Pterostichus laocaiensis Fedorenko, 2019

Fedorenko, D. N., 2019, New species of Pterostichus subgenus Circinatus (Coleoptera: Carabidae: Pterostichini) from Vietnam, Russian Entomological Journal 28 (4), pp. 358-369 : 368-369

4. Pterostichus laocaiensis Fedorenko , sp.n.

Figs 7 , 14–15 , 20 , 26–27 , 41–42 .

MATERIAL. Holotype ♂ ( ZMMU) and paratype ♂ ( SIEE), labelled: ‘N Vietnam, Lao Cai Prov., Hoang Lien Son Mt. Ridge, env. Fansipan [=Phahg Xi Pang] Mt , Tram Ton , h=1950–2100 [m], 15–30.VII.2007, leg. D. Fedorenko’.

DIAGNOSIS. As for P. phangxipang sp.n., except for pronotal posterolateral seta missing, body slender, pronotum barely longer than wide and subsinuate on sides in basal half, pronotal and elytral microsculpture consisting of dense transverse lines; elytral basolateral bead missing, parascutellar striole entire; meso- and metatarsomeres 1–3 carinate and sulcate laterally; distal femoral seta missing. For additional differences see description.

DESCRIPTION. Body slender ( Fig. 7 ), BL 13–13.2 mm.

Head: Frontal sulci straight and slightly diverging. Labrum sinuate apically.

Pronotum very slender, broadest two fifths from apex, barely longer than wide, PW/PL 0.93–0.97, PW/HW 1.28– 1.35, with sides subsinuate before base. Base subconvex to truncate, PB/PA 0.81–0.85; apex subconvex to truncate between slightly projecting apical angles, with apical bead entire or interrupted just medially. Disc with indistinct basal and apical transverse impressions; median line fine and nearly entire, ill-defined basally and almost reaching apex; sublateral line missing. Basal sulci γ- shaped combined, inner long, running in basal two fifths and S-shaped, anteriorly parallel, then diverging and afterwards incurved just in front of base; outer sulcus vestigial, very short and shallow, running close to lateral bead. Lateral bead obliterate just in front of basal angle.

Elytra oblong, EW/EL 1.78–1.80, EW/PW 1.26–1.33, broadest three fifths from base, with an indistinct preapical sinuation, apices truncate combined or rounded separately each. Base long, with lateral bead obliterate; humeral tooth vestigial. Intervals convex. Discal seta d2 about 3/5 distant from base. USS: 16–17.

Prosternal process with posterior declivity convex, narrow and subcostate at medial constriction.

Aedeagus ( Figs 14–15 , 20 , 26–27 ): median lobe ventrally carinate in basal half, with a convex ridge in middle third of right ventrolateral margin; apex in dorsal view fairly long, parallel-sided, rounded apically, with a shallow groove that is oblique to the right. Everted and inflated internal sac ( Figs 41–42 ) bent to the left and then ventrad.

NAME. Refers to Lao Cai Province, Vietnam, from where the species is described .

DISTRIBUTION. Known from the type locality only, Mt. Phang Xi Pang , northern Vietnam .

HABITATS AND HABITS. The species was hand collected under bark of a not quite fallen dead tree in a cloudy forest.

Acknowledgements. This study was funded by the Presidium of the Russian academy of sciences, Program No.41 “Biodiversity of natural systems and biological resources of Russia ”.

Figures

Figs 4–7. Dorsal habitus: 4 — Pterostichus phangxipang sp.n., female paratype; 5 — P. hoanglien sp.n., female paratype; 6 — Pterostichus batxatensis sp.n., holotype; 7 — P. laocaiensis sp.n., male paratype. Рис. 4–7. Габитус дорЗально: 4 — Pterostichus phangxipang sp.n., паратип самка; 5 — P. hoanglien sp.n., паратип самка; 6 — Pterostichus batxatensis sp.n., голотип; 7 — P. laocaiensis sp.n., паратип самец.

Figs 8–15. Median lobe of aedeagus: 8–9 — Pterostichus phangxipang sp.n.; 10–11 — P. hoanglien sp.n.; 12–13 — P. batxatensis sp.n.; 14–15 — P. laocaiensis sp.n.; 8, 10, 12, 14 — left view; 9, 11, 13, 15 — right view. Scale bars: 1 mm. Рис. 8–15. СреднЯЯ долЯ Эдеагуса: 8–9 — Pterostichus phangxipang sp.n.; 10–11 — P. hoanglien sp.n.; 12–13 — P. batxatensis sp.n.; 14–15 — P. laocaiensis sp.n.; 8, 10, 12, 14 — слева; 9, 11, 13, 15 — справа. Масштаб: 1 мм.

Figs 16–20. Median lobe of aedeagus: 16 — Pterostichus phangxipang sp.n.; 17–18 — P. batxatensis sp.n.; 19 — P. hoanglien sp.n.; 20 — P. laocaiensis sp.n.; 16–17, 20 — dorsal view; 18–19 — ventral view. Scale bar: 1 mm. Рис. 45–58. СреднЯЯ долЯ Эдеагуса: 16 — Pterostichus phangxipang sp.n.; 17–18 — P. batxatensis sp.n.; 19 — P. hoanglien sp.n.; 20 — P. laocaiensis sp.n.; 16–17, 20 — дорЗально; 18–19 — вентрально. Масштаб: 1 мм.

Figs 21–30. Right paramere: 21–22 — Pterostichus phangxipang sp.n.; 23–24 — P. batxatensis sp.n.; 25 — P. phiaoc sp.n.; 26–27 — P. laocaiensis sp.n.; 28–30 — P. hoanglien sp.n.; 21, 23, 25, 26, 28–29 — left view; 22, 24, 27, 30 — ventral view. Scale bar: 1 mm. Рис. 21–30. ПраваЯ парамера: 21–22 — Pterostichus phangxipang sp.n.; 23–24 — P. batxatensis sp.n.; 25 — P. phiaoc sp.n.; 26– 27 — P. laocaiensis sp.n.; 28–30 — P. hoanglien sp.n.; 21, 23, 25, 26, 28–29 — слева; 22, 24, 27, 30 — вентрально. Масштаб: 1 мм.

Figs 37–42. Everted and inflated internal sac of aedeagus: 37–38 — Pterostichus phangxipang sp.n.; 39–40 — P. hoanglien sp.n.; 41– 42 — P. laocaiensis sp.n.; 37, 41 — left view; 38 — dorso-apical view; 39 — dorsal view; 40 — left dorso-apical view; 42 — left ventrolateral view. Scale bar: 1 mm. Рис. 37–42. Вывернутый и надутый внутренний мешок Эдеагуса: 37–38 — Pterostichus phangxipang sp.n.; 39–40 — P. hoanglien sp.n.; 41–42 — P. laocaiensis sp.n.; 37, 41 — слева; 38 — дорЗоапикально; 39 — дорЗально; 40 — дорЗоапикально слева; 42 — вентрально слева. Масштаб: 1 мм.