Sweltsa exquisita (Frison)

Kondratieff, Boris C. & Baumann, Richard W., 2009, A Contribution To The Knowledge Of Sweltsa Exquisita (Frison) And S. Occidens (Frison) And Description Of A New Species Of Sweltsa From The Northern Rocky Mountains, U. S. A. (Plecoptera: Chloroperlidae), Illiesia 5 (3), pp. 20-29 : 21-24

Sweltsa exquisita (Frison)

( Figs. 2-9 , 26 )

Alloperla exquisita Frison 1935: 337 . Holotype ♂, Illinois Natural History Survey: Mt. Hood National Forest , near Welches (Clackamas Co.), Oregon.

Material examined. CANADA: British Columbia: Amadis Creek, above Maple Bay Campground, Cultus Lake Provincial Park, 9 June 1991, R. W. Baumann and B. P. Stark, 1 ♂, 2 ♀ ( BYUC) ; Turbid Creek, junction Squamish River , 15 July 1988, R.W. Baumann, S. A. Wells and M. F. Whiting, 1 ♂, 4 ♀ ( BYUC) ; stream, Hwy 99, S Whistler , 16 July 1988, R. W. Baumann, S.A. Wells and M.F. Whiting, 12 ♂, 10 ♀ ( BYUC) . USA: Oregon: Clackamas Co., Cedar Creek , 11 June 1986, G.R. Fiala, 1 ♀ ( BYUC) ; Cool Creek, tributary Still Creek , 13 June 1986, G.R. Fiala, 11 ♂, 14 ♀ ( BYUC) ; Still Creek, Still Creek Campground, 28 May 1983, G.R. Fiala, 1 ♂ ( BYUC) ; Clatsop Co., Quartz Creek, Hwy 26, 20 June 1986, G.R. Fiala, 1 ♀ ( BYUC) ; Deschutes Co., Quinn River Spring, Quinn River Campground , 10 June 2004, B.P. Stark and R.W. Baumann, 4 ♂, 1♀ ( BPSC) ; Douglas Co., headwaters, Bulldog Creek, Umpqua National Forest , 19 June 1998, R. W. Wisseman, 1 ♂ ( BYUC) ; Hood River Co., McGee Creek, N side of Mount Hood , 20 July 1986, G.R. Fiala, 2 ♂, 1 ♀ ( BYUC) ; Jackson Co., 15 mi. S Talent, Rogue River National Forest , 8 July 1979, B.P. Stark and K.W. Stewart, 10 ♂, 12 ♀ ( BPSC) ; Jefferson Co., headwater spring of Metolius River , 12 June 2004, B.P. Stark and R.W. Baumann, 2 ♂, 1 ♀ ( BPSC) ; Metolius River, Camp Sherman , 11 July 1979, B.P. Stark and K.W. Stewart, 2 ♀ ( BPSC) ; Josephine Co., Lake Creek , Oregon Caves National Monument , 9 July 1979, B.P. Stark and K.W. Stewart, 1 ♀ ( BPSC) ; Klamath Co., Creed National Trail, Crater Lake National Park , 12 August 1973, J. Schuh, 2 ♂, 3 ♀ ( BYUC) ; Sun Creek, Rim Drive, Crater Lake National Park , 7 October 1991, K.W. Stewart, 1 ♂ ( BYUC) ; Lane Co., Alder Springs Campground , 3 June 2000, B.P. Stark, I. Sivec and M. Zúñiga, 3 ♂, 4 ♀ ( BPSC) ; Linn Co., tributary, North Santiam River, Hwy 22, 0.2 mi N Minto Road , 3 June 2000, B.P. Stark, I. Sivec and M. Zúñiga, 2 ♀ ( BPSC) . Marion Co., North Santiam River, Hwy 22, 3 mi above Idanha, 3 June 2000, B.P. Stark, I. Sivec and M. Zúñiga, 1 ♂ ( BPSC) ; Tillamook Co., tributary, Neskowin Creek, Hwy 12, Neskowin Creek Campground, 4 June 1991, R.W. Baumann and B.P. Stark, 1 ♀ ( BYUC) ; Wasco Co., Barlow Creek , 6 July 1986, G.R. Fiala, 5 ♂, 20 ♀ ( BPSC) . Washington: Chelan Co., stream, Glacier View Campground, Lake Wenatchee , 15 June 2004, R. W. Baumann and B.P. Stark, 2♂, 7♀ ( BYUC) ; Bridge Creek, 2 mi. S Hwy 20, Fireweed Campground , 11 July 1994, G. Mackenzie, 15 ♂, 30 ♀ ( BYUC) ; Bridge Creek, below junction Copper Creek, Pacific Coast Trail , 1 mi. off Hwy 20, 11 July 1994, G. Mackenzie, 19 ♂, 29 ♀ ( BYUC) ; Minotaur Creek , 10 mi. W Wenatchee Lake, 4 August 1996, J.R. Wood, 3 ♂, 1 ♀ ( BYUC) ; Rainey