Pterostichus (Steropanus) pseudoglymmiger, Fairmaire, 1889

Fedorenko, D. N., 2020, New taxa of Pterostichini (Coleoptera: Carabidae) from Vietnam, Russian Entomological Journal 29 (1), pp. 38-52 : 44-45

Pterostichus (Steropanus) pseudoglymmiger Fedorenko, sp.n.

Figs 15 , 21–22 , 25, 28 .

MATERIAL. Holotype ♂ ( ZMMU), labelled: ‘N-Vietnam, 40 km WNW of Lao Cai, env. Y Ty , Bat Xat N[ational]P[ark], 22°37´– 37.6´N/ 103°37.3´–38.3´E, h= 1600–1900 m, 16–21.X.2018, leg. D.Fedorenko’ . Paratypes 2♀♀ ( SIEE), same data, except for 22°37´36´´N / 103°37´32´´E, h= 1850 m, 4–14.VI.2019 .

DIAGNOSIS. This species is similar to P. glymmiger in appearance, being distinctive chiefly in having tarsomere 5 setose ventrally, pronotum without basal sulci, and elytral basal ridge slightly shortened. The other consubgeners with ventrally setose tarsomere 5 are distributed much more westerly, but P. obliteratus Fedorenko, 2018 only occurs in Vietnam. It is smaller, with BL<14 mm, colouration black, pronotal sides convex towards base, with basal angles obtuse and rounded, elytral basal ridge entire, US being 18–19 in number, protibia less strongly armed, with 3–4 apical spinules along posterolateral edge, mesotibial brush one-row, etc. Metatibia with one external seta (vs. no seta) differentiates the new species from all eastern species of the subgenus.

DESCRIPTION. BL 15.3–16.3 mm. Body ( Fig. 15 ) shiny black, elytra slightly violaceous along sides. Legs, antennae and palps rather dark reddish brown, femora infuscated to nearly black but extreme apices. Dorsal microsculpture fine-meshed; isodiametric, obliterate or hardly traceable on head; superficial yet distinct over pronotum and elytra, being barely or moderately transverse, respectively; pronotal lateral groove and reflexed lateral margin of elytra with nearly granulate isodiametric (pronotum) or barely longitudinal (elytra) meshes. Head densely micropunctate, pronotal punctation sparser and finer, with admixture of some larger punctures here and there; elytra very sparsely micropunctate.

Head: Eyes lateral, hemispherical, medium-sized; gena short, meeting neck at obtuse angle. Frontal sulci short, deep, crenulate, diverging basad, not reaching level of anterior supra-ocular seta. Supra-ocular groove deep, slightly incurved anteriorly. Labrum subsinuate apically, sexsetose, inner four setae closely set. Antennae almost reaching (♀) or slightly extended beyond (♂) pronotal base.

Penultimate labial palpomere trisetose (ventro-apical seta missing), except for a minute outer apical seta. Terminal labial palpomere more triangular in male than in female, 0.81 or 0.56 times as wide at apex as long at inner margin, respectively.

Pronotum subcordate, PW / PL 1.30–1.35 (1.33, n=3), PW /HW 1.62–1.69 (1.66), broadest two fifths from apex, PLw/ PL 0.39–0.43 (0.41); sides evenly rounded in apical two thirds, indistinctly sinuate behind. Base sinuate medially, oblique towards obtuse and blunt basal angles, nearly as wide as apex, PB/PA 1.03–1.05 (1.04). Apex deeply sinuate, with almost indistinct convexity towards subrectangular and blunt apical angles. Apical bead fine, obliterate medially; lateral bead entire and fairly thick; lateral groove impunctate, narrow and deep, slightly extended inside base into a more or less distinct line. Median line very fine and shallow, slightly deeper near base, obliterate basally and apically. Disc smooth, basal sulci and transverse impressions, anterior and posterior, missing. Lateral setae, anterior and posterior, in lateral groove.

