Pterynotus (Pterymarchia) pelouatensis, Merle & Pacaud & Ledon & Goret, 2024

Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel & Goret, Bernard, 2024, New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe, Geodiversitas 46 (15), pp. 495-551 : 526

Pterynotus (Pterymarchia) pelouatensis n. sp.

( Figs 18 E-H; 27G, H)

urn:lsid:zoobank.org:act:4852FE61-DAEE-4316-B3D0-A7A7162A231E

Pterynotus (Pterymarchia) sp. – Merle et al. 2011: 424, pl. 90, fig.7a, b.

TYPE MATERIAL. — Holotype. France • Aquitaine Basin, Gironde, Saucats ( Peloua ); Faluns de Saucats ; Early Miocene ( lower Burdigalian ); MNHN.F.J09945 (Neuville coll.), figured specimen in Merle et al. (2011: 424, pl. 90, fig. 7), H: 21 mm ( Figs 18E, F ; 27G ). Paratype. France • 1 spm; same as for the holotype; MNHN.F.J17632 (Neuville coll.) ( Figs 18G, H ; 27H ).

ETYMOLOGY. — From the type locality, Peloua, Gironde, France.

TYPE HORIZON. — Faluns de Saucats, Early Miocene (lower Burdigalian), see Parize et al. (2008: 396).

TYPE LOCALITY. — France, Aquitaine Basin, Gironde, Saucats (Peloua), seeParize et al. (2008: fig. 1).

DISTRIBUTION. — Only known from the type locality.

DESCRIPTION

Two smooth protoconch whorls preserved delimited by sinusigera scar (protoconch tip not preserved) (see Merle et al. 2011: 122, text-fig. 46E). Teleoconch up to 21 mm in height, up to 12.5 mm in width, biconic in profile, composed of five whorls. Moderately high spire with subcarinate whorls. Last whorl up to 73% of total length. Apical angle 64°. Spiral sculpture with marked primary cords. First whorl: appearance of P1 and P2; second whorl: appearance of IP; third whorl: appearance of abis, adis and s2. Fourth whorl: no change or appearance of threads. Fifth whorl: IP, adis and abis, P1 to P6, s1 to s6; s3 and s4 more developed than other secondary cords; P6 slightly atrophied, on base of convex part of whorl; ADP, MP, ads and ms; ABP missing. Surface of shell with numerous threads appearing on fourth whorl. Axial sculpture with thick varices. On first whorl: 10-11 lamellose varices; on second whorl: 10-12 varices; on third whorl: 11-12 varices; on fourth whorl: 10-12 varices; on fifth whorl 8-10 varices. From second whorl to fifth whorl: thick varices. No evidence of intervarices on last whorl. From third to last whorl numerous, fine microlamellae forming scabrous microsculpture covering entire surface. Ovate aperture up to 31% of diameter and up to 72% of length of last whorl (including siphonal canal). Columellar lip adherent posteriorly, becoming progressively more erect anteriorly. One or two weak columellar tubercles. Parietal lip adherent. Outer lip with strong internal denticles: ID, D2 to D6 present, D1 missing; D4 split, D6 poorly developed. Pseudoumbilicus moderately large. Siphonal canal open, up to 43% of aperture length, slightly dorsally recurved.

COMPARISONS

The European Neogene contains two species of Pterynotus (Pterymarchia) : P. (P.) pelouatensis n. sp. and P. (P.) sandbergeri (Hörnes, 1856) n. comb. from the Langhian of Paratethys ( Austria, Hungary, Romania, Poland) and the Serravallian of Turkey. The type material of P. (P.) sandbergeri is housed in Naturhistorisches Museum of Wien and contains the figured syntype of the species (Hörnes 1856: pl. 51, fig. 5a, b, NHMW 1855/0045/0435). Here, we designate this specimen as the lectotype ( Fig. 18I, J ). Pterynotus (P.) sandbergeri was often attributed to Murexsul Iredale, 1915 ( Landau et al. 2013; Kovács et al. 2018; Kovács 2019). Although they share several similarities (biconic shells, internal denticles, columellar denticles), members of Murexsul have a more rounded aperture, their internal denticles are close together, ID is weaker and their siphonal canal is narrower. The spires of P. (P.) pelouatensis n. sp. and the lectotype of P. (P.) sandbergeri are very similar, but their last whorls are very different. The lectotype of P. (P.) sandbergeri has a larger aperture, a surface covered by numerous threads and atrophy of P2, P5 and P6; P4 is more developed and D6 is missing. The differences between the primary cords are great, but in other specimens attributed from P. (P.) sandbergeri ( Fig. 18K, L ), the atrophy of P2, P5 and P6 is less strong than in the lectotype from Austria (see Landau et al. 2013; Kovács et al. 2018; Kovács 2019). Compared to extant P. ( Pterymarchia ), P. (P.) pelouatensis n. sp. and P. (P.) sandbergeri share greatest morphological similarities with P. (P.) martinetana (Röding, 1798) and particularly with specimens from the Red Sea ( Merle et al. 2011: pl. 92, figs 11, 12), but P. (P.) sandbergeri differs by having develop cord spines on P1, P2, P4, P5, ADP and MP.

Figures

FIG. 18. — Spiral cords and internal denticles of the outer lip in Pterynotus (Pterymarchia) gaasensis n. sp. (A-D), P. (P.) pelouatensis n. sp. (E-H) and P. (P) sandbergeri (HÖrnes,1856) (I-L): A, B, holotype,MNHN.F.A32492 (Merle coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); C, D, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos);E, F, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); G, H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); K, L, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau.

FIG. 27. — New species of Timbellus De Gregorio, 1885, Pterynotus Swainson, 1833 (Pterymarchia Houart, 1995), Textiliomurex Merle, 2011, Ponderia Houart, 1986 and Flexopteron Shuto, 1969: A-C, Timbellus submicropterus n. sp.: A, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, France, Lutetian; B, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, France, Lutetian; C, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville; D, T. cf. submicropterus n. sp., MNHN.F.A90557 (Brébion coll.), Saint-Aignan-Grand-Lieu (Pierre-Aiguë), France, Bartonian; E, F, Pterynotus (Pterymarchia) gaasensis n. sp.: E, holotype, MNHN.F.A32492 (Merle coll.), Gaas (Espibos), France, Rupelian; F, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos); G, H, P. (P.) pelouatensis n. sp.: G, holotype, MNHN.F.J09945, Saucats (Peloua), France, Burdigalian; H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, Textiliomurex chodmonensis n. sp.: I, holotype, MNHN.F.A81144 (Pacaud coll.), Chaumont-en-Vexin (Carrière Darcy), France, Lutetian; J, paratype, MNHN.F.A81145 (Pons coll.), Chaumont-en- Vexin (Carrière Darcy); K, L, Ponderia remyi n. sp.: K, holotype, MNHN.F.A90548 (Remy coll.), Fresville, Lutetian; L, paratype, MNHN.F.A90555, Fresville (Remy coll.); M, N, Flexopteron constantinense n. sp.: M, paratype, MNHN.F.A77761 (MNHN coll.), Carentan-les-Marais (Cauvin), France, Priabonian; N, holotype, MNHN.F.A77760 (MNHN coll.), Carentan-les-Marais (Cauvin). Scale bars: A-D, I-L, 10 mm; E-H, N, 5 mm; M, 3 mm. Credits: A, B, D-F, N, P. Loubry (MNHN/ CNRS); C, E. Robert (UCBL).