Creek , 9 mi. W Wenatchee Lake, 6 July 1975, J.R. Wood, 20 ♂ ( BYUC) ; White Pine Campground, off Hwy 2, 16 June 2004, B.P. Stark and R.W. Baumann, 1 ♂, 3 ♀ ( BPSC) ; Stevens Creek , E Stevens Pass, FR 6980, 16 June 2004, B.P. Stark and R.W. Baumann, 3 ♂, 1 ♀ ( BPSC) ; Clallam Co., Falls Creek, Marymere Falls , Lake Crescent Lodge , Olympic National Park , 15 June 2005, B. Kondratieff and R.W. Baumann, 21 ♂, 14 ♀ ( BYUC, CSUC) ; Dungeness River, Hwy 101, near Sequim , 17 June 1967, R.W. Baumann, 1 ♂ ( BYUC) ; Clark Co., Dry Creek, tributary Kalama River , 26 May 1986, G.R. Fiala, 3 ♂, 3 ♀ ( BYUC) ; Panamake Creek, tributary North Lewis River , 28 May 1986, G. R. Fiala, 3 ♀ ( BYUC) ; tributary, Merrill Lake, near Goat Rock , 26 May 1986, G. R. Fiala, 6 ♂, 1 ♀ ( BPSC) ; Cowlitz Co., spring fed tributary, Kalama River, Weyerhauser Limnology Station , 20 June 1967, R.W. Baumann and R.N. Thut, 3 ♂, 1 ♀ ( BYUC) ; Jefferson Co., Spruce Creek, Olympic National Forest , 16 July 1979, B.P. Stark, 2 ♀ ( BPSC) ; Lewis Co., Falls Creek at Stevens Canyon Road , 6 July 1999, B. Kondratieff, 7 ♂, 13 ♀ ( CSUC) ; 8 July 1999, B. Kondratieff, 4 ♂ ( CSUC) ; 25 June 2000, R. Lechleitner, 2 ♂ ( CSUC) ; 7 June 2001, R. Lechleitner, 1 ♂, 1 ♀ ( CSUC) ; 20 June 2003, B. Kondratieff, 6 ♂, 2 ♀ ( CSUC) ; 17 June 2004, B. Kondratieff, 2 ♂, 1 ♀ ( CSUC) ; Glacier Creek, above junction Johnson Creek, Walupt Lake Road , 18 July 1988, R.W. Baumann, S.A. Wells and M.F. Whiting, 2 ♀ ( BYUC) ; Panther Creek, Hwy 123, 9 June 2001, R. Lechleitner, 1 ♂ ( CSUC) ; 13 July 2002, B. Kondratieff, 5 ♂, 4 ♀ ( CSUC) ; 13 July 2003, B. Kondratieff and J. Schmidt, 5 ♂, 5 ♀ ( BYUC, CSUC) ; 14 July 2004, B. Kondratieff, 7 ♂, 3 ♀ ( CSUC) ; Ohanapecosh River , N Hwy 123, 7 July 1999, B. Kondratieff, 10 ♂ ( BYUC, CSUC) ; Ohanapecosh Campground , 16 June 2004, M. Grove, 5 ♂, 4 ♀ ( CSUC) ; Purcell Creek , 5 mi NE Packwood, 15 June 2004, B.P. Stark and R.W. Baumann, 6 ♂, 5 ♀ ( BPSC) ; White Pass, Hwy 12, 28 June 1985, B.P. Stark, 1 ♂, 1♀ ( BPSC) ; King Co. Deception Creek, below Deception Falls, Hwy 2, jct. Tye River , 17 July 1988, R.W. Baumann, S.A. Wells and M.F. Whiting, 7 ♂, 25 ♀ ( BYUC) ; creek, Stevens Pass Ski Area, Hwy 2, 17 July 1988, R.W. Baumann, S.A. Wells and M.F. Whiting, 10 ♂, 7 ♀ ( BYUC) ; Mason Co., Jorsted Creek, FR24, NE Lake Cushman , 15 June 2005, B. Kondratieff and R.W. Baumann, 5 ♂, 3 ♀ ( BYUC, CSUC) ; Purdy Creek, George Adams Fish Hatchery , 5 mi. N Shelton, 22 May 1995, G.F. Kraft, 9 ♂, 1 ♀ ( BYUC) ; Rabbit Creek , 4 mi. NW Matlock, 23 April 1995, G.F. Kraft, 1 ♂ ( BYUC) ; Okanogan Co. stream, Loup Loup Campground, Hwy 20, Okanogan National Forest , 10 June 1991, R.W. Baumann and B.P. Stark, 2 ♂ ( BYUC) ; Boulder Creek 12 mi. NW Conconully, 20 July 1995, Mackenzie, Somera and Schultz, 3 ♂, 5 ♀ ( BYUC) ; Doe Creek , FR 150, 16 mi. N Winthrop, 5 July 1995, Mackenzie, 1 ♀ ( BYUC) ; Foggy Dew Creek , 8 mi. SW Carlton, 11 July 1995, G. Mackenzie, 1 ♂, 2 ♀ ( BYUC) ; Gold Creek, 6.5 mi. SW Carlton , 21 June 1995, G. Mackenzie, 2 ♂, 4 ♀ ( BYUC) ; Goat Creek , FR 300, 15.5 mi. NW Winthrop, 3 July 1995, G. Mackenzie, 9 ♂, 13 ♀ ( BYUC) ; Ortell Creek, FR 5220, Okanogan National Forest , 17 July 1995, G.F. Kraft, 