Sycophila (Sycophila) zebra Lotfalizadeh, 2024
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publication ID |
https://doi.org/ 10.11646/zootaxa.5401.1.1 |
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publication LSID |
lsid:zoobank.org:pub:376CAB04-43BE-4C9A-950B-80DBD676CFB0 |
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persistent identifier |
https://treatment.plazi.org/id/03FD6B5E-7248-61AD-5486-B4CDFA2FF87C |
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treatment provided by |
Plazi |
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scientific name |
Sycophila (Sycophila) zebra Lotfalizadeh |
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sp. n. |
Sycophila (Sycophila) zebra Lotfalizadeh & Rasplus sp. n.
( Figs 90–92 View FIGURE 90 View FIGURE 91 View FIGURE 92 )
Type material examined. Holotype ♀: United Republic of Tanzania, Soni, Mt Usambara West, 18.ii.1995, ex syconia of Ficus sur , Rasplus J.-Y. & Kerdelhué C. leg. Deposited in CBGP. Paratypes: same data as holotype, 5♀. Mialo, Mt Usambara West, ex syconia of Ficus vallis-choudae , 20.ii.1995, Rasplus J.-Y. & Kerdelhué C. leg., 7♀. Republic of Côte d’Ivoire, Lamto, 6.222197° -5.024070°, ex syconia of Ficus sur , 15.v.1994, Rasplus J.-Y. leg., 32♀ & 9♂. Cameroon, Nkolbisson, ex syconia of Ficus sycomorus , 17.iii.1993, Gibernau M. leg., 5♀. Mt Febe, ex syconia of Ficus sur , 19.ii.1993, Gibernau M. leg., 6♀ & 2♂. Guinea, Monts Nimba, Mifergui, ex syconia of Ficus sur , 17.iv.1993, Rasplus J.-Y. leg., 1♀.
Etymology. The specific name refers to striped colouration of the female gaster.
Diagnosis. Female body mainly brown with some yellowish bands, especially on tergites ( Fig. 90A View FIGURE 90 ). Funiculars short (fu2–fu5), as long as wide ( Fig. 90B View FIGURE 90 ). Costal cell ventrally densely setose, dorsally with a row of setae ( Fig. 91E View FIGURE 91 ). Petiole at most as long as broad dorsally ( Fig. 91D View FIGURE 91 ).
Male body mainly yellowish, with brown tergites ( Fig. 92A View FIGURE 92 ). Flagellum with fu2–fu4 at most slightly longer than wide, fu5 wider than long ( Fig. 92B View FIGURE 92 ).
Description of holotype ♀ ( Figs 90–91 View FIGURE 90 View FIGURE 91 ). Length 4.28mm. Body yellowish ( Fig. 90A View FIGURE 90 ), except head dorsally, antennal scrobe, clypeus, mesosoma dorsally, propodeum, mesopleuron anteriorly, metapleuron, tergites dorsally and posteriorly dark brown, antennal flagellum and pedicel yellowish-brown; pronotum anterolaterally testaceous; fore wing with a dark brown spot below mv, reaching base of mv, lower margin of mv not obscured by dark brown spot ( Fig. 91E View FIGURE 91 ); gaster yellowish with at least posterior third of the tergite dark brown, so it appears striped, dorsal edge with a dark brown band ( Fig. 91C View FIGURE 91 ); body setation whitish, setation of fore wing entirely dark.
Head in frontal view densely punctulate, in dorsal view coriaceous and sparsely punctulate; in frontal view 1.4× as broad as high [HW 182 & HH 130] ( Fig. 90C View FIGURE 90 ); in dorsal view stout, 1.79× as broad as long [HW 182 & HL 102] ( Fig. 90D View FIGURE 90 ); malar space 0.5× length of eye [ms 50 & EH 100]; malar sulcus distinct, straight ( Fig. 90E View FIGURE 90 ); genal margin regularly curved, edged but not carinate; antennal toruli widely separated; interantennal space not raised, carinate laterally; scrobal depression smooth, reaching the median ocellus, carinate on margin (at least dorsally); POL 1.82× as long as OOL [POL 40 & OOL 22] ( Fig. 90D View FIGURE 90 ); temple rounded, 0.27× the eye length (in dorsal view) [TL 24 & EL 90]; setation oriented downwards on face, outwards on the punctuate frons.
