Pterostichus batxatensis Fedorenko, 2019

Fedorenko, D. N., 2019, New species of Pterostichus subgenus Circinatus (Coleoptera: Carabidae: Pterostichini) from Vietnam, Russian Entomological Journal 28 (4), pp. 358-369 : 368

6. Pterostichus batxatensis Fedorenko , sp.n.

Figs 6 , 12–13 , 17–18 , 23–24 , 34–36 .

MATERIAL. Holotype ♂ ( ZMMU) and paratypes, ♂, ♀ ( SIEE), labelled: ‘N-Vietnam, 40 km WNW of Lao Cai, env. Y Ty, Bat Xat Natn. Park, h= 1600–1900 m, 22°37´–37.6´N/ 103°37.3´–38.3´E, 16–21.X.2018, leg. D. Fedorenko’; 3♂♂, 2♀♀, same data, except for 22°36´31´´N / 103°37´23´´E, h= 2000–2100 m, 4–14.VI.2019.

DIAGNOSIS. As for P. phangxipang sp.n., except for pronotal posterolateral seta missing and aedeagus distinctive. For additional differences see description.

DESCRIPTION. Body ( Fig. 6 ), BL 12.4–13.5 mm.

Pronotum: PW/PL 1.04–1.13 (mean 1.08, n=3), PW/HW 1.40–1.48 (1.44), PB/PA 0.79–0.88 (0.82). Apical bead entire to interrupted just medially.

Elytra more rounded at humeri than behind, EW/EL 1.59–1.64 (1.62), EW/PW 1.14– 1.18 (1.16); apices truncate combined to rounded separately each. Humeral tooth indistinct. Parascutellar striole missing or vestigial (entire unilaterally in the holotype). USS : 16–17.

Legs: distal metafemoral seta vestigial or missing.

Aedeagus ( Figs 12–13 , 17–18 , 23–24 ): median lobe with apex widely rounded. Right paramere short (lost through mounting). Everted and inflated internal sac ( Figs 34–36 ) dorsal and spiral, combined with a large horn emerged from dorsal membrane of median lobe.

NAME. Refers to the type locality of the species .

DISTRIBUTION. Known from the type locality only, the Bat Xat National Park , northern Vietnam .

HABITATS AND HABITS. All specimens were hand collected under fragments of rotten tree trunks on the ground or under bark of rotten tree trunks tilted at an angle in a cloudy mixed forest.

Figures

Figs 4–7. Dorsal habitus: 4 — Pterostichus phangxipang sp.n., female paratype; 5 — P. hoanglien sp.n., female paratype; 6 — Pterostichus batxatensis sp.n., holotype; 7 — P. laocaiensis sp.n., male paratype. Рис. 4–7. Габитус дорЗально: 4 — Pterostichus phangxipang sp.n., паратип самка; 5 — P. hoanglien sp.n., паратип самка; 6 — Pterostichus batxatensis sp.n., голотип; 7 — P. laocaiensis sp.n., паратип самец.

Figs 8–15. Median lobe of aedeagus: 8–9 — Pterostichus phangxipang sp.n.; 10–11 — P. hoanglien sp.n.; 12–13 — P. batxatensis sp.n.; 14–15 — P. laocaiensis sp.n.; 8, 10, 12, 14 — left view; 9, 11, 13, 15 — right view. Scale bars: 1 mm. Рис. 8–15. СреднЯЯ долЯ Эдеагуса: 8–9 — Pterostichus phangxipang sp.n.; 10–11 — P. hoanglien sp.n.; 12–13 — P. batxatensis sp.n.; 14–15 — P. laocaiensis sp.n.; 8, 10, 12, 14 — слева; 9, 11, 13, 15 — справа. Масштаб: 1 мм.

Figs 16–20. Median lobe of aedeagus: 16 — Pterostichus phangxipang sp.n.; 17–18 — P. batxatensis sp.n.; 19 — P. hoanglien sp.n.; 20 — P. laocaiensis sp.n.; 16–17, 20 — dorsal view; 18–19 — ventral view. Scale bar: 1 mm. Рис. 45–58. СреднЯЯ долЯ Эдеагуса: 16 — Pterostichus phangxipang sp.n.; 17–18 — P. batxatensis sp.n.; 19 — P. hoanglien sp.n.; 20 — P. laocaiensis sp.n.; 16–17, 20 — дорЗально; 18–19 — вентрально. Масштаб: 1 мм.

Figs 21–30. Right paramere: 21–22 — Pterostichus phangxipang sp.n.; 23–24 — P. batxatensis sp.n.; 25 — P. phiaoc sp.n.; 26–27 — P. laocaiensis sp.n.; 28–30 — P. hoanglien sp.n.; 21, 23, 25, 26, 28–29 — left view; 22, 24, 27, 30 — ventral view. Scale bar: 1 mm. Рис. 21–30. ПраваЯ парамера: 21–22 — Pterostichus phangxipang sp.n.; 23–24 — P. batxatensis sp.n.; 25 — P. phiaoc sp.n.; 26– 27 — P. laocaiensis sp.n.; 28–30 — P. hoanglien sp.n.; 21, 23, 25, 26, 28–29 — слева; 22, 24, 27, 30 — вентрально. Масштаб: 1 мм.

Figs 31–36. Everted and inflated internal sac of aedeagus: 31–33 — Pterostichus phiaoc sp.n.; 34–36 — P. batxatensis sp.n.; 31, 34 — left view; 32, 35 — dorsal view; 33, 36 — right view. Scale bar: 1 mm. Рис. 31–36. Вывернутый и надутый внутренний мешок Эдеагуса: 31–33 — Pterostichus phiaoc sp.n.; 34–36 — P. batxatensis sp.n.; 31, 34 — слева; 32, 35 — дорЗально; 33, 36 — справа. Масштаб: 1 мм.