Raveniola tarabaevi, Zonstein, 2024

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 75-77

Raveniola tarabaevi sp. nov.

urn:lsid:zoobank.org:act:E4DE9EC3-3D2A-4F4B-9167-3D80A69832E5

Figs 59 , 132 , 191 , 252 , 544–546 , 613 , 759

Diagnosis

The new species can be distinguished from other members of the group by having smaller PME (which are about 0.7 times as large as AME) and, correspondingly, by larger interdistances AME–PME and PLE–PME (which are approximately equal to two and one PME diameters, respectively). Within other related species, AME are subequal to PME, and the distances AME–PME and PLE–PME are clearly shorter than two and one PME diameter, respectively. The structure of the spermathecae in R. tarabaevi sp. nov. most closely resembles that in R. kirgizica sp. nov.; both species share a relatively short and subapically constricted spermathetical trunk, combined with a long lateral diverticulum. However, in R. tarabaevi , the main trunk is not so slender as in R. kirgizica ( Figs 544–546 cf. Fig. 537 ).

Etymology

The specific epithet is given in honour and memory of Chingiz Tarabaev (1951–1999), for his significant personal contribution to the modern study of the Central Asian spider fauna.

Material examined

Holotype

KAZAKHSTAN • ♀; Zhetyzhol Mts , Kastek Ridge , Uzunbulak Gorge, eastern slope of Mt Beriktas ; 42°54′ N, 75°33′ E; 1700–1900 m a.s.l.; 12 May 1988; S. Zonstein and S. Ovchinnikov leg.; SMNH.

Paratypes

KAZAKHSTAN • 6 ♀♀; same collection data as for preceding; SMNH .

Additional material

KAZAKHSTAN • 2 ♀♀; Trans-Ili Mts , Turgen Canyon 8 km S of Turgen Town, forest protection area; 43°18′ N, 77°38′ E; 1500 m a.s.l.; 12 Jul. 1993; S. Ovchinnikov and D. Milko leg.; SMNH .

Description

Female (holotype)

HABITUS. See Fig. 59.

MEASUREMENTS. TBL 15.75, CL 4.91, CW 4.54, LL 0.47, LW 0.98, SL 2.62, SW 2.36.

COLOUR. Carapace, sternum, labium, maxillae, palps and legs uniformly light ginger orange; eye tubercle blackish brown; chelicerae reddish orange; abdomen and spinnerets pale grayish brown; abdomen dorsally with slightly darker grayish cardiac mark followed behind by several very small and almost indistinct brownish chevrons.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 132 . Clypeus and eye group as in Fig. 191 . Eye diameters and interdistances: AME 0.12(0.16), ALE 0.27, PLE 0.14, PME 0.08; AME–AME 0.14(0.10), ALE–AME 0.12(0.10), ALE–PLE 0.11, PLE–PME 0.07, PME–PME 0.37. Cheliceral rastellum absent. Each cheliceral furrow with 9 promarginal teeth and 3–4 relatively large mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 252 . Maxillae with 22 cuspules each.

LEGS. Scopula: entire and distal on metatarsi I–II; narrowly divided on palpal tarsus and tarsi I–II; sparse and widely divided on tarsus III; absent on tarsus IV. Trichobothria: 2 rows of 8–9 each on tibiae, 14–16 on metatarsi, 11–13 on tarsi, 9 on palpal tarsus. Palpal claw with 4 promarginal teeth. PTC I–IV with 6–8 teeth on each margin.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	2.77	1.43	1.88	–	1.63	7.71
Leg I	3.43	2.07	2.79	1.94	1.58	11.81
Leg II	3.24	2.01	2.30	2.08	1.59	11.22
Leg III	3.05	1.82	2.03	2.65	1.67	11.22
Leg IV	3.89	2.03	2.91	3.66	1.93	14.42

SPINATION. Femora I–IV with one basodorsal spine and 3 dorsal bristles; palpal femur dorsally with 3 bristles; tarsi I–IV aspinose. Palp: femur pd1; patella p1; tibia v7; tarsus v2. Leg I: femur pd1; patella p1; tibia p1(0), v6; metatarsus v7. Leg II: femur pd1; patella p1; tibia p1, v6; metatarsus v7. Leg III: femur pd2, rd2; patella r1; tibia d1, p2, r3, v7; metatarsus d2, p3, r3, v7. Leg IV: femur pd2, rd1; patella p1, r1; tibia d1, p3, r2, v7; metatarsus d2, p4(3), r3, v8.

