Schistocarpha eupatorioides (Fenzl) Kuntze

MONRO, A. K., SANTAMARÍA-AGUILAR, D., GONZÁLEZ, F., CHACÓN, O., SOLANO, D., RODRÍGUEZ, A., ZAMORA, N., FEDELE, E. & CORREA, M., 2017, A first checklist to the vascular plants of La Amistad International Park (PILA), Costa Rica-Panama, Phytotaxa 322 (1), pp. 448-450 : 448-450

Schistocarpha eupatorioides (Fenzl) Kuntze Senecio phanerandrus Cufod.

Terrestrial herbs or shrubs. Distribution in PILA 0–1800 Terrestrial shrubs. Distribution in PILA 1300–3400 m, m, low elevation mixed forest, mixed forest in transition mixed forest in transition to cloud forest, cloud forest, low to low elevation forest, mixed forest in transition to elevation oak forest, páramo, pasture. Global distribution: cloud forest, cloud forest. Global distribution: Mexico to Costa Rica, Panama. Records: A.K. Monro 4112 (BM, Bolivia. Records: A. Rodríguez 11374 (BM, INB, MO, INB, MO), A.K. Monro 5175 (BM, INB, MO), A.K. PMA). Monro 5179 (BM, INB, MO), A.K. Monro 5312 (BM, INB, MO), A. Rodríguez 10662 (BM, INB, MO, PMA), A. ψ Senecio costaricensis R.M.King Rodríguez 11016 (BM, INB, MO, PMA), B.B. Klitgaard Terrestrial herbs or shrubs.Distribution in PILA2300–3500 760 (BM), B.B. Klitgaard 835 (BM, INB), E. Alfaro 3976 (INB), G. Davidse 25377 (MO), G. Davidse 25466 (MO), ε-PILA Talamancaster minuscula (Cuatrec.) Pruski F. Quesada 1907 (INB), L. Angulo 201 (INB), S. Knapp Terrestrial herbs. Distribution in PILA 3000–3300 m, 9897 (BM, INB), S. Knapp 9955 (BM, INB), S. Knapp high elevation oak forest, páramo. Global distribution: 9975 (BM, INB). Endemic to Panama. Records: Weston 10154 (MO).

Sigesbeckia jorullensis Kunth Talamancaster westonii (Cuatrec.) Pruski Mielilla (CR) . Terrestrial herbs. Distribution in PILA 3100–3200 m, low Terrestrial herbs. Distribution in PILA 800–3500 m, elevation oak forest, high elevation oak forest, páramo. mixed forest in transition to low elevation forest, mixed Global distribution: Endemic to Costa Rica. Records: forest in transition to cloud forest, cloud forest, low Alfaro 3719 (INB), Quesada 1999 (INB),. elevation oak forest, páramo. Global distribution: Mexico to Peru, Bolivia, Greater Antilles. Records: A.K. Monro Verbesina baruensis Hammel & D’Arcy 4161 (BM, INB), A. Picado 8 (CR, INB), B.B. Klitgaard Terrestrial shrubs. Distribution in PILA 3000–3100 m, low 790 (BM, INB), D. Solano 3957 (BM, INB, MO, PMA), elevation oak forest, high elevation oak forest, páramo. G. Davidse 25166 (MO), G. Herrera 3745 (INB, MO), G. Global distribution: Mexico to Panama, Venezuela. Herrera 6117 (CR, INB), M. Grayum 10356 (CR, INB, Records: N. Zamora, Jul. 29, 2008 (Field obs.). MO), S. Knapp 10003 (BM, INB), S. Knapp 10039 (BM, INB). Verbesina fuscasiccans (D’Arcy) D’Arcy Terrestrial shrubs. Distribution in PILA 3000–3100 m, low

Sinclairia polyantha (Klatt) Rydb. elevation oak forest, high elevation oak forest, páramo. Terrestrial or facultative epiphytic shrubs. Distribution Global distribution: Endemic to Panama. Records: A.K. in PILA 0–2200 m, low elevation mixed forest, mixed Monro 5334 (BM, INB, MO), A.K. Monro 4144 (BM, forest in transition to low elevation forest, mixed forest INB, MO). in transition to cloud forest, cloud forest, low elevation oak forest, dense thicket. Global distribution: Mexico to Verbesina oerstediana Benth. Panama. Records: A.K. Monro 4248 (BM), A. Rodríguez Torilla, tora (CR). 10954 (BM, INB, MO, PMA). Terrestrial shrubs. Distribution in PILA 100–2900 m, low elevation mixed forest, mixed forest in transition to low

