Ponerinae

K, Julia Baumann, 2024, An update on the knowledge about scutacarid mites as phoronts and inquilines, Acarologia 64 (1), pp. 3-17 : 14-17

Ponerinae

Brachyponera * chinensis * Scutacarus sp. ; Japan ; p; Lopes et al. (2015), citing Maruyama et al. (2013); attached to the head of the ant Ponera pennsylvanica * two not further identified Imparipes species ; USA ; p; Uppstrom (2010) one not further identified Scutacarus species ; USA ; p; Uppstrom (2010)

VI. SCUTACARID SPECIES ASSOCIATED WITH FORMICINAE (HEXAPODA, ECTOGNATHA, ACULEATA, FORMICIDAE )

Anoplolepsis * sp.

Scutacarus olszanowskii Khaustov et al., 2020 ; South Africa ; p; Khaustov et al. (2020); ants in an old nest of Trinervitermes trinervoides termites

Scutacarus variabilis Khaustov et al., 2020 ; South Africa ; p; Khaustov et al. (2020); ants in an old nest of Trinervitermes trinervoides termites

Camponotus sp.

Imparipes (I.) ignotus Khaustov & Chydyrov, 2004 ; Iran ; p; Navabi et al. (2018a)

Imparipes (I.) malus Khaustov, 2008 ; Iran ; p; Navabi et al. (2018a)

Camponotus chromaiodes *

Scutacarus sp. ; USA ; p; Uppstrom (2010)

Camponotus cingulatus *

Scutacarus cingulatus Silva, Khaustov & Oliveira, 2019 ; Brazil ; ip; Silva et al. (2019); females found phoretic on ants, males and larvae (but no females) found in the nest

Camponotus distinguendus *

Scutacarus camponoti Khaustov 2017 ; Chile ; ip; Khaustov (2017a); males found in the nest Camponotus oasium *

Scutacarus pseudoapodemi Navabi & Hajiqanbar, 2018 ; Iran ; p; Navabi et al. (2018a)

Imparipes (I.) morosus Khaustov, 2008 ; Iran ; p; Navabi et al. (2018a)

Cataglyphis sp.

Scutacarus subquadratus Khaustov & Chydyrov, 2004 ; Iran ; p; Sobhi et al. (2017a)

Formica sp.

Scutacarus avarus Khaustov, 2008 ; Iran ; p; Navabi et al. (2018a)

Thaumatopelvis lorasiacus Navabi & Hajiqanbar, 2018 ; Iran ; p; Navabi et al. (2018b)

Formica spp.

Imparipes sp. ; USA ; p; Uppstrom 2010

Imparipes (I.) obsoletus Rack, 1966 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Imparipes (I.) nescius Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Imparipes (I.) robustus Karafiat, 1959 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus atypicus Karafiat, 1959 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016); attached to the anterior surface of ant abdomen

3 not nearer identified Scutacarus spp. ; USA ; p; Uppstrom (2010)

Scutacarus carsticus Mahunka & Mahunka-Papp, 1980 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Scutacarus pilatus Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus rotundus (Berlese, 1903) ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus subterraneus (Oudemans, 1913) ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Formica fusca

Scutacarus karafiati Khaustov, 2015 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Formica rufa

Scutacarus (tackei?) suborbiculatus Rack, 1964; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)* Formica sanguinea

Scutacarus pseudospinosus Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Lasiophanes * picinus *

Scutacarus neotropicus Khaustov, 2017 ; Chile ; p; Khaustov (2017a)

Lasius sp.

Imparipes (I.) hajiqanbari Rahiminejad et al., 2022 ; Iran ; p; Rahiminejad et al. (2022); found on alates

Lasius spp.

5 not nearer identified

Imparipes spp. ; USA ; p; Uppstrom (2010)

Imparipes (I.) histricinus Berlese, 1903 ; Iran ; p; Rahiminejad et al. (2022)

Imparipes (I.) lituanicus Lazauskene & Sevastianov, 1974 ; Iran ; p; Rahiminejad et al. (2022); new record as phoretic species (so far only found in forest litter)

Imparipes (I.) obsoletus Rack, 1966 ; Russia, Iran ; p; Khaustov & Tolstikov (2016), Rahiminejad et al. (2022)

Imparipes (I.) sklyari Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

17 not nearer identified

Scutacarus spp. ; USA ; p; Uppstrom (2010)

Scutacarus heterotrichus Khaustov, 2015 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus lasiophilus Khaustov, 2015 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016); on lateral parts of ant abdomen

Scutacarus longisetus (Berlese, 1903) ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus pseudospinosus Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016); attached to tibia of leg I of Lasius niger

Scutacarus spinosus Štorkán, 1936 ¸ Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Lasius alienus

Diveripes sp.; USA ; p; Uppstrom (2010); attached to the ant head

Pygmodispus sp. ; USA ; p; Uppstrom (2010); attached to the ant head

Lasius flavoniger *

Imparipes (I.) sevastianovi Khaustov, 2008 ; Iran ; p; Hajiqanbar & Arjomandi (2019)

Lasius flavus

Imparipes (I.) brevitarsus Ebermann, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus insolitus Khaustov, 2015 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus molnari Mahunka, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016); on lateral parts of ant abdomen

