Quedius (Raphirus) hebes, Assing, 2017

Assing, Volker, 2017, On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae), Linzer biologische Beiträge 49 (1), pp. 207-228 : 219-221

Quedius (Raphirus) hebes nov.sp. (Figs 10-13, 18, 25, 34-36, 46-47, 54, 58, Map 1 )

T y p e m a t e r i a l: Holotype 3: "N 37°35'23 E 034°53'05, TR Adana Kamisli , 1350 m, 18.4.2011, Brachat & Meybohm (15) / Holotypus 3 Quedius hebes sp.n., det. V. Assing 2017" (cAss) . Paratypes: 1♀ [slightly teneral]: "TR - Adana (21), ca. 11 km nö Feke, sw Cumhurlu, 710 m, 21.IV.2011, N37°53'11, E35°59'37'', leg. Meybohm & Brachat " (cAss); 13 [slightly teneral]: "TR. - Mersin [51], road to Arslanköy, 5 km SE Aladag, 700 m, 36°54'45N, 34°31'44E, 10.V.2004, C. Besuchet " (cAss); 13: "TR - Niğde (9), Umg. Çiftehan, ö. Maden, 1200 m, 17.IV.2011, N37°28'52, E34°41'36, leg. Meybohm & Brachat " (cAss); 1♀: "TR - Niğde (11), Umg. Çiftehan, ö. Maden, 1870 m, 17.IV.2011, N37°27'08, E34°37'05, leg. Meybohm & Brachat " (cAss); 1♀: "N 36°36'59 E 32°24'36 (36), TR Antalya, Karapinar, Alanya → Taşkent, 1090 m, 25.IV.2008, leg. Meybohm & Brachat " (cAss); 1♀: "N 36°07'16 E 32°34'11 (33), TR Antalya Kalandere , Anamur - Gazipasa , 40 m, Brachat & Meybohm 24.4.2008 " (cAss); 13: "N36°8', E32°46', Türkei Umg. Anamur, Kösekbasi 150 m, Meybohm 18.5.2000 " ( ZMC); 13 [teneral]: "TR. - Antalya, Nr. 7, 60 km SSW Antalya, Çiralı , 220 m, 36°25'54N, 30°25'59E, 25.III.2002, V. Assing " ( ZMC); 13: "TR [6] - Afyon, Sultan Dağları , 15 km SE Çay, 38°31'39''N, 31°11'07''E, 1430 m, oak forest, 18.IV.2011, P. Wunderle " (cWun); 13: "TR - Muğla, 19, 600 m, Gölgeli Dağları , 20 km NE Köyceğiz, below Ağla , 37°01'20N, 28°44'27E, 6.X.2002, V. Assing " (cAss); 13: "TR - Muğla, 20, 1000 m, 20 km NNE Fethiye, oak forest, N-slope, 36°47'27N, 29°11'29E, 8.X.2002, V. Assing " (cAss) ; 1♀: "Bürücek, Toros, 29-31.7.47, Exp. N Mus. ČSR / Quedius (Raph.) coloratus Fauv. , det. P. Štourač 2000" ( NMP) .

E t y m o l o g y: The specific epithet (Latin, adjective: obtuse) alludes to the apically convex median lobe of the aedeagus (lateral view).

D e s c r i p t i o n: Measurements (in mm) and ratios (range, arithmetic mean) (n = 13): TL: 8.6-12.9, 10.7; HL: 1.21-1.48, 1.34; HW: 1.44-1.81, 1.63; PL: 1.67-2.04, 1.86; PW: 1.83-2.25, 2.03; EL: 1.09-1.32, 1.21; HTiL: 1.44-1.83, 1.63; HTaL: 1.16-1.37, 1.27; ML: 1.32-1.46, 1.39; HW/HL: 1.17-1.26, 1.21; PW/HW: 1.21-1.29, 1.25; PL/PW: 0.89- 0.94, 0.92; EL/PL: 0.61-0.68, 0.65; HTiL/HTaL: 1.20-1.40, 1.29. Female of slightly larger average size than male.

External and sexual characters similar to those of Q. coloratus , except as follows:

Legs somewhat darker, dark-reddish to reddish-brown; antennae dark-brown with the basal three antennomeres reddish. Head (Figs 11-12) with additional 1-3 smaller punc- tures near posterior frontal puncture and postero-dorsal margin of eye on either side (occasionally, these additional punctures may be completely absent on one side). Body on average larger ( Fig. 58 ). Elytra (Fig. 10) on average slightly shorter (see ratio EL/PL). Protarsomeres more strongly dilated both in male and in female (Fig. 13).

3: aedeagus (Figs 25, 34-36, 46-47, 54) larger than in Q. coloratus (see measuments ML and Fig. 58 ; little overlap); median lobe apically stouter, not acute but convex, and somewhat curved dorsad in lateral view, broader in ventral view; paramere (Figs 36, 54) broader and with approximately 35 peg-setae arranged in a pair of distinctly separated clusters.

♀: tergite X (Fig. 18) apically with much smaller and less deep posterior excision; styli of segments IX-X shorter and less slender (similar to Fig. 22).

HL*HW+PL*PW

C o m p a r a t i v e n o t e s: Quedius hebes is reliably distinguished from Q. coloratus particularly by the presence of additional punctures near the posterior frontal puncture on the head, by the different morphology of the aedeagus (see above), and by the much smaller posterior excision of the female tergite X. For characters separating it from the geographically close Q. spiculatus see the following section.

D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: Thedistributionrangesacross southern Turkey from Muğla in the west to Adana in the east ( Map 1 ), where it appears to parapatrically border on that of Q. coloratus . A previous record of Q. coloratus from Mersin ( SMETANA 1967) refers to Q. hebes (see the paratype from "Bürücek"); the same probably applies to a record of Q. coloratus from Isparta (JAPOSHVILI & ANLAŞ 2011). The type specimens were found under stones or sifted from leaf litter at altitudes of 40- 1870 m. All of them were collected as singletons, suggesting that the records are accidental and that this species is not an inhabitant of the leaf litter, but of a subterranean habitat. Three teneral specimens were collected in March, April, and May.

Figures

Map 1: Distributions of Quedius hellenicus (black diamonds), Q. sp. 2 (white stars), Q. carpathius (black stars), Q. spiculatus (white triangles), Q. hebes (white circles), and Q. coloratus (black circles) in the East Mediterranean.

Fig. 58: Length of median lobe of aedeagus in relation to body size, given as an approximation of the combined area of head and pronotum, in Quedius coloratus and allied species.