Smallanthus maculatus (Cav.) H.Rob. elevation forest, mixed forest in transition to cloud forest, Purca, Tora (CR). cloud forest, low elevation oak forest, high elevation oak Terrestrial herbs. Distribution in PILA 1000–2400 forest. Global distribution: Mexico to Panama, Venezuela.

m, mixed forest in transition to low elevation forest, Records: A. Rodríguez, Feb. 25, 2007 (Field obs.). mixed forest in transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Mexico to Verbesina pleistocephala (Donn.Sm.) B.L.Rob. Panama. Records: C. Hamilton 740 ( MO). Terrestrial shrubs or trees. Distribution in PILA – 1700 m. Global distribution: Costa Rica, Panama border. Records :

℮ Sonchus oleraceus L. T. Croat 26997 (MO). Cerraja , Cerrajilla, Lechuguilla ( CR) . Terrestrial herbs. Distribution in PILA 700–3400 m, Verbesina turbacensis Kunth low elevation mixed forest,mixed forest in transition to low Tora ( CR) . elevation forest, mixed forest in transition to cloud forest, Terrestrial shrubs. Distribution in PILA 600–2600 m, cloud forest, low elevation oak forest, páramo. Global low elevation mixed forest, mixed forest in transition to low distribution: USA to Argentina, Greater Antilles , Lesser elevation forest, mixed forest in transition to cloud forest, Antilles , Lesser Antilles , Europe , Africa, Madagascar. cloud forest, low elevation oak forest, high elevation oak Records: C. Hamilton 736 ( MO) . Uses : medicinal. forest. Global distribution: Mexico to Panama, Venezuela. Records: A. Rodríguez 9754 ( INB) , D. Santamaría 3472

Tagetes filifolia Lag. (INB) , D. Solano 2887 (INB), F. Quesada 1897 (INB). Anisillo, Manzanilla (CR). Terrestrial herbs. Distribution in PILA 800–2200 m, Vernonanthura patens (Kunth) H.Rob. mixed forest in transition to low elevation forest, mixed Tuete, Tuetillo, Dawaska, Tuete blanco, Dowás (CR). forest in transition to cloud forest, cloud forest, low Terrestrial shrubs. Distribution in PILA 0–1700 m, elevation oak forest. Global distribution: Mexico to Peru, low elevation mixed forest, mixed forest in transition to Bolivia, Argentina. Records: A. Rodríguez 9813 (INB), low elevation forest, mixed forest in transition to cloud D. Santamaría 3529 (INB), D. Solano 2874 (INB). forest, cloud forest. Global distribution: Mexico to Brazil. Records: A. Soto 1306 (INB). Uses: medicinal.

Vernonia arborescens (L.) Sw. low elevation mixed forest, mixed forest in transition to Tuete, tuetillo (CR). low elevation forest, mixed forest in transition to cloud Terrestrial herbs or shrubs. Distribution in PILA forest. Global distribution: Mexico to Panama. Records: 400–1800 m, low elevation mixed forest, mixed forest M. Alfaro 11 (INB). in transition to low elevation forest, mixed forest in transition to cloud forest. Global distribution: Mexico to Panama, Venezuela, Bolivia, Lesser Antilles. Records: A. BALANOPHORACEAE Rodríguez 11928 (BM, INB, MO, PMA), A. Rodríguez 9762 (INB), A. Rodríguez 9846 (INB), D. Santamaría Corynaea crassa Hook.f. 3742 (INB), D. Santamaría 7469 (PMA), D. Solano Terrestrial herbs. Distribution in PILA 1700–3200 m, 2844 (INB), D. Solano 5113 (BM, INB, MO, PMA), M. cloud forest, low elevation oak forest, páramo. Global Moraga 682 (INB). distribution: Costa Rica to Bolivia, Venezuela. Records: A. Fernández 670 (CR, INB, MO), A. Fernández 825

Viguiera sylvatica Klatt (CR, INB) , A. Fernández 826 (INB), A. Fernández 826 Terrestrial shrubs. Distribution in PILA 100–2200 m, low (INB, MO), E. Alfaro 3225 (INB), E. Alfaro 4657 (INB), elevation mixed forest, mixed forest in transition to low G. Davidse 25205 (CR, MO), G. Davidse 25847 (CR, elevation forest, mixed forest in transition to cloud forest, MO), G. Davidse 29196 (CR, MO), J. Bittner 1884 (INB, cloud forest, low elevation oak forest. Global distribution: MO), J. Bittner 1929 (INB, MO). El Salvador, Costa Rica, Panama. Records: C. Hamilton 3666 (MO). Helosis cayanensis (Sw.) Spreng. Terrestrial herbs. Distribution in PILA 0–2400 m, low