Antenna. Scape not reaching median ocellus, 2.05× as long as broad [SL 82 & SW 20]; fu1 1.52× as long as broad [fu1L 32 & fu1W 21], distinctly longer than pedicel, not bulging dorsally; funiculars (fu2–fu4) with one row of mps and 2–3 whorls of adpressed setae; fu2–fu4 not longer than broad; fu5 as not longer than broad ( Fig. 90B View FIGURE 90 ).
Mesosoma weakly arched dorsally, 1.75× as long as wide (175:100), with deep sculpture ( Fig. 91A View FIGURE 91 ); pronotum with coarse umbilicate punctures; mesepisternum raised medially, with piliferous sculpture on epicnemium, being closer to anteroventral margin of mesepisternum than to mesocoxa; mesoscutum with dense umbilicate punctures, transversely strigose anteriorly; notauli conspicuous, slightly obliterated by mesoscutum sculpture; prepectus bare, with raised sculpture; mesoscutum 0.6× as long as wide (70:110), 0.8× longer than length of pronotal collar (70:80), distinctly umbilicate laterally and with coriaceous sculpture on mesoscutellar disc; mesepimeron transversally strigose; metepimeron subtriangular and areolate with long white hairs; propodeum ( Fig. 91B View FIGURE 91 ) with deep median depression, with a smooth median band, without any median carina; lateral rows of areolae short (extending at most to half-length of propodeum); basal carina inverted V, truncate mesally; lateral branches of propodeal basal carina slightly convex; distance between lateral carinae less than 0.5× width of nucha.
Fore wing with short marginal fringe, apically subtruncate (pointed anteriorly); dark brown spot only behind mv; basal cell bare or nearly so; mv distinctly enlarged, rounded, lower margin convex, not longer than pmv; stv not curved; pmv distinct, longer than stv; ventral side of costal cell densely setose, dorsally bare; speculum present; parastigmal bristles absent ( Figs 91 E, F View FIGURE 91 ).
Legs. Fore coxa without depression on frontal surface, hind coxa bare basodorsally; fore (45:15) and hind femora (55:20); hind tibia dorsally with more than 10 setae, setae shorter than hind tibia width.
Metasoma. Petiole shorter than broad dorsally, ventral transverse carina between petiole and St1 present. Metasoma 1.7× as long as wide (170:100) ( Fig. 91C View FIGURE 91 ), longer than mesosoma, strongly compressed laterally and dorsally angulate ( Fig. 91D View FIGURE 91 ); ventral transverse carina between petiole and St1 present; petiole in dorsal view not longer than wide, shorter than half length of hind coxa; tergites smooth; Gt4 (in lateral view) slightly as long as Gt3, sinuous posteriorly; Gt5 partly covering Gt6, sinuous posteriorly; Gt5 with few setae dorsally, Gt6 sparsely setose ventrally; ovipositor oriented dorsad but not distinctly upturned ( Fig. 91C View FIGURE 91 ).
Male ( Fig. 92 View FIGURE 92 ). Length 2.42–2.50 mm. Similar to female with longer petiole ( Fig. 92A View FIGURE 92 ); but with a yellowish colouration, with funiculars brown, gaster dark brown; antennal formula 11143; each funicular with 1–2 rows of mps and 3 whorls of relatively short erected setae ( Fig. 92B View FIGURE 92 ); fore wing more setose than in female, speculum present, costal cell dorsally distinctly setose, ventrally sparsely setose ( Fig. 92D View FIGURE 92 ).
Variation. Length 2.20–6.30 mm. Brown lateral bands on tergites may vary in size.
Host plants. Ficus sur and F. vallis-choudae .
Geographical distribution. Cameroon, Guinea, Republic of Côte d’Ivoire and United Republic of Tanzania.
Subgenus Tineomyza Rondani, 1872 stat. rev.
Type species: Tineomyza pistacina Rondani, 1872 View in CoL by monotypy.
Subgeneric diagnosis. In female flagellum with fu1 at most as long as pedicel (except in kestraneura View in CoL species-group), not curved. Epicnemium flattened. Fore wing with mv slightly widened, ventrally not convex, smv straight, pmv distinct. Propodeum slightly depressed medially. Petiole long or short, ventral transverse carina between petiole and St1 absent or present.
Description. Size and colour. Length varies between 1.6–6.6 mm in female and 1.8–6.2 mm in male; body entirely yellowish to brown or black, sometimes with different colouration; fore wing with a dark brown spot below mv, reaching base of mv or not, lower margin of mv obscured by dark brown spot or not; tergites uniformly coloured or with bands yellowish to dark coloured.
