Tesnus soekhnandanae, Makhan, 1995

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium IV. Staphylinine Group (Part 1) Euaesthetinae, Leptotyphlinae, Megalopsidiinae, Oxyporinae, Pseudopsinae, Solieriinae, Steninae, Bulletin of the American Museum of Natural History 2440 (265), pp. 1807-2440 : 1807-2440

soekhnandanae Makhan, 1995 , see: collaris Sharp, 1876 .

sogai Jarrige, 1970: 53 ( Stenus ; subgenus Hypostenus ; Type locality: Madagascar:

Mt. Tsaratanana, versant Sud, Andohanananila, 1850 m).

— Puthz, 1972: 20 ( Stenus ; characters; Madagascar).

— Puthz, 1992a: 13 ( Stenus ; Madagascar).

- DISTRIBUTION: Madagascar.

soleatus Puthz, 1973 n: 2 ( Stenus ; subgenus Parastenus ; Type locality: Ceylon: Central Prov.: Horton Plains, 2100 m, 12 mi. SSE Nuwara-Eliya).

- DISTRIBUTION: Sri Lanka.

solidus L. Benick, 1921c: 192 ( Stenus ; replacement name for curvipes Bernhauer ).

— Scheerpeltz, 1933: 1166 ( Stenus ; subgenus Stenus ; catalog).

— L. Benick, 1938c: 260 ( Stenus ; characters).

— Puthz, 1969j: 57 ( Stenus ; aedeagus).

— Puthz, 1974a: 68 ( Stenus ; characters).

- DISTRIBUTION: Venezuela.

curvipes Bernhauer, 1916f: 173 ( Stenus ; subgenus Stenus ; [preoccupied]; Type locality: Caracas).

— L. Benick, 1917b: 295 ( Stenus ; subgenus Stenus ; notes).

— Scheerpeltz, 1933: 1166 ( Stenus ; synonym of solidus ).

— Puthz, 1974a: 68 ( Stenus ; synonym of solidus ).

solieri Coiffait and Saiz, 1968 , see: gayi Solier, 1849 .

solitarius Sharp, 1886: 647 ( Stenus ; Type locality: Guatemala, Cerro Zunil 4000 to 5000 feet). — Bernhauer and Schubert, 1911: 171 ( Stenus ; subgenus Tesnus ; catalog).

— L. Benick, 1938c: 267 ( Stenus ; characters).

— L. Benick, 1939: 21 ( Stenus ; subgenus Tesnus ; characters; Costa Rica).

— L. Benick, 1949b: 567 ( Stenus ; subgenus Tesnus ; characters).

— Hromádka, 1986: 151 ( Stenus ; aedeagus).

- DISTRIBUTION: Guatemala, Costa Rica.

solstitialis Zheng, 1994d: 79 ( Stenus ; subgenus Parastenus ; Type locality: China: Sichuan: Wolong).

- DISTRIBUTION: China.

solutus Erichson, 1840: 734 ( Stenus ; Type locality: Lutetiae; [Note: See lectotype designation by Puthz, 1967h]).

— Leprieur, 1851: 199 ( Stenus ; characters).

— Fairmaire and Laboulbène, 1856: 595 ( Stenus ; characters; France).

— Rye, 1864: 110 ( Stenus ; characters; England).

— Fauvel, 1873: 110 [= 1873a: 268] ( Stenus ; characters; France; Britain).

— Solsky, 1874: 198 ( Philonthus ; characters; Turkestan).

— Rey, 1884: 395 [= 1884a: 243] ( Stenus ; subgenus Hypostenus ; characters; France).

— Fowler, 1888: 358 ( Stenus ; characters; Britain).

— Ganglbauer, 1895: 585 ( Stenus ; subgenus Hypostenus ; characters; western central Europe).

— Reitter, 1909: 162 ( Stenus ; subgenus Hypostenus ; characters; Germany).

— Bernhauer and Schubert, 1911: 177 ( Stenus ; subgenus Hypostenus ; catalog).

— Johansen, 1914: 527 ( Stenus ; subgenus Hypostenus ; characters; Denmark).

— L. Benick, 1929a: 56 ( Stenus ; subgenus Hypostenus ; characters in key; central Europe; Caspian region; Turkestan).