Werneria nubigena Kunth elevation mixed forest, mixed forest in transition to Terrestrial herbs. Distribution in PILA 3200–3300 m, low elevation forest, mixed forest in transition to cloud páramo. Global distribution: Mexico, Guatemala, Costa forest, cloud forest, low elevation oak forest. Global Rica to Bolivia. Records: A.K. Monro 5184 (BM, MO), distribution: Mexico to Bolivia, Greater Antilles, Lesser L.D. Gómez 22448 (MO). Antilles. Records: A. Fernández 774 (INB), A. Fernández 827 (INB), A.K. Monro 5948 (PMA), A. Rodríguez 11860

Westoniella eriocephala (Klatt) Cuatrec. (BM, INB, MO, PMA) , D. Solano 3947 (BM, INB, MO, Terrestrial herbs or shrubs. Distribution in PILA 3100– PMA), D. Solano 4268 (BM, INB, MO, PMA), E. Alfaro 3500 m, high elevation oak forest, páramo. Global 5575 (BM), F. González 649 (BM, INB, MO, PMA), J. distribution: Endemic to Costa Rica. Records: G. Davidse Bittner 1916 (INB, MO), J. Lezcano 353 (PMA), L. Acosta 26043 (CR, MO). 2384 (INB), L.D. Gómez 23103 (MO), V. H. Ramírez 627 (INB).

□ Westoniella kohkemperi Cuatrec. Terrestrial herbs to shrubs. Distribution in PILA 3400– Langsdorffia hypogaea Mart. 3800 m, páramo. Global distribution: Costa Rica, Panama. Terrestrial herbs. Distribution in PILA 1100–2300 m, Records: A.K. Monro 4089 (BM, INB), A.K. Monro 5361 mixed forest in transition to low elevation forest, mixed (BM, INB), B.B. Klitgaard 825 (BM, INB). forest in transition to cloud forest, cloud forest, low elevation oak forest. Global distribution: Mexico to

Westoniella lanuginosa Cuatrec. Paraguay. Records: S. Knapp 1502 (MO). Terrestrial herbs. Distribution in PILA 3200– 3200 m, low elevation oak forest, páramo. Global distribution: Costa BALSAMINACEAE Rica , Panama. Records: A. S. Weston 10169 (MO), L.D. Gómez 22471 (MO). Impatiens turrialbana Donn. Sm. China de monte (CR).

℮ Youngia japonica (L.) DC. Terrestrial herbs. Distribution in PILA 900–3000 m, Terrestrial herbs. Distribution in PILA 600–2200 m, low mixed forest in transition to low elevation forest, mixed elevation mixed forest, mixed forest in transition to low forest in transition to cloud forest, cloud forest, low elevation forest, mixed forest in transition to cloud forest, elevation oak forest, high elevation oak forest. Global cloud forest, low elevation oak forest. Global distribution: distribution: Costa Rica, Panama. Records: A. Soto 309 USA to Venezuela, Guyanas, Greater Antilles, Lesser (INB), E. Alfaro 3489 (INB), G. Davidse 25808 (CR, Antilles, Africa, Madagascar. Records: E. Navarro 184 MO), G. Davidse 28645 (CR, MO), G. Herrera 3635 (INB). (INB, MO), G. Herrera 3735 (INB, MO), G. Herrera 6250 (INB, MO), F. Quesada 1495 (CR, INB), L. Angulo

Zexmenia virgulta Klatt 217 (CR, INB), M.A. Thomas 1658 (MO), M. Moraga Terrestrial shrubs. Distribution in PILA 200–1700 m, 401 (INB).

elevation mixed forest, mixed forest in transition to low

℮ Impatiens walleriana Hook. f. elevation forest, mixed forest in transition to cloud forest, China, China, Sultana (CR) . cloud forest, low elevation oak forest, high elevation oak

Terrestrial herbs. Distribution in PILA 800–1800 m, forest, pasture, dense thicket. Global distribution: Mexico mixed forest in transition to low elevation forest, mixed to Peru, Greater Antilles, Lesser Antilles. Records: A.K. forest in transition to cloud forest. Global distribution: Monro 5737 (BM, INB, MO, PMA), D. Santamaría 5972 Africa, introduced in Mexico, Nicaragua, Panama, (BM, INB, MO, PMA), D. Santamaría 7419 (PMA), D. Ecuador, Africa, Asia. Records: C. Hamilton 722 (MO). Solano 4323 (BM, INB, MO, PMA), E. Alfaro 17 (MO),

J. Lezcano 343 (PMA). BEGONIACEAE Begonia heydei C.DC.

Terrestrial herbs. Distribution in PILA 1000–1600 m,

Begonia carletonii Standl. mixed forest in transition to low elevation forest. Global Terrestrial herbs. Distribution in PILA 100–400 m, low distribution: Guatemala to Panama.Records: A. Rodríguez elevation mixed forest. Global distribution: Costa Rica, 11233 (BM, INB, MO, PMA), A. Soto 315 (INB), G. Panama. Records: A. Rodríguez 11420 (BM, INB, MO, Herrera 8108 (CR). PMA). Begonia involucrata Liebm.