Scutacarus moseri Khaustov, 2015 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus ponticulus Mahunka, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus sibiriensis Khaustov, 2015 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016); attached to the ant head

Scutacarus tutus Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Lasius fuliginosus

Imparipes (I.) brevibasis (Sevastianov, 1983) ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Imparipes (I.) fuliginosophilus Khaustov, 2016 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Imparipes (I.) sevastianovi Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus flexisetus Karafiat, 1959 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus heterotrichus Khaustov, 2015 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Lasius niger

Lophodispus irregularis (Mahunka, 1971) ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus carsticus Mahunka & Mahunka-Papp, 1980 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Scutacarus crinitus Khaustov, 2015 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus ellipticus Karafiat, 1959 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus expectatus Karafiat, 1959 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus hauseri Mahunka, 1977 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus hystrichocentrus Sevastianov, 1983 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Scutacarus (gratus?) hortobagyensis Mahunka, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Scutacarus kassaii Mahunka, 1965 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus velutinosus Sevastianov, 1983 ¸ Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Lasius obscuratus *

Imparipes (I.) placidus Khaustov & Chydyrov, 2004 ; Iran ; p; Sobhi et al. (2017a)

Lasius umbratus

Scutacarus aequalis Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus flexisetosimilis Khaustov, 2016 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus levis Mahunka, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016); new record as phoretic species (so far reported from grass sample)

Scutacarus velutinosus Sevastianov, 1983 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

VII. SCUTACARID SPECIES ASSOCIATED WITH MYRMICINAE (HEXAPODA, ECTOGNATHA, ACULEATA, FORMICIDAE )

Aphaenogaster spp.

2 not nearer identified Imparipes spp. ; USA ; p; Uppstrom (2010)

4 not nearer identified Scutacarus spp. ; USA ; p; Uppstrom (2010) Messor sp.

Imparipes (I.) histricinus Berlese, 1903 ; Iran ; p; Sobhi et al. (2017a)

Scutacarus ebermanni Sobhi & Hajiqanbar, 2017 ; p; Iran ; Sobhi et al. (2017b)

Scutacarus remissus Khaustov, 2008 ; Iran ; p; Sobhi et al. (2017a)

Scutacarus shivicki Lazauskene & Sevastianov, 1974 ; Iran ; p; Sobhi et al. (2017a); new record as phoretic species (so far reported from soil and litter)

Messor caducus *

Imparipes (I.) gagati Khaustov, 2008 ; Iran ; p; Navabi et al. (2018a)

Myrmica ruginodis

Imparipes (I.) charkoviensis Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Imparipes (I.) comatosimilis Metwali, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Scutacarus ellipticus Karafiat, 1959 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus hystrichocentrus Sevastianov, 1983 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Scutacarus kassaii Mahunka, 1965 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus latus Karafiat, 1959 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus myrmecophilus Metwali, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus myrmicinus Khaustov, 2015 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus ovoideus Karafiat, 1959 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus pseudospinosus Khaustov, 2008 ¸ Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Scutacarus spinosus Štorkán, 1936 ¸ Russia ; nk (presumably ip); Khaustov & Tolstikov (2016); on lateral parts of ant abdomen

Scutacarus subterraneus (Oudemans, 1913) ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Pheidole sp.

Thaumatopelvis lorasiacus Navabi & Hajiqanbar, 2018 ; Iran ; p; Navabi et al. (2018b)

Solenopsis molesta *

2 not nearer identified Imparipes spp. ; USA ; p; Uppstrom (2010)

Tetramorium sp.

Imparipes (I.) imaginatus Mahunka, 1981 ; Iran ; p; Sobhi et al. (2017a)

Imparipes (I.) lentus Khaustov, 2008 ; Iran ; p; Sobhi et al. (2017a)

Imparipes (I.) malus Khaustov, 2008 ; Iran ; p; Navabi et al. (2018a)

Imparipes (I.) placidus Khaustov & Chydyrov, 2004 ; Iran ; p; Sobhi et al. (2017a)

Imparipes (I.) tenuis Mahunka, 1981 ; Iran ; p; Sobhi et al. (2017a)

Scutacarus shajariani Sobhi & Hajiqanbar, 2017 ; Iran ; p; Sobhi et al. (2017b)

Tetramorium caespitum

Imparipes sp. ; USA ; p; Uppstrom (2010)

Imparipes (I.) parapicosimilis Metwali, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Imparipes (I.) histricinus Berlese, 1903 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Imparipes (I.) imaginatus Mahunka, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Imparipes (I.) lentus Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Imparipes (I.) hortobagyensis Mahunka, 1981 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

Lophodispus irregularis (Mahunka, 1971) ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus claviger (Paoli, 1911) ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016) Scutacarus pseudospinosus Khaustov, 2008 ; Russia ; nk (presumably ip); Khaustov & Tolstikov (2016)

VIII. SCUTACARID SPECIES ASSOCIATED WITH MAMMALIA

Cricetidae

Pygmodispus odontotarsus Sevastianov, 1975 ; Ukraine ; i; Sosnina & Sevastianov (1975)