Sagola brumfieldi Park and Carlton

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 141

Sagola brumfieldi Park and Carlton , new species

( Figs. 68o , 69o 70o , 73 )

Type Material. Holotype. NEW ZEALAND: Taupo: 1♂ ( NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand: Tongariro N.P., Mangawhero R. Rd. near Ohakune, S. Slope of Mt. Ruapehu TO”, “Forest litter berlese at FIT 4 , 10 Mar 2000 C Carlton, A Weir, #034”, “ HOLOTYPE Sagola brumfieldi Park and Carlton 2013 ” . Paratypes (5 females). NEW ZEALAND: Waikato : 2♀♀ ( NZAC) , Mahoenui, Gribbons Rd , 26 VI 1977, G.W. Ramsay, litter ; 1♀ ( NZAC) , Mahoenui, Gribbons Rd , 26 VI 1977, N.H. Mancer, litter ; 2♀♀ ( NZAC) , nr Waitomo, Mathers Rd, Fred Catchment , 23 V 1983, litter .

Etymology. This species is named for one of the enthusiastic supporters of this study, Robb T. Brumfield. He was a member of the first author’ s PhD advisory committee and provided valuable advice during this study, a major component of his dissertation.

Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.4–2.6 mm; temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; elytra subquadrate; shape of antennomeres and genitalia unique to species.

Description. Length 2.4–2.6 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68o ). Head: Male head triangular, widest across temples ( Fig. 69o ). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-fourth length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface ( Fig. 69o ). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae ( Fig. 69o ). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate ( Fig. 68o ). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Aedeagus: Median lobe of genitalia widest at middle, with flat apical margin ( Fig. 70o ). Phallobase of median lobe symmetrical and rounded ( Fig. 70o ). Parameres symmetrical and slender, with sparse setae at apex and midpoint ( Fig. 70o ).

Distribution. Taupo, Waikato ( Fig. 73 : black squares).

Habitat. Specimens of this species were collected by sifting leaf litter.

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 73. Known collection localities of New Zealand Sagola species. S. cuvierensis: black circle; S. buckleyi: triangle; S. brumfieldi: black squares; S. otteai: stars; S. boudreauxae: white circle; S. taegyui: white squares.