1 ♂ ( BYUC) ; Pierce Co., creek, junction Nisqually River, near Nisqually Glacier , Mount Rainier National Park , 15 June 1969, R.W. Baumann, 1 ♂ ( BYUC) ; Carbon River Entrance, Rainforest Loop Trail , 27 May 1997, B. Kondratieff, 1 ♂ ( CSUC) ; Carbon River Entrance, 27 May 1997, B. Kondratieff, 1 ♂, 1 ♀ ( CSUC) ; 20 June 2003, B. Kondratieff, 2 ♂, 2 ♀ ( CSUC) ; 12 July 2003, B. Kondratieff, 1 ♂, 1♀ ( CSUC) ; Carbon River at Chenuis Falls Trailhead, 6 June 1997, R. Lechleitner, 8 ♂ ( CSUC) ; Edith Creek, Fourth Crossing , 13 September 1997, R. Lechleitner, 2 ♂ ( CSUC) ; Fryingpan Creek, Sunrise Road , 13 July 2003, B. Kondratieff and J. Schmidt, 5 ♂, 5 ♀ ( BYUC, CSUC) ; Nisqually River, Glacier View Bridge , 7 July 1999, B. Kondratieff, 2 ♂ ( CSUC) ; Paradise River, Fourth Crossing, Valley Road , 16 August 1999, B. Kondratieff, 1 ♂ ( CSUC) ; 23 July 2000, R. Lechleitner, 3 ♂ ( CSUC) ; St. Andrews Creek, Westside Road , 14 July 2000, R. Lechleitner, 10 ♂, 5 ♀ ( CSUC) ; boggy stream, 17 mi N Paradise Road, Westside Road , 17 June 2001, R. Lechleitner, 1 ♂ ( CSUC) ; St. Andrews Creek, Westside Road , 24 July 1997, B. Kondratieff, 3 ♂, 1 ♀ ( CSUC) ; Tahoma Creek, Westside Road , 22 May 2003, B. Kondratieff, 1 ♂ ( CSUC) ; Skagit Co., unnamed creek, Hwy 11, Lake Tyee Community , 17 June 2004, B.P. Stark and R.W. Baumann, 9 ♂, 4 ♀ ( BPSC) ; Skamania Co., Falls Creek , 14 August 1983, G.R. Fiala, 6 ♂, 8 ♀ ( BYUC) ; Mosquito Creek , 14 August 1983, G.R. Fiala, 3 ♀ ( BYUC) ; Smoky Creek , 14 August 1983, G.R. Fiala, 1 ♀ ( BYUC) ; unnamed creek, 2 mi. below Big Creek Falls Viewpoint , FR 90, 14 June 2004, B.P. Stark and R.W. Baumann, 1 ♂, 2 ♀ ( BPSC) ; Snohomish Co., Owl Creek , FR 23, 16 mi. SE Darrington, 9 June 1994, G. Mackenzie, 3 ♀, 2 ♀ ( BYUC) ; unnamed creek, 1.6 mi. up FR 23, 11 mi. SE Darrington , 9 June 1994, G. Mackenzie, 4 ♂, 4 ♀ ( BYUC) ; Whatcom Co., unnamed creek, 100m S ski lift, Mount Baker , 14 July 1994, G. Mackenzie, 10 ♂ ( BYUC) ; stream, North Pass Road , 5 mi. NE Everson, 11 May 1967, K.E. Vander Mey, 2 ♂, 1 ♀ ( BYUC) ; Boulder Creek, Maple Falls , 15 May 1967, K.E. Vander Mey, 2 ♂ ( BYUC) ; Canyon Creek , NE Welcome, 27 July 1966, K.E. Vander Mey, 1 ♂ ( BYUC) ; Cornell Creek, Mt. Baker Road , 29 June 2008, D.E. Ruiter, 3 ♂ ( CSUC) ; Galena Creek, MP 53 Hwy 542, 12 mi. E Glacier, 14 July 1994, G. Mackenzie, 3 ♂, 15 ♀ ( BYUC) ; Happy Creek, Hwy 20, North Cascades National Park , 17 June 2004, B.P. Stark and R.W. Baumann, 3 ♂, 2 ♀ ( BPSC) ; Newhalem Creek, North Cascades National Park , 17 June 2004, B.P. Stark and R.W. Baumann, 11 ♂, 11 ♀ ( BPSC) ; Padden Creek, outlet Lake Bellingham , 18 May 1994, G. Mackenzie, 1 ♀ ( BYUC) ; Razor Hone Creek, Hwy 542, 13 mi. E Glacier, 14 July 1994, G. Mackenzie, 5 ♂, 17 ♀ ( BYUC) ; Ruth Creek, Hannegan Campground , 14 July 1994, G. Mackenzie, 6 ♂, 8 ♀ ( BYUC) ; Sygitowicz Creek , 2 mi SW Van Zandt, 5 May 1994, Kraft and Sammeth, 1 ♀ ( BYUC) ; Swamp Creek, Mount Baker National Forest , 11 August 1966, K.E. Vander May, 4 ♂, 10 ♀ ( BYUC) ; Yakima Co., Ahtanum Creek, Ahtanum Meadows Campground , 25 June 2000, R.L. Newell, 4 ♂, 5 ♀ ( BYUC) ; seep, 5 mi E White Pass, Hwy 12, 15 June 2004, B.P. Stark and R.W. Baumann, 3 ♂ ( BPSC) .