SPERMATHECAE. Each of paired spermathecae Y-shaped with short spermathecal trunk carrying relatively long lateral diverticulum with fairly pronounced terminal fundus ( Fig. 544 ).

SPINNERETS. See Fig. 613 . PMS: length 0.29, diameter 0.13. PLS: maximal diameter 0.59; length of basal, medial and apical segments 0.81, 0.54, 0.48; total length 1.83; apical segment triangular.

Male

Unknown.

Variation

Carapace length in females (n =6) varies from 4.37 to 6.25. Abdomen dorsally with a weak brownish chevron-like pattern that ranges from poorly discernible to almost indistinct throughout the specimens. Within the type series, only a minor variation in the structure of the spermathecae is observed, as shown in Figs 545–546 .

Ecology

All spiders were found hiding under stones in small mountain gorges covered with open low forest and shrubs.

Distribution

Kazakhstan: Northern Tien-Shan (Trans-Ili Mt Ridge, including its western spur, Zhetyzhol Mts). See Fig. 759 .

Notes

Logunov & Gromov (2012: text figure on p. 28) depicted a female belonging to this species as Raveniola sp.

Figures

Figs 54–62. Raveniola spp., ♀♀, paratypes (except 59–60), habitus, dorsal views. 54. R. ferghanensis (Zonstein, 1984) (SMNH). 55. R. kirgizica sp. nov. (SMNH). 56–57. R. mikhailovi Zonstein, 2021, from Aflatun (ZMMU) and Sary-Chelek Reserve (SMNH), respectively. 58. R. nenilini sp. nov. (SMNH). 59. R. tarabaevi sp. nov., holotype (SMNH). 60. R. virgata (Simon, 1891), ♀ from Arslanbob (SMNH). 61–62. R. vulpina sp. nov., from Sary-Chelek Reserve and Tereksai (both SMNH), respectively. Scale bars: 5.0 mm.

Figs 127–135. Raveniola spp., ♀♀, paratypes (except 132–133), cephalothorax, dorsal views. 127. R. zyuzinisp. nov. (SMNH). 128. R. ferghanensis(Zonstein, 1984) (SMNH). 129. R. kirgizicasp. nov. (SMNH). 130. R. mikhailovi Zonstein, 2021, from Sary-Chelek Reserve (SMNH). 131. R. nenilini sp. nov. (SMNH). 132. R. tarabaevi sp. nov., holotype (SMNH). 133. R. virgata (Simon, 1891), from Arslanbob (SMNH). 134–135. R. vulpina sp. nov., from Sary-Chelek Reserve and Tereksai (both SMNH), respectively. Scale bars: 2.0 mm.

Figs 184–195. Raveniola spp., ♀♀, paratypes (except 191–193), clypeus and eye tubercle, dorsal views. 184. R. pallens sp. nov. (SMNH). 185. R. zyuzini sp. nov. (SMNH). 186. R. ferghanensis (Zonstein, 1984) (SMNH). 187. R. kirgizica sp. nov. (SMNH). 188–189. R. mikhailovi Zonstein, 2021, from Aflatun (ZMMU) and Sary-Chelek Reserve (SMNH), respectively. 190. R. nenilini sp. nov. (SMNH). 191. R. tarabaevi sp. nov., holotype (SMNH). 192–193. R. virgata (Simon, 1891), ♀♀ from Arslanbob (SMNH) and Gava (SMF), respectively. 194–195. R. vulpina sp. nov., from Sary-Chelek Reserve and Tereksai, respectively (both SMNH). Scale bars: 0.5 mm.

