Rhaphaulus tonkinensis Pall-Gergely , Hunyadi & Maassen, 2014

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Rhaphaulus tonkinensis Pall-Gergely, Hunyadi & Maassen, 2014

Figs 13E, F , 14A

Rhaphaulus tonkinensis Páll-Gergely et al. 2014: 567, 569, fig. 1. Type locality: rocky wall, left side of the road nr. 6, 156 km towards Moc Chau, Ha Noi, Son La Province, Vietnam. Do et al. 2015: 128, fig. 7d, Son La Province, Vietnam. Páll-Gergely et al. 2017: fig. 1a-e. Raheem et al. 2017: 6 (plate figure).

Type material examined.

Holotype HNHM 98757 from Ha Noi, Son La Province, Vietnam (Fig. 13E ).

Other material examined.

CUMZ 12163 (4 shells; Figs 13F , 14A ) from Luang Cave, Mae Sai District, Chiang Rai Province, 23 Oct. 2015. CUMZ 12164 (2 shells) from Pha Mee Cave, Mae Sai District, Chiang Rai Province, 23 Oct. 2015.

Diagnosis.

Shell elongated ovate; body whorls slightly bulging. Tube thick and flat, turning first straight upward then abruptly downward, highly widening and extending to nearly the entire last whorl height.

Differential diagnosis.

Rhaphaulus tonkinensis can be distinguished from all other species from mainland Southeast Asia by a distinctive tube that is thick and flat, turning first straight upward then abruptly downward, greatly widening and extending to nearly the entire last whorl height.

Distribution.

Northern Vietnam ( Do et al. 2015) and Chiang Rai Province, northern Thailand.

Remarks.

The tube of one specimen from Tham Luang, Mae Sai District, Chiang Rai Province when turning downward does not adhere to the apertural margin (Fig. 13F ). However, the tube of another specimen from the same locality adheres to the apertural margin (Fig. 14A ), identical to the holotype ( Páll-Gergely et al. 2014). Thus, the extent of tube adherence to the apertural margin is treated as an infraspecific variation.

Figures

Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)

Figure 14. A Rhaphaulus tonkinensis, specimen CUMZ 12163 / 2 from Luang Cave, Chiang Rai. B-D Rhaphaulus chrysalis B possible syntype NHMUK 2013.04.16 from Siam C specimen NHMUK 1871.9.23.52 from Burma, and D specimen NHMUK 1903.7.1.3073 from Molmein.