving in horizontal plane, its tip in situ resting close to base of pmp; apical part: lamellar lobe (ll) flattened, bent down; a small slender spine (ss) at base of apical part, opposite the origin of tibial spine; palette (pa) simple, flattened, with a longitudinal rounded crest (cr) near tip; distally with about ten brownish blepharochaetae (bp).

Distribution ( Fig. 8 ): Known from several localities in THAILAND, Phang Nga Province, and also from Phuket Island ( THAILAND, Phuket Province).

Figures

FIGURE 2. Thyropygus enghoffi, holotype, gonopods. A: anterior view, right telopodite removed. B: posterior view, right telopodite removed. C: right telopodite, posterior-mesal view. D: right telopodite, anterior-lateral view.

FIGURE 8. Map of peninsular Thailand, showing the known distribution of the species of the T. bifurcus subgroup.

FIGURE 7. A: Thyropygus demangei, living male. B: Thyropygus enghoffi, living male. C: Thyropygus casjeekeli, living male. D: Thyropygus quadricuspis, living male. E: Thyropygus richardhoffmani, living male.