Figs 247–255. Raveniola spp., ♀♀, paratypes (except 252–253), sternum, labium and maxillae, ventral views. 247. R. ferghanensis (Zonstein, 1984) (SMNH). 248. R. kirgizica sp. nov. (SMNH). 249–250. R. mikhailovi Zonstein, 2021, from Aflatun (ZMMU) and Karangitun (SMNH), respectively. 251. R. nenilini sp. nov. (SMNH). 252. R. tarabaevi sp. nov., holotype (SMNH). 253. R. virgata (Simon, 1891), ♀ from Arslanbob (SMNH). 254–255. R. vulpina sp. nov., from Sary-Chelek Reserve and Tereksai, respectively (both SMNH). Scale bars: 1.0 mm.

Figs 537–554. Raveniola spp., ♀♀, spermathecae, dorsal views. 537. R. kirgizica sp. nov., paratype (SMNH). 538. R. mikhailovi Zonstein, 2021, female from Chapchama Pass (SMNH). 539– 540. R. mikhailovi, paratypes from Aflatun (ZMMU) and Sary-Chelek Reserve (SMNH), respectively. 541. R. nenilini sp. nov., female from Kamchik Pass (SMNH). 542–543. R. nenilini, paratypes from Kainarsai (SMNH). 544. R. tarabaevi sp. nov., holotype (SMNH). 545. R. tarabaevi, paratype from Uzunbulak (SMNH). 546. R. tarabaevi, paratype from Turgen (SMNH). 547. R. virgata (Simon, 1891), paralectotype (MNHN 6506-B361). 548–550. R. virgata, females from surroundings of Arslanbob (SMNH). 551–552. R. vulpina sp. nov., paratypes from Tumanyak (SMNH). 553–554. R. vulpina, from Tereksai (SMNH).Abbreviations: ld = lateral diverticulum; st = spermathecal trunk. Scale bars: 0.5 mm.

Figs 611–618. Raveniola spp., ♀♀ (611, 613, 615, 617–618) and ♂♂ (612, 614, 616). Spinnerets, ventral (611–617) and subcaudal (618) views. 611. R. nenilini sp. nov., paratype from Mazarsai (SMNH). 612. R. ovchinnikovi sp. nov., holotype (SMNH). 613. R. tarabaevi sp. nov., holotype (SMNH). 614– 615. R. virgata (Fet, 1984), from vicinity of Arslanbob (SMNH). 616. R. vulpina sp. nov., holotype (SMNH). 617–618. R. vulpina, paratype from Tumanyak (SMNH). Strictly reduced PMS indicated by white arrows. Scale bars: 1.0 mm.

Figs 751–760. Distribution records of Raveniola spp. 751. R. alajensis sp. nov. (), R. cucullata sp. nov. () and R. dolosa sp. nov. (). 752. R. hirta sp. nov. (), R. ignobilis sp. nov. () and R. insolita sp. nov. (). 753. R. karategensis sp. nov. (), R. ornata sp. nov. () and R. ornatula sp. nov. (). 754. R. pamira sp. nov. (), R. sororcula sp. nov. () and R. diluta sp. nov. (). 755. R. fedotovi (Charitonov, 1946) (), R. pallens sp. nov. () and R. zyuzini sp. nov. (). 756. R. ferghanensis (Zonstein, 1984) () and R. kirgizica sp. nov. (). 757. R. kopetdaghensis (Fet, 1984) (). 758. R. mikhailovi Zonstein, 2021 () and R. nenilini sp. nov. (). 759. R. ovchinnikovi sp. nov. () and R. tarabaevi sp. nov. (). 760. R. virgata (Simon, 1891) (;? = the questionable type locality) and R. vulpina sp. nov. (). Country abbreviations as in Figs 747–750.