Systaria lanna, Dankittipakul, Pakawin & Singtripop, Tippawan, 2011

Dankittipakul, Pakawin & Singtripop, Tippawan, 2011, Seven new species of Systaria Simon, 1897 from Southeast Asia (Araneae, Clubionidae, Systariinae), Zootaxa 2905, pp. 16-32 : 23

Systaria lanna sp. nov.

Figs 15–19

Type material. Holotype: 3, THAILAND: Mae Hong Son Province: Tham Lod [= incorrect transliteration by collectors as Tham Lok], 5 km north of Mae Lang, 14 November 1985, leg. B. & I. Löbl ( MHNG, Sys-01).

Paratypes: 13, same data as holotype ( THNM, Sys-01). THAILAND: Mae Hong Son Province: 1Ƥ, Pang Ma Pha District, Tham Lod, 450 m, cave entrance, 31 December 2006, leg. P. Dankittipakul ( MHNG, Sys-02). Chiang Mai Province: 2Ƥ, Chiang Dao District, Tham Krab, 400 m, cave entrance, 23 December 2004, leg. P. Dankittipakul ( MHNG, Sys-03; THNM).

Diagnosis. Males of S. lanna sp. nov. are recognized by the hook-like process on the complex RTA ( Figs 15– 17 ); females by the elongate-ovoid anterior bursae, and the elongate-ovoid posterior spermathecae ( Fig. 19 ).

Etymology. The specific epithet is a noun in apposition, referring to the old independent state of Lanna (modern northern Thailand) where the types of this new species were collected.

Description. Male (holotype): Total length 11.0. Prosoma 5.02 long, 3.56 wide. Opisthosoma 5.98 long, 3.30 wide. Leg formula: 4123; I 10.24 (2.92, 1.56, 2.84, 1.88, 1.04); II 9.66 (2.86, 1.42, 2.58, 1.82, 0.98); III 9.16 (3.14, 0.98, 1.96, 2.16, 0.92); IV 13.58 (3.82, 1.48, 2.98, 3.52, 1.78). Spination. Femora: I d1-1-1, p1-0-1, r1-0-1; II d1-1- 1, p1-1-0, r0-1-2; III d1-1-1, p1-0-1, r1-1-1; IV d1-1-1, p1-1-1, r0-0-1. Tibiae: I v2 -0-2; II v2-2 -0; III p1-1-0, r1-0- 1, v2-2 -2; IV p1-1-0, r1-0-1, v2-2 -2. Metatarsi: II v1 -0-0; III p1-1-1, r1-1-2, v1-1 -2; IV p1-1-2, r1-1-2, v2-2 -2. Eye size and interdistances: AME 0.16, ALE 0.12, PME 0.12, PLE 0.10; AME–AME 0.18, AME–ALE 0.16, PME– PME 0.24, PME–PLE 0.18, ALE–PLE 0.14; MOA 0.26 long, anterior width 0.38, posterior width 0.36.

Palp ( Figs 15–17 ). Ventral surface of tibia proximally with distal swelling ( Fig. 16 ). RTA complex, with sharply pointed, posteriorly directed process ( Fig. 15 ) and dorsal flange ( Figs 16–17 ). Cymbial fold partially membranous. Cymbial ridge slightly elevated. Basal part of embolus triangular; embolic base with broad membranous area ( Fig. 15 ); distal part of embolus filiform. Conductor broad, concave in lateral view ( Fig. 16 ).

Female (paratype, MHNG, Sys-02): Total length 10.82. Prosoma 4.94 long, 3.52 wide. Opisthosoma 5.88 long, 3.22 wide. Leg formula: 4123; I 9.88 (2.78, 1.52, 2.72, 1.88, 0.98); II 8.92 (2.72, 1.32, 2.40, 1.56, 0.92); III 8.92 (3.04, 0.98, 1.92, 2.02, 0.96); IV 13.66 (3.86, 1.48, 2.96, 3.54, 1.82). Spination. Femora: I d1-1-1, p1-1-0, r0-1-1; II d1-1-1, p0-1-1, r0-1-2; III d1-1-1, p1-1-0, r1-1-1; IV d1-1-1, p1-1-1, r0-0-1. Tibiae: I v2-1 -1; II v0-1-1; III p1-1-0, r1-1-0, v2-2 -2; IV p1-0-1, r1-1-0, v2-2 -1. Metatarsi: II v1 -0-0; III p1-1-1, r1-1-1, v2-1 -1; IV p1-1-2, r1-1-2, v2-2 -2. Eye size and interdistances: AME 0.16, ALE 0.12, PME 0.12, PLE 0.10; AME–AME 0.18, AME–ALE 0.16, PME–PME 0.24, PME–PLE 0.18, ALE–PLE 0.14; MOA 0.26 long, anterior width 0.38, posterior width 0.36.

Genitalia ( Figs 18–19 ). Epigynal atrium oval, broad, its margin ill-defined posteriorly. Copulatory orifices situated on posterior margin of atrial border ( Fig. 18 ). Anterior bursae convergent, elongate-ovoid, heavily sclerotized ( Fig. 19 ). Insemination ducts plugged with dark material in the described specimen, spermathecae bacilliform. Fertilization ducts subulated.

Natural history. Types of S. lanna sp. nov. were directly taken from their retreats on the wall at the cave entrance.

Distribution. Northern Thailand (Fig. 43).

Figures

FIGURES 15 – 19. Systaria lanna sp. nov., holotype male (15 – 17) and paratype female (18 – 19). 15 – 17 Left male palp (15 ventral, 16 retrolateral, 17 dorsal); 18 Epigyne, ventral; 19 Vulva, dorsal. Scale lines = 0.1 mm.