Scopaeus diversilobatus Frisch, 2022

Frisch, Johannes & Narakusumo, Raden Pramesa, 2022, Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species, Soil Organisms 95 (1), pp. 23-73 : 42-48

publication ID	https://doi.org/ 10.25674/so95iss1id311
DOI	https://doi.org/10.5281/zenodo.10724307
persistent identifier	https://treatment.plazi.org/id/03B7C77A-FFD3-FFED-BF14-FAB50FAB07D0
treatment provided by	Felipe
scientific name	Scopaeus diversilobatus Frisch
status	sp. nov.

Treatment

Scopaeus diversilobatus Frisch , spec. nov.

( Figs 15 View Figures 14–16 , 39 View Figures 35–41 , 89–91 View Figures 86–91 , 133 View Figures 119–133 , 165, 166 View Figures 163–172 )

Type specimens: Holotype ♂, Indonesia, Jawa Barat, Sukabumi, N Djampang Kulon: Cikarang River (07°14’03’’S, 106°36’49’’E), 250 m, 25.9.2015, leg. Frisch ( MZB) GoogleMaps . Paratypes (10 specimens): 3 ♂, 4 ♀, same data as holotype ( MFNB, MZB). Indonesia, Bali GoogleMaps : 1 ♂, Jembrana: Cekik, 20.6. 1994, 300 m, leg. Wunderle ( PWCM). Malaysia, Sarawak : 1 ♂, Undong ( San Uncap Undong ?), 17.4.1981, leg. Rougemont ( HECO) ; 1 ♂, Mt Mulu NP, near base camp, 50 – 10 m 5.8.1978, leg. Hammond & Marshall ( NHML) .

Description: Habitus and coloring as in Fig. 15 View Figures 14–16 . Head subquadrate. Penultimate antennal segment elongate. Mesotibia moderately thickened. Body surface with fine, dense, setose punctation, without microreticulation, shiny. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body dark brown except for slightly lighter pronotum, yellow-brown posterior fourth of elytra, medium brown abdomen with darkened disk of tergites and light brown tip, yellow brown maxillary palpi with darker penultimate segment, medium brown antennae with darker median segments, and yellow-brown legs. Total body length 3.6 – 3.9 mm; forebody length 2.1 – 2.3 mm.

Male: Abdominal sternite VII with unmodified posterior margin. Emargination in posterior fifth of abdominal sternite VIII strongly convex in median third ( Fig. 133 View Figures 119–133 ). Aedeagus ( Figs 39 View Figures 35–41 , 89 – 91 View Figures 86–91 ) about 1.0 mm long with distal lobes asymmetrical; dextral apical lobe strongly reduced, as long as dorsal lobe, in lateral view with undulate, convex apical and ventral margins ( Fig. 39 View Figures 35–41 , 89 View Figures 86–91 ), in ventral and dorsal view almost vestigial with truncate apex ( Figs 90, 91 View Figures 86–91 ); sinistral apical lobe well developed, testaceous, twice as long as dextral apical lobe, in lateral view triangular ( Fig. 39 View Figures 35–41 , 89 View Figures 86–91 ), in ventral and dorsal view convex laterally with somewhat retracted, acute apex ( Figs 90, 91 View Figures 86–91 ); flagellum curved ventrosinistrad with thin end projecting from apical lobes ventrally ( Figs 39 View Figures 35–41 , 89 View Figures 86–91 ); lateral lobes reduced, indicated by group of setae ( Figs 89, 90 View Figures 86–91 ); ventral lobe long, slender, adjacent to aedeagus ( Figs 39 View Figures 35–41 , 89 View Figures 86–91 ), in ventral view narrow and shifted sinistrad ( Fig. 90 View Figures 86–91 ); transverse ridge of median foramen not projecting ventrally ( Fig. 89 View Figures 86–91 ).

Female: Sperm pump with process segment and apophysis of chamber long, slender and curved at ends ( Fig. 166 View Figures 163–172 ); bursal duct about twice as long as sperm pump; bursa membranous ( Fig. 165 View Figures 163–172 ).

Phylogeny: Scopaeus diversilobatus is to be classified in the S. gracilis species group ( Frisch et al. 2002: 39).

Distribution: Hitherto recorded from western Java, Bali and northern Borneo, Scopaeus diversilobatus is widespread in the Sunda Islands.

Etymology: The epithet diversilobatus [adjective, Latin, composed of the adjectives diversus (different) and lobatus (indicating a characteristic/feature of the lobe)] describes the different size and shape of the apical lobes of the aedeagus of this new species, one of which is strongly reduced unlike the other ( Figs 87, 88 View Figures 86–91 ).