Rhammatophyllum gaudanense (Litv.) Al-Shehbaz & O. Appel, Novon, 2002

Moazzeni, Hamid, Zarre, Shahin, Assadi, Mostafa, Joharchi, Mohammad Reza & German, Dmitry A., 2014, Erysimum hezarense, a new species and Rhammatophyllum gaudanense, a new record of Brassicaceae from Iran, Phytotaxa 175 (5), pp. 241-248 : 244

Rhammatophyllum gaudanense (Litv.) Al-Shehbaz & O. Appel, Novon 12: 3. 2002 [2 Apr. 2002]

Basionym: Erysimum gaudanense Litvinov, Trudy Bot. Muz. Imp. Akad. Nauk 1: 33. 1902.

Synonyms: Prionotrichon gaudanense (Litv.) Botschantzev, Novosti Sist. Vyssh. Rast. 3: 125. (1966).

Rhammatophyllum gaudanense (Litv.) Kamelin, Bot. Zhurn. 87, 12: 109. 2002 [24 Dec. 2002], comb. superfl.

TYPE: TURKMENISTAN. Turcomania. In declivibus herbosis montium pr. Gaudan , 28 IV 1898 , D. I. Litwinow 518 (lectotype, LE!, designated by Botschantzev (1966: 125, “type”); isolectotypes, B!, BRNM!, JE!, LE!, WU!). Figs. 1D , 2 .

Distribution: Iran and Turkmenistan (endemic to Kopet-Dagh).

Plants subshrubs. Trichomes on stems, leaves, pedicels, sepals, and fruits softly malpighiaceous or submalpighiaceous, with two (rarely three) equal or unequal, unbranched or rarely branched principal rays, usually crisped and predominantly bearing a few lateral minute branches; simple on petals. Stems erect to ascending, (20) 30–45 cm tall, usually several at base, branched below (annual shoots unbranched) and slightly swollen at nodes. Basal leaves fasciculate, spatulate, 5–15 × 1–2(3) mm, round at apex, attenuate at base. Cauline leaves sessile or shortly petiolate, linear to oblanceolate, entire, often longitudinally plicate, 10–35 × 1–2 mm. Racemes 6–16-flowered, ebracteate, corymbose, considerably elongated in fruit. Fruiting pedicels straight, slender, divaricate-ascending to suberect, (4)5– 8(10) mm. Sepals oblong, membranous at margin, 3–4 mm long. Petals creamy white, becoming yellow when drying, 5–8 × 2–3 mm, pubescent outside. Stamens 6, tetradynamous; filaments not dilated or slightly flattened at base; anthers oblong, apiculate at apex. Fruits dehiscent siliques 25–50 × 1.5–2 mm, linear, latiseptate, sessile or shortly stipitate, suberect, straight or slightly flexuous, shallowly constricted between seeds; valves torulose, with a narrow midvein; style obsolete or distinct, up to 2 mm long; stigma capitate, 2-lobed. Seeds winged at top, oblong, strongly flattened; seed coat not mucilaginous when wetted; cotyledons accumbent.

Specimen examined: IRAN. Khorasan, S Kalat-e Naderi , between Baghkand and Ortekand. 1900 m, 10 July 2002, Joharchi 34207 ( FUMH!)

Figures

FIGURE 1. Erysimum hezarense sp. nova.A: habit, B: flower, C: mature fruit and orientation of pedicel, D: distribution map of Erysimum hezarense (○ colored in red), E. laxiflorum (◊), E. subulatum (♦), E. iraqense (G) and Rhammatophyllum gaudanense ().

FIGURE 2. Rhammatophyllum gaudanense. Drawing prepared from Jouharchi 34907 (FUMH). A: part of fruit, B: seed with wing, C: plant. Scale bar in A: 2 mm; B: 0.5 mm; C: 1 cm.