Head coriaceous to punctured, in frontal view 1–1.4× as broad as high, in dorsal view from 1.6× (stout) to 2× (transverse) as broad as long; malar space less than half-length of eye; malar sulcus absent or present; genal margin regularly curved, edged but not carinate; antennal toruli widely or narrowly separated; interantennal space generally not raised and in lateral view, carinate laterally or not; scrobal depression smooth or slightly reticulate, not carinate; POL more than 2× as long as OOL (1.5× in Sycophila (T.) platygastra sp. n. and 1.9× in S. (T.) wiebesi sp. n.), temple generally round, 0.14 to 0.45× the eye length (in dorsal view).
Antenna. Scape reaching median ocellus or not, more than 3× as long as broad; fu1 several times as long as broad, longer than pedicel or not, not bulging dorsally; funiculars longer than broad or not, with one to three rows of mps.
Mesosoma with sculpture varying from coriaceous to punctuated; mesepisternum without any process or raised sculpture on epicnemium; mesoscutum sparsely umbilicate, smooth or with sculpture varying from coriaceous to strigose or umbilicate anteriorly; prepectus bare (except in punctum species-group); sculpture of mesoscutellum varying from coriaceous to entirely umbilicate punctures; propodeum with shallow median depression, with or without smooth median band, median carina and rows of areolae; basal carina inverted V-, U- shaped or truncate mesally.
Fore wing with short or long marginal fringe, apically rounded or truncate; dark brown spot below mv small to longer than wide; basal cell bare or setose; mv slightly broadened, not convex in lower margin; stv not curved; pmv distinct, longer than stv or not; costal cell ventrally and dorsally bare to setose; speculum present; parastigmal bristles generally absent except in binotata species-group.
Metasoma generally longer than mesosoma, slightly to distinctly compressed laterally; ventral transverse carina between petiole and St1 present; petiole in dorsal view short to long, generally longer than wide and shorter than hind coxa; tergites smooth to finely sculptured, in some case Gt 4 in lateral view much as long as Gt3 (at least 1.5× longer), sinuous posteriorly or not; ovipositor distinctly to slightly upturned.
Comments. We reinstated Tineomyza as a valid subgenus of Sycophila based on characters listed here above and comparison with the type species of Sycophila . In contrast to Zerova (1995) who separated Sycophila and Tineomyza based on the shape of femora and on their biological association ( Ficus versus Quercus and others), we concluded that the two taxa only deserve a subgeneric status and observed that species of Sycophila subgenus Tineomyza can also be associated with Ficus .
We hereafter describe 11 new species that belong to the subgenus Tineomyza and transfer seven other species to this subgenus: Sycophila (Tineomyza) flaviclava Bouček, 1981 , Sycophila (T.) modesta Bouček, 1981 , Sycophila (T.) naso Bouček, 1981 , Sycophila (T.) punctum Bouček, 1981 , Sycophila (T.) ruandensis Risbec, 1957 , Sycophila (T.) sessilis Bouček, 1981 , Sycophila (T.) setulosa Zerova, 2009 .
Based on our observation of European species, we hypothesize that most species associated with Cynipidae ( Hymenoptera ) and Cecidomyiidae ( Diptera ), could be classified in subgenus Tineomyza all species. However, this must be validated by analyzing subgeneric characters for all described species.
We also defined six species-groups for the species studied: binotata group, platygastra group, modesta group, naso group, punctum group and sessilis group. These groups were erected mostly based on characters proposed by Bouček et al. (1981) for specific identification of S. kestraneura , S. modesta , S. naso , S. punctum and S. sessilis .