— Portevin, 1929: 406 ( Stenus ; characters; France).

— Scheerpeltz, 1933: 1182 ( Stenus ; subgenus Hypostenus ; catalog).

— Wüsthoff, 1934a: Tafel III ( Stenus ; illustration of aedeagus).

— Palm, 1953: 11 ( Stenus ; ecological notes; Sweden).

— Smetana, 1961b: 150 ( Stenus ; subgenus Hypostenus ; characters; Czechoslovakia).

— Szujecki, 1961: 47 ( Stenus ; subgenus Hypostenus ; characters; Poland).

— Horion, 1963: 368 ( Stenus ; subgenus Hypostenus ; Britain; Denmark; Sweden; Holland; Belgium; France; Switzerland; Italy; Czech Republic; Slovakia; Hungary; Romania; Serbia; Poland; Germany; Austria). — Lohse, 1964: 121 ( Stenus ; subgenus Hypostenus ; characters; central Europe). — Smetana, 1964c: 165 ( Stenus ; subgenus Hypostenus ; Czechoslovakia). — Focarile, 1964: 63 ( Stenus ; Italy). — Puthz, 1967h: 298 ( Stenus ; lectotype designation). — Szujecki, 1968a: 721 ( Stenus ; Poland). — Puthz, 1971v: 98 ( Stenus ; notes). — Puthz, 1972s: 263 ( Stenus ; Yugoslavia). — Pope, 1977: 26 ( Stenus ; Britain). — Lucht, 1987: 97 ( Stenus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark). — Schülke and Uhlig, 1988: 7 ( Stenus ; notes; Germany). — Ryvkin, 1990: 174 ( Stenus ; notes; Ukraine). — Hromádka, 1993: 45 ( Stenus ; subgenus Hypostenus ; Czech Republic; Slovakia). — Puthz and Zanetti, 1995: 19 ( Stenus ; Italy). — Betz, 1996: 25 ( Stenus ; labial adhesion capture apparatus). — Betz, 1998 ( Stenus ; comparative predatory behavior; use of labial apparatus). — Betz, 1999 ( Stenus ; inventory and discussion of feeding, reproduction, grooming, resting, protection behavior). — Jelínek, 1999: 158 ( Stenus ; subgenus Hypostenus ; collecting note; Czech Republic). - DISTRIBUTION: Europe, “Caspian region”.

soma Puthz, 1986 h: 6 ( Stenus ; subgenus Parastenus ; Type locality: Nepal: Provinz Bagmati: Jangtang Ridge N.E. Barahbise, 3250 m). - DISTRIBUTION: Nepal.

sondaicus Bernhauer, 1911: 58 ( Stenus ; subgenus Hypostenus ; Type locality: Sumatra: Si Rambé). — Cameron, 1930d: 326 ( Stenus ; subgenus Hypostenus ; Sumatra). — Scheerpeltz, 1933: 1183 ( Stenus ; subgenus Hypostenus ; catalog). — Puthz, 1970n: 306 ( Stenus ; notes). — Rougemont, 1984: 238 ( Stenus ; Bali). — Herman, 2001: 33 ( Stenus ; synonymic note). - DISTRIBUTION: Indonesia.

plicatus Bernhauer, 1926a: 127 ( Stenus ; subgenus Hypostenus ; Type locality: Sumatra: Brashagi). — Cameron, 1930d: 327 ( Stenus ; subgenus Hypostenus ; Sumatra). — Scheerpeltz, 1933: 1181 ( Stenus ; subgenus Hypostenus ; catalog). — Puthz, 1970n: 306 ( Stenus ; synonym of sondaicus ). — Rougemont, 1984: 238 ( Stenus ; synonym of sondaicus ).

gibbifrons L. Benick, 1928c: 459 ( Stenus ; Type locality: Bandar Baro, Sumatra, East Coast, 850 metres). — L. Benick, 1931: 138 ( Stenus ; notes). — Scheerpeltz, 1933: 1178 ( Stenus ; subgenus Hypostenus ; catalog). — Puthz, 1970n: 306 ( Stenus ; synonym of sondaicus ). — Rougemont, 1984: 238 ( Stenus ; synonym of sondaicus ).