Begonia convallariodora C.DC. Terrestrial herbs. Distribution in PILA 1300–2700 m, Terrestrial herbs. Distribution in PILA 400–2800 m, low mixed forest in transition to cloud forest, cloud forest, elevation mixed forest, mixed forest in transition to low low elevation oak forest, high elevation oak forest. Global elevation forest, mixed forest in transition to cloud forest, distribution: Guatemala to Panama.Records: A. Rodríguez cloud forest, low elevation oak forest, high elevation oak 11757 (BM, INB, MO, PMA), D. Santamaría 7266 (BM,

forest. Global distribution: Mexico to Panama. Records: INB, MO, PMA), D. Solano 5048 (BM, INB, MO, PMA), A. Fernández 897 (INB), A. Picado 3 (CR, INB), A. D. Solano 5158 (BM, INB, MO, PMA), D. Solano 5170 Rodríguez 10762 (BM, INB, MO, PMA), A. Rodríguez (BM, INB, MO, PMA), N. Zamora 4062 (BM, INB, MO, 10906 (BM, INB, MO, PMA), A. Rodríguez 11929 (BM, PMA). Uses: medicinal. INB, MO, PMA), C. Hamilton 3648 (MO), D. Santamaría 5831 (BM, INB, MO, PMA), D. Santamaría 7092 (BM, ε-PILA Begonia mucronistipula C.DC. INB, MO, PMA ), D. Solano 3948 (BM, INB, MO, PMA), Epiphytic herbs. Distribution in PILA 1200–2100 m, E. Alfaro 5497 (BM, INB, MO), G. Gallardo 83 (INB). mixed forest in transition to low elevation forest, mixed forest in transition to cloud forest, cloud forest, low

Begonia cooperi C.DC. elevation oak forest. Global distribution: Endemic to Ala de ángel (CR). Panama. Records: T. Cochrane et al. 6272 (MO), W.

Terrestrial herbs to shrubs. Distribution in PILA D’Arcy 10106 (MO), J. Dwyer et al. 441 (MO), J. Folsom 1000–1600 m, mixed forest in transition to low elevation et al. 2256 (MO), B. Hammel 5682 (MO), W. Hoover 554 forest. Global distribution: Costa Rica, Panama. Records: (MO), H. Pittier 3172 ( US), R. Wilbur & j. Luteyn 19247 D. Santamaría 5642 (BM, INB, MO, PMA), J. Bittner (DUKE), R. Woodson & R. Schery 620 (MO). 1752 (INB, MO). Begonia oaxacana A.DC.

Begonia copeyana C.DC. Terrestrial herbs. Distribution in PILA 1400–3200 m,

Terrestrial herbs. Distribution in PILA 1300–2200 m, mixed forest in transition to cloud forest, cloud forest, mixed forest in transition to cloud forest, cloud forest, low elevation oak forest, páramo. Global distribution: low elevation oak forest. Global distribution: Endemic Mexico to Panama. Records: A.K. Monro 4063 ( BM,

to Costa Rica. Records: D. Solano 3831 (BM, INB, MO, INB, MO), A.K. Monro 5565 (BM, INB, MO, PMA), PMA). D. Solano 3953 (BM, INB, MO, PMA), E. Alfaro 3240

(INB), G. Davidse 25098 (CR, MO), G. Davidse 25765

Begonia estrellensis C.DC. (MO) , G. Davidse 25858 (CR, MO), G. Davidse 26139 Facultatively epiphytic herbs. Distribution in PILA (CR, MO), G. Davidse 28652 (CR), G. Davidse 28687 700–2000 m, low elevation mixed forest, mixed forest in (MO), G. McPherson 15906 (MO), J. Bittner 1897 (INB, transition to low elevation forest, mixed forest in transition MO), F. Quesada 1493 (INB), F. Quesada 1911 (INB),

to cloud forest. Global distribution: Costa Rica, Panama. M.A. Thomas 1607 (MO), M. Grayum 10311 (INB, MO), Records: G. Davidse 28323 (CR). S. Knapp 9984 (BM, INB).