Key to Afrotropical species-groups and species of Sycophila (Tineomyza) associated with Ficus View in CoL
1 Antenna with 5 funiculars, funiculars with short and appressed hairs ( Figs 93B View FIGURE 93 , 95B View FIGURE 95 , 97E View FIGURE 97 , 100B View FIGURE 100 ); metasoma longer than mesosoma, with petiole relatively short, at most as long as hind coxa ( Figs 100A View FIGURE 100 ., 105A View FIGURE 105 , 107A View FIGURE 107 ).......... go to female key
- Antenna with 4 funiculars, funiculars with relatively long and erect hairs ( Figs 99B View FIGURE 99 , 102B View FIGURE 102 ); metasoma shorter than mesosoma, petiole longer than hind coxa ( Figs 99A View FIGURE 99 , 117A View FIGURE 117 , 123A View FIGURE 123 )............................................. go to male key
Female
1 Fore wing with two dark spots below mv ( Fig. 94E View FIGURE 94 ), parastigmal bristles present............................................................................................ binotata View in CoL species group [ S. (T.) setulosa Zerova, 2009 )]
- Fore wing only one dark spot below mv or not; parastigmal bristles absent ( Figs 96E View FIGURE 96 , 98E View FIGURE 98 , 101E View FIGURE 101 )..................... 2
2(1) Flagellum with fu1 as long as or shorter than pedicel ( Figs 124B View FIGURE 124 , 127B View FIGURE 127 , 130B View FIGURE 130 ).................................... 3
- Flagellum with fu1 distinctly longer than pedicel ( Figs 115B View FIGURE 115 , 118B View FIGURE 118 , 121B View FIGURE 121 )................... platygastra species group 6
3(2) Prepectus bare ( Figs 93F View FIGURE 93 , 95F View FIGURE 95 , 97F View FIGURE 97 ); fore wing with dark brown spot reaching ( Figs 96F View FIGURE 96 , 98F View FIGURE 98 ); or not reaching base of mv (in sessilis View in CoL group) ( Figs 131F View FIGURE 131 , 137F View FIGURE 137 )......................................................................... 4
- Prepectus pilose ( Figs 124F View FIGURE 124 , 127F View FIGURE 127 ); fore wing with dark brown spot not reaching base of mv ( Figs 125F View FIGURE 125 , 128F View FIGURE 128 )............................................................................................ punctum View in CoL species-group 9
4(3) Petiole shorter than broad, distinctly shorter than hind coxa ( Figs 131C, D View FIGURE 131 , 134C, D View FIGURE 134 , 137C, D View FIGURE 137 ); dark brown spot of fore wing not reaching the base of mv ( Figs 125F View FIGURE 125 , 128F View FIGURE 128 )............................................ sessilis View in CoL species-group 9
- Petiole longer than broad, as long as hind coxa ( Figs 96C View FIGURE 96 , 98C–D View FIGURE 98 , 101C–D View FIGURE 101 ); dark brown spot of fore wing reaching base of mv ( Figs 96F View FIGURE 96 , 98F View FIGURE 98 )....................................................................................... 5
5(4) Costal cell dorsally with a row of setae, fore wing round apically ( Figs 104E View FIGURE 104 , 106E View FIGURE 106 ); all funiculars longer than broad ( Figs 95B View FIGURE 95 , 97E View FIGURE 97 ); Gt4 much as long as Gt3 or not ( Figs 96C View FIGURE 96 , 98C View FIGURE 98 , 101C View FIGURE 101 )............................... modesta View in CoL species-group 11
- Costal cell without row of setae, fore wing not round apically ( Figs 104E View FIGURE 104 , 106E View FIGURE 106 , 108E View FIGURE 108 ); Funiculars long or as long as broad ( Figs 103B View FIGURE 103 , 105B View FIGURE 105 , 107B View FIGURE 107 ); Gt4 much as long as Gt3 ( Figs 104C View FIGURE 104 , 105C View FIGURE 105 , 108C View FIGURE 108 ).................... naso View in CoL species-group 13
6(2) Costal cell densely setose ventrally ( Fig. 122E View FIGURE 122 ).......................... S. (T.) wiebesi Lotfalizadeh & Rasplus sp. n.
- Costal cell sparsely setose ventrally ( Figs 116E View FIGURE 116 , 119E View FIGURE 119 )........................................................ 7
7(6) Smooth median band of propodeum elongate, reaching the posterior margin, basal carina not wide, less than nucha width, propodeum areolate posteriorly ( Fig. 116B View FIGURE 116 ); ventral area of interantennal projection reticulate ( Fig. 115C View FIGURE 115 )................................................................................ S. (T.) maldesi Lotfalizadeh & Rasplus sp. n.
- Smooth median band of propodeum short, at most reaching basal half of propodeum, basal carina of propodeum wide basally, propodeum posteriorly rugulose transversally ( Fig. 119B View FIGURE 119 ); ventral area of interantennal space smooth ( Fig. 118C View FIGURE 118 )....................................................................... S. (T.) platygastra Lotfalizadeh & Rasplus sp. n.