gibbifrons L. Benick, 1928b: 243 ( Stenus ; [preoccupied]; Type locality: Bandar Baro, Sumatras, O.-K., 850 metres). — Herman, 2001: 33 ( Stenus ; synonym of sondaicus ).

prahoeensis L. Benick, 1938: 41 ( Stenus ; subgenus Hypostenus ; Type locality: Java: G. Goentoer; Batoerraden, G. Slamat; G. Patoeha, Preanger, 5000 Voet; G. Tangkoeban Prahoe, Preanger, 4000–5000 Voet). — Puthz, 1970n: 306 ( Stenus ; synonym of sondaicus ).

sordidatus Puthz, 1998e: 94 ( Stenus ; subgenus Parastenus ; Type locality: Ukraine: Karpaten: Svidovec).

- DISTRIBUTION: Ukraine.

sordidipes Puthz, 1988: 661 ( Stenus ; subgenus Parastenus ; Type locality: Sabah: Kinabalu National Park, Layang Layang, 2610 m).

- DISTRIBUTION: Malaysia.

sordidus Puthz, 1988b: 152 ( Stenus ; subgenus Nestus ; Type locality: Neufundland: 3 mi. SE L’Anse-au-Meadow. Also cited from Northwest Territories, Alberta, Manitoba, Ontario, Quebec, and Vermont and subfossil in Alaska).

— Campbell and Davies, 1991: 112 ( Stenus ; Canada).

— Downie and Arnett, 1996: 495 ( Stenus ; characters; USA).

- DISTRIBUTION: Canada, USA.

spangleri Puthz, 1971 k: 141 ( Stenus ; subgenus Hypostenus ; Type locality: Kenya:

Lake Naivasha).

- DISTRIBUTION: Kenya.

spanglerianus Puthz, 1984: 137 ( Stenus ; subgenus Hypostenus ; Type locality: Guatemala:

20 miles S. Guatemala City. Also cited from Costa Rica and Mexico).

- DISTRIBUTION: Costa Rica, Guatemala, Mexico.

sparsepilosus Puthz, 1969a: 66 ( Stenus ; subgenus Hypostenus ; Type locality: Haldwani District, Kumaon).

- DISTRIBUTION: India.

sparsus Fauvel, 1874: 333 [= 1874b: 39] note ( Stenus ; Type locality: Corse; [ Note : See lectotype designation by Puthz, 1969]).

— Sainte-Claire Deville, 1906: 84 ( Stenus ; Corsica).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog; Spain).

— L. Benick, 1929a: 76 ( Stenus ; subgenus Parastenus ; characters in key).

— Scheerpeltz, 1933: 1197 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1969: 39 ( Stenus ; subgenus Parastenus ; lectotype designation; notes).

— Puthz, 1970c: 56 ( Stenus ; aedeagus).

— Puthz and Zanetti, 1995: 19 ( Stenus ; Italy?).

- DISTRIBUTION: Corsica, Spain.

spathifer Puthz, 1971: 248 ( Stenus ; subgenus Hypostenus ; Type locality: Nigeria: Iseri, Lagos. Also cited from Cameroon).

- DISTRIBUTION: Nigeria.

spathipenis Puthz, 1994f: 397 ( Stenus ; subgenus Hypostenus ; Type locality: Zaire:

Parc National Albert, Mutsora, à 5 km au Sud de Mutwanga, 1200 m).

- DISTRIBUTION: Congo.

speculator Lacordaire, 1835 , see: clavicornis Scopoli, 1763 .

speculifer Fauvel, 1873: 130 [= 1873a: 288] ( Stenus ; Type locality: Hautes Pyrénées, Gavarnie).

— Rey, 1884: 360 [= 1884a: 208] ( Stenus ; subgenus Mesostenus ; characters; France).

— Bernhauer and Schubert, 1911: 185 ( Stenus ; subgenus Hemistenus ; catalog).

— L. Benick, 1929a: 81 ( Stenus ; subgenus Parastenus ; characters in key).

— Portevin, 1929: 404 ( Stenus ; characters; France).

— Puthz, 1969: 40 ( Stenus ; subgenus Parastenus ; notes).

— Puthz, 1974a: 67 ( Stenus ; characters).

