Poljanka hirsuta, Yang, Guang, Yao, Yunzhi & Ren, Dong, 2012

Yang, Guang, Yao, Yunzhi & Ren, Dong, 2012, A new species of Protopsyllidiidae (Hemiptera, Sternorrhyncha) from the Middle Jurassic of China, Zootaxa 3274, pp. 36-42 : 38-41

Poljanka hirsuta sp.n.

( Figs. 1–4 )

Diagnosis: Scapus 1.50 times thicker than pedicelli, pedicelli two times thicker than flagellomeres, apical flagellomere swollen; femora about two times thicker than corresponding tibiae, hind tibiae 1.46–1.57 times longer than fore and mid tibiae, first tarsomere of fore legs 1.33 times longer than second, first tarsomere of mid legs 1.50 times longer than second, first tarsomere of hind legs 2.31 times longer than second; R1 reduced, vein M+CuA about 1.57 times longer than vein R, branches of M about 2.22–2.39 times longer than M.

Description: Body 2.89 times as long as wide.

Head: Head 1.27 times as wide as long. Antennae 9.47 times length of head, its bases inserted in round antennal sockets, scapus enlarged, 1.50 times as thick as pedicelli, pedicelli cylindrical, twice as thick as flagellomeres, flagellomeres with dense long hairs, apical flagellomere swollen. Clypeus approximately triangular, junction between clypeus and rostrum broken. Rostrum stout, first segment longest, second 1.25 times as long as third ( Fig. 1 J).

Thorax: Pronotum 2.88 times as wide as long, flat to moderately deflexed; mesothorax barrel-shaped.

Legs: With approximate square trochanters; femora stout, nearly subequal in length, about two times as thick as corresponding tibiae; tibiae and tarsi with dense setae. Fore tibiae about 1.34 times as long as corresponding femora, first tarsomere of fore tarsi long, 1.33 times as long as second; mid legs similar to fore legs, tibiae slightly longer than fore tibiae, tarsi 0.50 times length of tibiae; hind legs distinctly longer than fore and mid legs, tibiae 2.10 times as long as femora, hind tibiae with three visible metatibial apical spurs ( Fig. 1 K), first tarsomere distinctly longer than fore and mid first tarsomeres, tarsi 0.45 times as long as tibiae, with two claws at apex of tarsi.

Wings: Forewings length 4.94 mm, distinctly extending beyond abdominal tip, width 2.26 mm, with marginal fringe of short setae. Most veins with setae of 0.06–0.23 mm length. R2 weekly sinuous. Rs branching off R very proximally and bent to costal margin of wing at apex abruptly. CuA connected to M creating the common part M+CuA and M+CuA connected to R forming the commom stem R+M+CuA. M+CuA stem about 1.57 times length of vein R. M straight, with two branches. M1+2 bent to wing fore margin, M3+4 bent to wing hind margin. M1+2 slightly longer than M3+4, both M1+2 and M3+4 2.22–2.39 times longer than M. Cell m1+2 3.25 times longer than cell cua1. CuA very long, about 1.89 times longer than M stem. CuA forked into CuA1 and CuA2. CuA1 long and curved, about 0.53 times length of CuA stem. CuA2 short, almost straight, 0.28 times length of CuA1. CuP nearly parallel to CuA. A1 and A2 merged into a point.

Abdomen: Abdomen roughly oval.

Measurements (in mm): Holotype, 3, CNU-PSY-NN2011008pc: Body length 5.00, maximal width of body 1.73; head length 0.45, width 0.57; antennal segments length I–X: 0.19, 0.21, 0.66, 0.60, 0.42, 0.51, 0.60, 0.18, 0.48, 0.42; length rostrum I–III: 0.66, 0.45, 0.36; length pronotum 0.33, width 0.95; length fore leg: femur 1.25, tibia 1.67, tarsomeres I–II: 0.48, 0.36; length mid leg: femur 1.22, tibia 1.79, tarsomeres I–II: 0.54, 0.36; length hind leg: femur 1.25, tibia 2.62, tarsomeres I–II: 0.83, 0.36; forewing length 4.94, width 2.26; R 0.42, R2 2.83, Rs 3.81, M+CuA 0.66, M 1.07, M1+2 2.56, M3+4 2.38, CuA 2.02, CuA1 1.07, CuA2 0.30; cell m1+2 2.50, cell cua1 0.77. Paratype, 3, CNU-PSY-NN2011018: Body length 6.31, maximal width of body 1.50; head length 0.25, width 0.56; antenna length 2.13; length fore leg: femur 1.19, tibia 2.08, tarsomeres I–II: 0.59, 0.40; length mid leg: tibia 2.28, tarsomeres I–II: 0.59, 0.40; length hind leg: femur 1.49, tibia 3.66, tarsomeres I–II: 0.84, 0.40; forewing length 3.81, width 1.88; R 0.31, R2 2.19, Rs 2.88, M+CuA 0.81, M 0.69, M1+2 1.94, M3+4 1.63, CuA 1.06, CuA1 0.81, CuA2 0.28; cell m1+2 1.88, cell cua1 0.63.