8(3) Gt3 partly covering Gt4 ( Fig. 125C View FIGURE 125 ), Gt4 sinuous posteriorly ( Fig. 125D View FIGURE 125 ); lateral ocelli far from eyes, distinctly more than ocellus diameter (about 2×) ( Fig. 124D View FIGURE 124 ); propodeum without median longitudinal carina ( Fig. 125B View FIGURE 125 ).................................................................................. S. (T.) busseicola Lotfalizadeh & Rasplus sp. n.
- Gt3 not covering Gt4 ( Fig. 128C View FIGURE 128 ), Gt4 straight posteriorly ( Fig. 128D View FIGURE 128 ); lateral ocelli close to eyes, less than 2× ocellus diameter ( Fig. 127C View FIGURE 127 ); propodeum with a short median longitudinal carina basally ( Fig. 128B View FIGURE 128 )......... S. (T.) punctum Bouček, 1981 View in CoL
9(4) Funiculars slightly longer than wide, F5 at most as long as wide ( Fig. 130B View FIGURE 130 ); malar sulcus absent ( Fig. 130E View FIGURE 130 )........................................................................... S. (T.) beninensis Lotfalizadeh & Rasplus sp. n.
- Funiculars distinctly longer than wide, at least 2× as long as wide; malar sulcus present............................. 10
10(9) Propodeum with a median smooth band ( Fig. 134B View FIGURE 134 ); body mostly dark brown; fore wing with large dark brown spot below mv, the spot slightly extending into the posterior half of the wing ( Fig. 134E View FIGURE 134 )..... S. (T.) luteacola Lotfalizadeh & Rasplus sp. n.
- Propodeum with two rows of large areoles medially ( Fig. 137B View FIGURE 137 ); body mostly yellowish ( Fig. 136A View FIGURE 136 ); fore wing with small dark brown spot below mv, not extending into the posterior half of the wing ( Fig. 137E View FIGURE 137 )............ S. (T.) sessilis Bouček, 1981 View in CoL
11(5) Basal carina of propodeum as an inverted V, truncate mesally ( Fig. 101B View FIGURE 101 )...................................................................................................... S. (T.) zebrogastra Lotfalizadeh & Rasplus sp. n.
- Basal carina of propodeum inverted V-shaped ( Figs 96A View FIGURE 96 , 98B View FIGURE 98 )................................................ 13
12(11) Fore wing with small and less extensive dark brown spot below mv ( Fig. 96E View FIGURE 96 ); front margin of mesoscutum strigose ( Fig. 96B View FIGURE 96 ); dorsellum smooth ( Fig. 96A View FIGURE 96 )...................................................... S. (T.) modesta Bouček, 1981 View in CoL
- Fore wing with large dark brown spot below mv, slightly extending into posterior half of wing ( Fig. 98E View FIGURE 98 ); front margin of mesoscutum smooth ( Fig. 98A View FIGURE 98 ); dorsellum sculptured...................... S. (T.) risbeci Lotfalizadeh & Rasplus sp. n.
13(5) All funiculars longer than broad ( Figs 103B View FIGURE 103 , 107B View FIGURE 107 )......................................................... 14
- At least some funiculars shorter than broad ( Figs 105B View FIGURE 105 , 110B View FIGURE 110 , 112B View FIGURE 112 )........................................... 15
14(13) Flagellum with fu1 obconical, narrower than pedicel, its basal half without longitudinal sensilla ( Fig. 107B View FIGURE 107 ); basal carina of propodeum as an inverted V truncate mesally ( Fig. 108B View FIGURE 108 ); dark brown spot of fore wing not reaching base of mv ( Fig. 108F View FIGURE 108 )................................................................. S. (T.) minuta Lotfalizadeh View in CoL & Rasplus sp. n.
- Flagellum with fu1 not obconical, more or less as wide as pedicel, its basal half with longitudinal sensilla ( Fig. 104B View FIGURE 104 ); basal carina of propodeum inverted V-shaped ( Fig. 105B View FIGURE 105 ); dark brown spot of fore wing reaching base of mv ( Fig. 105E View FIGURE 105 )................................................................................... S. (T.) flaviclava Bouček, 1981 View in CoL
15(13) Front margin of mesoscutum smooth ( Fig. 113A View FIGURE 113 )................................... S. (T.) ruandensis Risbec (1957) View in CoL
- Mesoscutum with coarse umbilicate punctures ( Figs 106A View FIGURE 106 , 111A View FIGURE 111 ).............................................. 16
16(15) Basal carina of propodeum inverted V-shaped ( Fig. 106B View FIGURE 106 )................. S. (T.) glumosae Lotfalizadeh & Rasplus sp. n.