- DISTRIBUTION: France.

speculifrons Fauvel, 1877 : xxv ( Stenus ; Type locality: Pampas; [Note: See lectotype designation by Puthz, 1969]. Also cited from Montevideo).

— Lynch, 1884: 341 ( Stenus ; subgenus Astenus ; characters; Argentina).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— Scheerpeltz, 1933: 1197 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1969: 28 ( Stenus ; subgenus Hypostenus ; lectotype designation: Pampas; characters).

— Puthz, 1974m: 136 ( Stenus ; notes).

- DISTRIBUTION: Argentina, Uruguay.

bonariensis Bernhauer, 1912e: 171 ( Stenus ; subgenus Hypostenus ; Type locality: Argentinien: Prov. Buenos Aires).

— L. Benick, 1928d: 47 ( Stenus ; characters).

— Scheerpeltz, 1933: 1174 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1969: 28 ( Stenus ; synonym of speculifrons ).

— Puthz, 1974a: 46 ( Stenus ; synonym of speculifrons ).

— Puthz, 1974m: 136 ( Stenus ; synonym of speculifrons ).

topalianus Scheerpeltz, 1972: 99 ( Stenus ; subgenus Hypostenus ; Type locality: Argentinien: Rio Negro, El Bolson, Fuss des Mt. Piltriquitron, 350 m).

— Puthz, 1974m: 136 ( Stenus ; synonym of speculifrons ).

speculus L. Benick, 1941, see: splendidus Bernhauer, 1914 .

spenceri Puthz, 1983: 4 ( Stenus ; subgenus Hypostenus ; Type locality: Vietnam: Dalat 6 km s., 1400–1500 m. Also cited from Laos).

- DISTRIBUTION: Vietnam.

sphaerops Casey, 1884: 68 ( Stenus ; Type locality: Tyngsborough , Massachusetts).

— Bernhauer and Schubert, 1911: 167 ( Stenus ; catalog).

— Campbell and Davies, 1991: 112 ( Stenus ; Canada).

— Downie and Arnett, 1996: 491 ( Stenus ; characters; USA).

- DISTRIBUTION: USA, Canada.

spiculus Zheng, 1993: 227 ( Stenus ; subgenus Parastenus ; Type locality: China: Sichuan Province: Wolong).

- DISTRIBUTION: China.

spinicollis Puthz, 1971 k: 143 ( Stenus ; subgenus Hypostenus ; Type locality: Abyssinie).

— Rougemont, 1981: 122 ( Stenus ; characters; Ethiopia).

- DISTRIBUTION: Ethiopia.

spinifer Puthz, 1967 j: 45 ( Stenus ; subgenus Hypostenus ; Type locality: Mushumangabo, Nyamuragira, 2075 m).

— Puthz, 1971: 310 ( Stenus ; subgenus Hypostenus ; characters; Congo).

- DISTRIBUTION: Congo.

spinosus L. Benick, 1921c: 193 ( Stenus ; subgenus Hypostenus ; replacement name for parvipennis Bernhauer ).

— L. Benick, 1929: 241 ( Stenus ; subgenus Hypostenus ; characters; Philippines).

— Scheerpeltz, 1933: 1183 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1971u: 533 ( Stenus ; Philippines).

— Puthz, 1985: 95, 96 ( Stenus ; subgenus Hypostenus ; characters).

- DISTRIBUTION: Philippines.

parvipennis Bernhauer, 1919: 361 ( Stenus ; [preoccupied]; Type locality: Luzon:

Mt. Makiling).

— L. Benick, 1929: 241 ( Stenus ; synonym of benicki ).

— Scheerpeltz, 1933: 1183 ( Stenus ; synonym of spinosus ).

— Puthz, 1985: 95 ( Stenus ; synonym of spinosus ).

benicki Bernhauer, 1923c: 63 ( Stenus ; replacement name for parvipennis Bernhauer, 1919 ; [preoccupied]).

— L. Benick, 1929: 241 ( Stenus ; synonym of spinosus ).

— Scheerpeltz, 1933: 1183 ( Stenus ; synonym of spinosus ).

— Puthz, 1985: 95 ( Stenus ; synonym of spinosus ).

spissicollis L. Benick, 1921: 5 ( Stenus ; Type locality: S. Paulo, Brasilien).