Stenomacrus valvator, Varga, 2024

Varga, Oleksandr, 2024, Review of the genus Stenomacrus Förster, 1869 (Hymenoptera: Ichneumonidae: Orthocentrinae) from Kenya and Burundi: a first step to understanding the diversity of the genus in the Afrotropics, European Journal of Taxonomy 958, pp. 177-202 : 195-197

Stenomacrus valvator sp. nov.

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Fig. 8

Diagnosis

Stenomacrus valvator sp. nov. is characterized by the combination of the following: face aciculate ( Fig. 8A–B ); antenna with 18–19 flagellomeres, first flagellomere 2.8–3.4 × as long as wide; temples 0.6 × the maximum diameter of eye in dorsal view, strongly narrowed ( Fig. 8F ); propodeum with apical transverse, lateral and lateromedian longitudinal carinae present ( Fig. 8G, J ); fore wing with vein 2 rs-m about 0.6 × the distance between 2 rs-m and 2 m-cu; vein 3rs-m absent ( Fig. 8H ); first tergite 1.2 × as long as its apical width, rugulose on the granulate background, with latero-median longitudinal carinae indistinct in female ( Fig. 8I ) or clearly visible in male ( Fig. 8J ); second tergite 0.7 × as long as its apical width, weakly rugulose on granulate background ( Fig. 8I–J .); ovipositor long and upcurved, ovipositor sheath about 0.9 × as long as hind tibia ( Fig. 8A ).

Stenomacrus valvator sp. nov. differs from all known Afrotropical species in having a long and upcurved ovipositor and strongly sculptured metasomal tergites 1–2. The species also exhibits a unique character never found in the genus Stenomacrus before: absence (or at least very strong reduction) of the inner mandibular tooth which is the only feature used in defining the genus Chilocyrtus . This supports the assumption of the future synonymy of these two genera.

Etymology

The new species is named after the long ovipositor.

Type material

Holotype

KENYA • ♀; Coast Prov., Taita Hills , Vuria Forest ; 3.41428° S, 38.29178° E; 2162 m a.s.l.; 8–22 Aug. 2012; R. Copeland leg.; Malaise trap, just inside indigenous forest; ICIPE.

Paratypes KENYA • 1 ♂, 1 ♀; same data as for holotype; ICIPE • 1 ♂; Coast Prov., Taita Hills , Ngangao Forest ; 3.36100º S, 38.34186º E; 1848 m a.s.l.; 10–24 Jan. 2012; R. Copeland leg.; Malaise trap, indigenous forest; ICIPE .

Description

Female

Body length 3.0– 3.1 mm. Fore wing 2.7–2.8 mm.

Head smooth and sparsely pubescent. Antenna with 18–19 flagellomeres, first flagellomere 2.8–3.4 × as long as wide. Face about 0.5 × as long as wide, aciculate; inner orbits parallel. Malar space 2.5 × basal width of mandible; subocular sulcus distinct. Clypeus 0.6 × as long as wide, weakly convex. Mandible unidentate, strongly bent outwards. Temples 0.6 × maximum diameter of eye in dorsal view, strongly narrowed. Frons and vertex smooth; length of ocellar-ocular distance about 1.4 × maximum diameter of lateral ocellus; occipital carina absent.

Propleuron sparsely pubescent. Pronotum smooth; epomia present. Mesoscutum more-or-less evenly densely pubescent; notauli absent. Scutellum densely pubescent, with carinae present only on basal 0.1. Mesopleuron smooth; epicnemial carina present laterally. Metapleuron smooth; pleural and submetapleural carinae present. Propodeum weakly granulate, with apical transverse, lateral and lateromedian longitudinal carinae present, rugulose between lateromedian longitudinal carinae.

Legs relatively stout; hind femur 3.5 × as long as wide, third tarsomere of hind tarsus about 0.9 × length of fifth tarsomere; tarsal claws simple.

Fore wing with vein 2 rs-m about 0.6 × distance between 2 rs-m and 2 m-cu; vein 3rs-m absent; vein 1cu-a weakly distad M & Rs; hind wing with nervellus not intercepted, weakly reclivous.

Metasoma with first tergite 1.2 × as long as apical width, rugulose on granulate background, with latero-median longitudinal carinae clearly visible only basally; lateral oblique grooves distinct and deep. Second tergite 0.7× as long as apical width, granulate, with basolateral and subapical transverse grooves; thyridium elongate. Third tergite with traces of granulation. Remaining part of metasoma smooth, not laterally compressed. Ovipositor long and upcurved, ovipositor sheath about 0.9 × as long as hind tibia, apical part widened and pubescent.

Body generally brownish except scape and pedicel, face narrowly below antennal sockets, clypeus partly, and mandible (except apices) yellow. Legs generally yellow: fore and mid femora, tibiae and tarsi darker dorsally; hind coxa brownish basally; hind tibia reddish with fuscous line dorsally; hind tarsus fuscous.

Male

Generally resembles female, but has wider temples, longer and more strongly sculptured metasoma with more distinct carination ( Fig. 8B, J ).

Distribution

So far only known from Kenya.

Figures

Fig. 8. Stenomacrus valvator sp. nov. A, C, E-I. Holotype, ♀ (ICIPE). D. Paratype, ♀ (ICIPE). B, J. Paratype, ♂ (ICIPE). A–B. Lateral view of habitus. C. Frontal view of face. D. Dorsal view of mandible: absence of inner tooth arrowed with red. E. Lateral view of head and mesosoma. F. Dorsal view of head and mesoscutum. G. Dorsal view of propodeum. H. Wings. I. Dorsal view of metasomal tergites 1–2. J. Dorsal view of propodeum and metasomal tergites 1–2. Scale bars: A–B = 0.5 mm; C–J = 0.1 mm.