Male. General color yellow in life, head with black ocellar rings, pronotum yellow except for wide black lateral margins, abdomen with wide black stripe covering terga 1 to 8, black mark on tergum 8 much broader than long as shown in Frison (1935) and sometimes even slightly divided medially. Tergum 9 with a definite, bare elevated transverse process on the anterior margin ( Figs. 2 & 3 ). The anterior margin of the process varies from entire to slightly emarginate ( Figs. 2 & 3 ). In dorsal aspect, the epiproct is erectile, elongate, and subparallel ( Fig. 4 ); in lateral view, slightly enlarged subapically ( Figs. 6 & 7 ), in dorsal aspect, apex sharp and bare ( Fig. 5 ), laterally, rounded and bare at edge ( Figs. 6 & 7 ); in dorsal aspect, epiproct surface covered with dense pile of appressed long setae, extending to base ( Figs. 4-7 ), in lateral aspect, ⅔ or more of epiproct, covered by pile-like hairs, ventrally bare ( Fig. 6 ). Aedeagus tubular basally, dorsal aspect subtriangular, with two small dorsal lobes, bearing long thin setae ( Fig. 8 ), lateral, terminal aspect exhibiting paired curled hornlike lobes ( Fig. 9 ); surface with pores ( Fig. 9 ).

Female. Color and dark markings similar to male. Subgenital plate large, nearly covering sternum 9, sides subparallel, with a broadly rounded apex, that can sometimes appear truncate ( Fig. 26 ).

Figures

Figs. 2-9. Sweltsa equisita, male terminalia and aedeagus. 2. male terminalia, dorsal, Cool Creek, Oregon; 3. tergum 9, with elevated transverse process, Fryingpan Creek, Washington; 4. epiproct, dorsal, Cool Creek, Oregon; 5. epiproct apex, dorsal, Fryingpan Creek, Washington; 6. epiproct, lateral, Fryingpan Creek, Washington; 7. epiproct apex, lateral, Fryingpan Creek, Washington; 8. aedeagus, dorsal, Bridge Creek, Washington; 9. epiproct, terminal, Bridge Creek, Washington.

Figs. 26-27. Female subgenital plate, ventral. 26. Sweltsa equisita, Fryingpan Creek, Washington; 27. Sweltsa durfeei, Van Ness Creek, Montana.