- Basal carina of propodeum inverted V, truncate mesally ( Fig. 111B View FIGURE 111 )......................... S. (T.) naso Bouček, 1981 View in CoL
Male
1 Prepectus pilose ( Figs 126A View FIGURE 126 , 129A View FIGURE 129 )....................................................................... 2
- Prepectus bare........................................................................................ 3
2(1) All funiculars longer than wide, pedicel + funicle longer than head width ( Fig. 126C View FIGURE 126 ); dark brown spot of fore wing longer than wide, basal cell bare medially ( Fig. 126D View FIGURE 126 )............................. S. (T.) busseicola Lotfalizadeh & Rasplus sp. n.
- Only fu1 longer than wide, fu2–fu4 wider than long ( Fig. 129B View FIGURE 129 ), pedicel + funicle shorter than head width; dark brown spot of fore wing wider than long, basal cell sparsely setose ( Fig. 129D View FIGURE 129 )......................... S. (T.) punctum Bouček, 1981 View in CoL
3(1) All funiculars long, longer than wide ( Figs 102B View FIGURE 102 , 109B View FIGURE 109 , 120D View FIGURE 120 )................................................. 4
- Only fu1 longer than wide, fu2–fu4 at most as long as wide ( Figs 114B View FIGURE 114 , 123B View FIGURE 123 , 132B View FIGURE 132 )............................... 9
4(3) Dark brown spot of fore wing short and wider than long ( Figs 102C View FIGURE 102 , 109D View FIGURE 109 )....................................... 5
- Dark brown spot of fore wing longer than wide ( Figs 120A, 120C View FIGURE 120 , 135D View FIGURE 135 )......................................... 6
5(4) Costal cell dorsally bare, ventrally at most with two rows of setae ( Fig. 109D View FIGURE 109 )... S. (T.) minuta Lotfalizadeh View in CoL & Rasplus sp. n.
- Costal cell dorsally with a row of setae, ventrally with more than two rows of setae ( Fig. 102C View FIGURE 102 )..................................................................................... S. (T.) zebrogastra Lotfalizadeh & Rasplus sp. n.
6(4) Costal cell densely setose ventrally ( Figs 120A, C View FIGURE 120 )..................... S. (T.) platygastra Lotfalizadeh & Rasplus sp. n.
- Costal cell sparsely setose ( Fig. 135D View FIGURE 135 ).................................................................... 7
7(6) Body mostly dark brown and partly yellowish ( Fig. 135A View FIGURE 135 )................. S. (T.) luteacola Lotfalizadeh & Rasplus sp. n.
- Body entirely yellowish ( Figs 117A View FIGURE 117 , 99A View FIGURE 99 )................................................................. 8
8(7) Funiculars much longer than wide ( Fig. 99B View FIGURE 99 ), funicle distinctly longer than head width................................................................................................ S. (T.) risbeci Lotfalizadeh & Rasplus sp. n.
- Funiculars slightly longer than wide ( Fig. 117C View FIGURE 117 ), funicle at most slightly longer than head width....................................................................................... S. (T.) maldesi Lotfalizadeh & Rasplus sp. n.
9(3) Body mostly dark or dark brown ( Fig. 123A View FIGURE 123 ); fore wing entirely setose, without speculum, basal cell setose ( Fig. 123D View FIGURE 123 ).................................................................... S. (T.) wiebesi Lotfalizadeh & Rasplus sp. n.
- Body at least partly yellowish; fore wing not entirely setose, with distinct speculum, basal cell bare at least medially..... 10
10(9) Body partly yellowish, dorsally dark or dark brown ( Fig. 114A View FIGURE 114 ); mesosoma flattened; pronotum and mesoscutum at the same level...................................................................... S. (T.) ruandensis ( Risbec, 1957) View in CoL
- Body entirely yellowish ( Fig. 132A View FIGURE 132 ); mesosoma arched; pronotum and mesoscutum not at the same level.............................................................................. S. (T.) beninensis Lotfalizadeh & Rasplus sp. n.
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