Elytra connate, oblong, broadest at middle, EW/EL 1.62– 1.65 (1.64), EW/ PW 1.12–1.18 (1.15), apices rounded combined, both preapical sinuation and internal plica slight. Base slightly oblique, humeri rather widely rounded, humeral tooth vestigial, being only traceable as a denticle between lateral bead and basal ridge in lateral view; basal ridge almost reaching stria 1. Striae deep, impunctate, those 2–6 adjoining basal ridge; parascutellar striole missing. Intervals slightly convex, nearly flat a third from apex, all merging apically but intervals 4–6. Reflexed lateral margin narrow. Parascutellar setigerous pore at base of stria 2, discal setae missing, stria 7 with one preapical seta, anterior, distant far from apex; USS: 24–25.

Underside. Prosternum almost smooth along middle. Prosternal process in ventral view truncate, with fairly sharp angles; in lateral view subrectangular; inclination wide, flat or barely concave, slightly constricted at middle, with lateral edges faintly beaded in dorsal half; in lateral view nearly vertical and barely concave. Abdominal lateral bead distinct and entire; sternite VII similar in both sexes. Mesepisternum very finely and densely punctate, metepisternum less so or smooth, abdominal sternites II–III indistinctly punctate or shallowly rugulose-punctate; ventral side otherwise smooth.

Legs: profemur posteriorly bisetose (basal seta missing), protibia distinctly dilated apicad, with 5–7 spinules at posterolateral edge. Tarsi laterally neither carinate nor sulcate; Tarsomere 5 with 2–3 pairs of ventral setae. Protarsomeres 1– 3 dilated and biseriately squamose ventrally in male; strongly dentate at apical angles in female. Mesotibia with 2–3 additional setae just above one-row apical setal brush. Metatibia with one external seta, preapical.

Aedeagus ( Figs 21–22 , 25, 28 ): Median lobe geniculate. Apex narrow, subtriangular, apically rounded. Right paramere moderately long, boomerang-like, sinuate ventrally before a narrow apex. Internal sac without distinct apical sclerite (which may be a result of slightly teneral condition of the male specimen).

DISTRIBUTION. Known from the type locality only.

NAME. Refers to the similarity of this species to another consubgener, P. glymmiger .

HABITATS AND HABITS. All specimens were hand collected at the edge of a disturbed cloudy forest.

Figures

Figs 14–16. Dorsal habitus, holotypes: 14 — Pterostichus (Steropanus) securipenis sp.n; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n. Рис. 14–16. Габитус дорЗально, голотипы: 14 — Pterostichus (Steropanus) securipenis sp.n.; 15 — P. (S.) pseudoglymmiger sp.n.; 16 — P. (S.) fossifrons sp.n.

Figs 17–22. Aedeagus of Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 17–20 — median lobe; 17, 19, 21 — left lateral aspect; 18, 20, 22 — right lateral aspect. Scale bars: 1 mm. Рис. 17–22. Эдеагус Pterostichus (Steropanus) spp.: 17–18 — P. (S.) fossifrons sp.n.; 19–20 — P. (S.) securipenis sp.n.; 21–22 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 17–20 — среднЯЯ долЯ; 17, 19, 21 — слева; 18, 20, 22 — справа. МасШтаб: 1 мм.

Figs 23–29. Aedeagus of Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (apical margin slightly damaged); 25, 28 — P. (S.) pseudoglymmiger sp.n., median lobe with everted and inflated internal sac; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — median lobe, ventral aspect; 26–28 — inside of right paramere. Scale bars: 1 mm. Рис. 23–29. Эдеагус Pterostichus spp.: 23, 26 — P. (Steropanus) fossifrons sp.n.; 24, 27 — P. (S.) securipenis sp.n. (апикальный край слегка обломан); 25, 28 — P. (S.) pseudoglymmiger sp.n., среднЯЯ долЯ с вывернутым и раЗдутым внутренним меШком; 29 — P. (Vietosteropus) kuznetsovi sp.n.; 23–25 — среднЯЯ долЯ? вентрально; 26–28 — праваЯ парамера, иЗнутри. МасШтаб: 1 мм.