Sinochloritis lii Wu & Chen, gen. &

Wu, Min, Chen, Zheyu & Zhu, Xiaoran, 2019, Two new camaenid land snails (Eupulmonata) from Central China, ZooKeys 861, pp. 129-144 : 132-136

publication ID	https://dx.doi.org/10.3897/zookeys.861.35430
publication LSID	lsid:zoobank.org:pub:DAD076A6-B8B7-4FEE-94C4-CFB56E0B6338
persistent identifier	https://treatment.plazi.org/id/FCC544B8-3DD5-4785-A1C0-5A2E2240C55E
taxon LSID	lsid:zoobank.org:act:FCC544B8-3DD5-4785-A1C0-5A2E2240C55E
treatment provided by	ZooKeys by Pensoft
scientific name	Sinochloritis lii Wu & Chen, gen. &
status	sp. nov.

Treatment

Sinochloritis lii Wu & Chen, gen. & sp. nov. Figs 1 View Figure 1 , 2 View Figure 2 , 3 View Figure 3 , 4 View Figure 4 , 5 View Figure 5 , 6 View Figure 6

Type material.

Holotype, fully matured animal (HBUMM08294). Sichuan Province, Dujiangyan, Qingchenghoushan, 30°56'39.38"N, 103°28'47.21"E, 1500 m a. s. l., 2018-XI-8, coll. Li, Chenliang & Zhu, Xiaoran. A sample of foot muscle tissue was preserved in 99.7% ethanol at -20 °C (HBUMM08295). Paratypes, 1 old fms (HBUMM10008), Sichuan, Dujiangyan, Qinchenghoushan, 1500 m a. s. l., 2018-V, coll. Liu, Zhengping; 1 broken fully matured shell (HBUMM10009), Sichuan, Dujiangyan, Qinchenghoushan, 1500 m a. s. l., 2017-X, coll. Liu, Zhengping.

Description.

Shell ( Fig. 2 View Figure 2 ). Depressed; thick and solid; dextral. Whorls convex. Suture rather impressed. Umbilicus closed by reflexed columellar lip. Columella oblique. Proto conch and teleoconch densely and uniformly covered with fine granules, without spiral furrows. Aperture oblique; not sinuate at peristome. Body whorl not descending behind aperture. Shell surface without ribs. Growth lines fine. Adult shell not hairy or scaly. Adult body whorl rounded at periphery; basally convex. Ring-like thickening within aperture absent. Peristome thin; abruptly angled at top; narrowly and uniformly reflexed; brownish purple. Callus thin and transparent. Shell glossy; uniformly reddish brown. Measurements (type material): shell height 16.0-17.1 (16.5 ± 0.55) mm, shell breadth 25.0-30.6 (27.0 ± 3.10) mm, aperture height 11.5-12.5 (11.9 ± 0.51) mm, aperture width 13.4-16.8 (14.9 ± 1.72) mm, embryonic shell whorls 1.375-1.500 (1.458 ± 0.072), whorls 4.750-4.875 (4.833 ± 0.072), shell height/ breadth ratio 0.56-0.65 (0.62 ± 0.049).

General anatomy ( Fig. 3 View Figure 3 ). A heart-shaped head gland between ommatophore insertions present on inner body wall ( Fig. 3C View Figure 3 , arrowed), externally with a visible gland pore ( Fig. 3A View Figure 3 , arrowed). On internal body wall, at the base of ommatophore with two groups of glands each consisted of numerous small sacs ( Fig. 3C View Figure 3 ). On left side of mantle edge, a leaf-shaped appendage present ( Fig. 3D, E View Figure 3 ). Body blueish purple with scattered lighter spots ( Fig. 3H View Figure 3 ). Sole dirty white. Jaw arcuate; with twelve more or less projecting ribs ( Fig. 3F View Figure 3 ).

Genitalia ( Figs 4 View Figure 4 , 5 View Figure 5 ). Penis sheath absent. Penis thick; externally simple; internally with five thick and high plicae/pilasters ( Fig. 5D View Figure 5 ). Epiphallus longer than penis; with section between penial retractor muscle and epiphallic papilla one-third thickness of penis; section between penial retractor muscle and vas deferens insertion much thicker than proximal part but thinner than penis ( Fig. 5A View Figure 5 ); internally with a large peach-shaped epiphallic papilla (approximate size 3.5 ×4.0× 6.0 mm3) entering penis ( Fig. 5A View Figure 5 , B); externally, partially connected with distal penis by strong muscles that insert on penis just opposite to penial retractor muscle ( Figs 4C View Figure 4 , 4D View Figure 4 , 5A View Figure 5 , arrowed). Flagellum cylindrical; tapering. Inside flagellum and epiphallus, a long pilaster running from tip of flagellum to epiphallic papilla and a much shorter wavy pilaster running from tip of flagellum to vas deferens insertion ( Fig. 5C View Figure 5 ). Vas deferens thin; of even thickness ( Fig. 4A View Figure 4 ). Vagina subequal to penis in length ( Fig. 4A View Figure 4 ). Base of bursa copulatrix duct expanded and ball-shaped ( Fig. 4A View Figure 4 ); internal wall strongly corrugated ( Fig. 5E View Figure 5 ). Measurement of holotype: P– 13.6 mm; Ep– 19.3 mm; Fl– 8.6 mm; VD– 31.4 mm; PR– 4.6 mm; Va– 18.8 mm; FO– 5.4 mm; BC plus BCD– 37.9 mm.

Etymology.

This species is named in honor of Dr. Chenliang Li who collected and sent us the holotype (HBUMM08294).

Distribution.

Sichuan (Qingchengshan), only known from the type locality ( Fig. 1 View Figure 1 ).

Ecology.

This species was found living in the well-developed forest ( Fig. 6 View Figure 6 ).

Taxonomic remarks.

The new species has a closed umbilicus, otherwise is very close to Trichochloritis percussa in shell size (Table 2), the general shape and the microsculpture of shell. Camaena hemiclista (Schmacker & Böttger, 1894) known only from Hubei (Lytschouan=Lichuan), which has a closed umbilicus and is bluntly shouldered (but not visible in Yen 1939: pl. 12, fig. 42) resembles the new species; however, the new species has fewer whorls and a clearly rounded periphery.