Saximormia jelineki, Ježek, 2010

Ježek, Jan, 2010, Further new taxa of non-biting moth flies (Diptera: Psychodidae: Psychodinae) from Malaysia, Acta Entomologica Musei Nationalis Pragae 50 (1), pp. 235-252 : 237-240

Saximormia jelineki sp. nov.

( Figs. 1–18 , 53 )

Type locality. Malaysia, Perak, Hulu, Belum Camp (5°30′07″N, 101°26′21″E).

Type material. HOLOTYPE: J ( NMLS), MALAYSIA: PERAK: Hulu, Belum Expedition, Belum Camp , iv.–vi.1994, light trap (Rothamsted). Slide , Canada Balsam , dissected specimen . PARATYPES: 4JJ ( NMPC:Cat No.34494–34497, Inv. No. 17656–17659), the same locality and data, dissected. The figures are based mainly on the holotype.

Description. Male. Head prolonged ( Fig. 1 ), vertex conspicuously enlarged, eyes touching for 3.3 diameters of one facet, eye bridge of four facet rows, however, in the touch of eyes only three facets on both sides ( Fig. 53 ). Frontoclypeus with hexagonal patch of insertions of hairs, upper area of patch with shallow cleft. Antennae probably with 16 antennomeres (terminal flagellomeres missing). Scape very short ( Fig. 2 ), widened distad, 1.2 times as long as pedicel, the latter almost globular. Basal flagellomeres excentrically bulbous with rather short and broad necks. Ascoids of flagellomeres approximately rectangular, embracing their constricted parts ( Fig. 10 ), paired. Upper side of sensory rectangle inconspicuously curled. Length ratios of maxillary palpomeres equal to 1: 1.3: 1.3: 1.4. Palpomere 4 annulate, connected basally with subapical area of palpomere 3 ( Fig. 3 ). Labium and labial lobes with many folds terminally ( Fig. 11 ). Cibarium 2.3 times as long as epipharynx.

Thoracic sclerites as in Fig. 12 . Wing ( Fig. 13 ) widely lancet shaped, 0.9–1.2 mm long (1.2 mm in holotype), wing membrane bare, clear, radial and medial forks complete; the following veins or their parts strengthened: Sc inconspicuously at base, R 1 distally (but not at apex), R 2+3+4, basis of M 1+2, entire M 4 and Cu. Basal costal nodes distinct. Sc uninterrupted. Bases of M 3 and M 4 at large distance to base of Cu. R 5 extending distally to reach wing margin below the rounded apex of the wing. Veins r-r, r-m and m-m not developed. Medial wing angle 152° (BCD). Wing 2.5 times as long as wide. Length ratios of femur, tibia and first tarsomere: P 1 = 1.9: 21.1: 1, P 3 = 2: 2.8: 1 (ratio for P 3 measured in a different specimen), P 2 missing in examined specimens. Fore claws as in Fig. 14 .

Basal apodeme of male genitalia rather short and thin, a little bent from lateral view, straight from dorsal one. Aedeagal complex ( Figs. 5–7 , 17–18 ) propped upon a sclerite of complicated shape at large distance to basal apodeme (lateral view). Male copulatory organ formed by two jointed phallomeres and one ventral declinated phallomere, with a similar overall morphology to the genitalia of the tribe Psychodini (incorrectly interpreted as asymmetrical copulatory organ by some authors). Gonocoxites ( Figs. 4 , 15 ) rather slender, expanded at base, gonostyli long, gradually tapering to apex, sharply pointed and a little bent terminally. Epandrium ( Fig. 8 ) almost bare, several insertions of hairs caudally on both sides of a shallow epandrial notch. Basal paired apertures visible. Remainders of tergite 10 and sternite 10 inside epandrium conspicuous, largely V-shaped. Hypandrium narrow, bare. Hypoproct large, long, triangular, 2.3 times as long as epiproct ( Fig. 9 ); epiproct small, short, almost hemicircular, tongue-shaped, both parts haired. Surstyli cylindrical ( Figs. 9 , 16 ), straight (dorsal view), a little bent from lateral view. Seven retinaculi present.

Female unknown.

Differential diagnosis. Saximormia jelineki sp. nov. is characterized by the following characters: curled upper side of the sensory rectangles (ascoids) ( Fig. 10 ), short and thin basal apodeme of the male genitalia, aedeagal complex formed by two jointed phallomeres and one ventral declinated phallomere and propped upon a sclerite of complicated shape at large distance to basal apodeme ( Figs. 5–7 , 17–18 ), gonostyli straight, not corrugate ( Figs. 4 , 15 ), and surstyli with seven retinaculi ( Fig. 9 ).

Saximormia saxicola ( Tokunaga & Komyo, 1955) differs from the new species by having the upper side of sensory rectangles not curled. Unfortunately, the type material of S. saxicola is probably lost (M. Sasakawa, pers. comm.) and there is no illustration of the aedeagal complex in the original paper of TOKUNAGA & KOMYO (1955). The gonostyli of S. saxicola are little corrugated and the surstyli have six retinaculi.

Etymology. The species is dedicated to my dear colleague and friend Josef Jelínek (National Museum, Prague).

Bionomy. Unknown. The males were collected by a light trap in a primeval rainforest.

Distribution. Peninsular Malaysia: Perak state.

Figures

Figs. 1–9. Saximormia jelineki sp. nov. (J). 1 – head; 2 – basal antennomeres; 3 – maxilla and maxillary palpus (palpomere 4 deformed); 4 – gonocoxite and gonostylus, dorsal view; 5 – aedeagal complex, dorso-lateral view; 6 – the same, dorso-ventral view; 7 – the same, diagonal view; 8 – epandrium, dorsal view; 9 – surstyli (retinaculi omitted), dorsal view. Scales = 0.05 mm (Figs. 2, 5–7) and 0.1 mm (Figs. 1, 3–4, 8–9).

Figs. 10–18. Saximormia jelineki sp. nov. (J). 10 – middle flagellomere, ascoids, axial view; 11 – terminal lobes of labium and epipharynx; 12 – thoracic sclerites, lateral view; 13 – wing; 14 – claw of P1; 15 – gonocoxite and gonostylus, lateral view; 16 – epandrium and surstylus, lateral view; 17 – aedeagal complex, lateral view; 18 – same, dorsal view. Scales = 0.05 mm (Figs. 10–11, 14, 17–18), 0.1 mm (Figs. 12, 15–16) and 1 mm (Fig. 13).

Figs. 53–55. 53 – Saximormia jelineki sp. nov. (J), eye bridge; 54 – Nototelmatoscopus (Jozifekia) sasakawai sp. nov.(J), wing (phase contrast observation); 55 – Perakomyia sifneri sp.nov. (J) – wing (phase contrast observation). Scales = 0.05 mm (Fig. 53) and 1 mm (Figs. 54–55).