Scaphisoma signaticolle, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1433-1434

5.2.21.2. Scaphisoma signaticolle nov.sp. ( Figs 231-233 )

T y p e m a t e r i a l: Holotype 6: Luzon: Mountain Prov. Mount Data Lodge 2200-2300m 8.I.1980 leg. J. Orousset (MHNG) [154, litière près de la lodge]. Paratypes: with the same data, 1♀ [161, humus dans un ravin près d’un ruisseau]; Luzon, Mountain Prov., N & NE Sagada, 15.- 19.XII.1979, leg. L. Deharveng & J. Orousset, 1♀ [39, en aval de Sagada, pied de paroi calcaire]; same data, [51, doline sombre au nord de Sagada, mousses sur tronc de caféier], 3♀♀; with same data, 1♀ [72, doline au nord-est de Sagada, litière humide, éclairée]; with same data, 1♀ [143, vallon à l’est du cimetière de Sagada, litière épaise humide]. (All in MHNG).

D e s c r i p t i o n: Length 2.05-3.20 mm, width 1.37-1.82 mm. Head dark reddishbrown to blackish. Most of body very dark brown to blackish. Hypomera blackish to ochreous. Elytra each with apical fourth ochreous, not sharply delimited from darker anterior area. Abdominal apical sternites brown. Antennae yellowish, legs ochreous. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 15/11: IV 50/8: V 65/8: VI 58/8: VII 60/14: VIII 53/9: IX 66/12: X 55/12: XI 70/13. Pronotum with lateral margins evenly rounded, lateral margin carinae not exposed in dorsal view, lateral striae punctate; pronotal punctation conspicuously dense, punctures variably large, often coarse, mostly about half as large to as large as puncture intervals. Tip of scutellum exposed. Elytra with lateral margin carinae entirely and well visible in dorsal view, apical angles situated posterior level of outer angles; sutural margin not raised, sutural striae fairly deep, starting posterior level of scutellar tip, parallel except near apices, finely punctate; adsutural areas flat, densely and finely punctate, with punctures smaller than those on elytral disc, lateral striae distinctly punctate. Elytral disc with punctation coarse, punctures often well delimited, punctures intervals mostly about as to twice as large as puncture diameters. Hind wings fully developed. Hypomera and mesanepisterna smooth not microsculptured, usually distinctly punctured. Mesepimera shorter than intervals to mesocoxae, about four times as long as wide. Median part of metaventrite slightly convex, with shallow apicomesal impressions and distinct strigulate microsculpture; punctation fine except in impressions. Lateral parts of metaventrite not microsculptured, with distinct punctures; antecoxal puncture rows present, in impressed striae. Submesocoxal areas 0.03-0.05 mm, about as sixth to eighth of intervals to metacoxae, submesocoxal lines subparallel to convex, coarsely punctate. Metanepisternum somewhat convex, weakly narrowed anteriad, inner margin rounded, impressed. Metepimera punctate and microsculptured. Abdomen with strigulate microsculpture. Sternite 1 distinctly and densely punctate basomedian area and usually with fairly coarse punctures on lateral area; submetacoxal areas 0.06-0.09 mm, about as fourth of intervals to apical margin of sternite 1, submetacoxal lines convex, coarsely punctate. M a l e: Protibiae and metatibiae straight, mesotibiae distinctly curved, thicker than metatibiae. Protarsomeres 1 to 3 and mesotarsomeres 1 and 2 strongly widened, tarsomeres 1 wider than apices of respective tibiae. Lobe of sternite 6 flat. Aedeagus with apical process of median lobe ( Figs 231-233 ) strongly inflexed and sinuate, in lateral view gradually narrowed, in dorsal view almost evenly broad in apical half.

F e m a l e: Elytra with apical margins oblique or slightly uneven; inner apical angles prominent, each forming about 0.02-0.10 mm long tooth.

E t y m o l o g y: The species epithet is a Latin adjective referring to the well characterized pronotum.

C o m p a r a t i v e n o t e s: This species may be readily distinguished from other members of the S. tricolor group by its comparatively coarse and very dense pronotal punctation, and distinct punctation on mesanepisterna and metanepisterna. The hypomera are finely punctate in the sole male available, but quite distinctly punctate in females. The specimens were inadequately conserved and became extremely fragile. The aedeagus felt in to pieces while dissection, and only the apical process of the median lobe and to part also the internal sac remained suitable for description and illustration. The shape of the apical process is, however, diagnostic.

Figures

Figs 228 - 233: Aedeagi of Scaphisoma; (228); S. pulchrum nov. sp., median lobe in dorsal view; (229), ditto, aedeagus in lateral view, without proximal part of basal bulb; (230), ditto, parameres in ventral view; (231, 232), S. signaticolle nov. sp., apical process of median lobe in dorsal and lateral views; (233), internal sac. Scale = 0.2 mm.