Material: Holotype, male, CNU-PSY-NN2011008pc (part and counterpart), a well-preserved specimen, with body mainly preserved and a rather clear forewing; paratype, male, CNU-PSY-NN2011018.

Etymology: The name of the new species refers to the setae on the antennae, legs and wings, which derives from Latin hirsuta (“hairy”).

Horizon and locality: Jiulongshan Formation, Middle Jurassic, Daohugou Village, Ningcheng County, Inner Mongolia, China.

Remarks: The new species is assigned to Poljanka due to the following features: apical flagellomere swollen; first tarsomere longer than second; R1 reduced; vein R slightly shorter than M+CuA; CuA with two branches. Poljanka differs, in addition, from Cicadellopsis Martynov, 1937 in the reduced R1 and from Carpenterella Bekker- Migdisova, 1968 and Karatavopsyllidium Bekker-Migdisova, 1968 in CuA bearing two branches.

Compared with P. shurabensis : forewing length 4.94 mm (vs. 6.50 mm); Rs bent at the apex abruptly (vs. Rs slightly bent at the apex); M1+2 bent (vs. M1+2 straight); branches of M short, about 2.22–2.39 times longer than M (vs. M branches longer, about 2.93–3.07 times as long as M).

Compared with P. ventriculosa : veins with short or long setae, (vs. without setae); first tarsomere of hind legs slightly longer than first tarsomere of mid legs (vs. the former seven times longer than the latter); forewing length 4.94 mm (vs. 4.00– 4.20 mm); branches of M 2.22–2.39 times length of vein M (vs. branches of M 3.40–3.50 times length of vein M); CuA1 curved (vs. CuA1 straight).

Compared with P. kukalovae : vertex without any setae (vs. with dense hair); hind tibiae 2.10 times as long as hind femora (vs. 1.27 times); forewing length 4.94 mm (vs. 3.00 mm); Rs bent at the apex abruptly (vs. Rs nearly straight).

Compared with P. sharovi : apical flagellomere slightly shorter than flagellomere seven (vs. apical flagellomere longer than flagellomere seven); hind tibiae 2.10 times as long as hind femora (vs. 1.57 times); forewing length 4.94 mm (vs. 2.80–3.20 mm); vein Rs bent at the apex abruptly (vs. Rs nearly straight); bifurcation of M before bifurcation of CuA (bifurcation of M after bifurcation of CuA); branches of M longer than vein M (vs. M3+4 slightly shorter than M); CuA1 curved (vs. CuA1 straight).

Compared with P. s p. 1: tarsi of hind legs distinctly longer than tarsi of mid legs (vs. tarsi of hind and mid legs subequal in length); forewing length 4.94 mm (vs. 3.10 mm); veins M1+2, M3+4 and CuA2 all bent more or less (vs. veins M1+2, M3+4 and CuA2 nearly straight).

Compared with P. s p. 2: rostrum stout (vs. rostrum relatively slim); femora nearly subequal in length (vs. hind femora longer than fore and mid femora); first tarsomere of fore legs slightly longer than second segment (vs. first about twice as long as second); forewing length 4.94 mm (vs. 4.00 mm); CuA1 3.60 times as long as CuA2 (vs. CuA1 2.50 times as long as CuA2).

Figures

FIGURE 1. A-F (after Bekker-Migdisova 1985): Forewing line drawings of Poljanka spp. A. P. shurabensis; B. P. ventriculosa; C. P. s p. 1; D. P. kukalovae; E. P. s h a ro v i; F. P. s p. 2. Scale bar = 2 mm. G-K: Poljanka hirsuta sp. n. Line drawings, CNU-PSY-NN 2011008. G. Right forewing; H. Left fore leg and antennae; I. Left middle leg; J. Rostrum; K. Right hind leg. Scale bar = 1 mm.

FIGURE 2. P. hirsuta sp. n. Photographs. A. Holotype, CNU-PSY-NN 2011008 p; B. Holotype, CNU-PSY-NN 2011008 c. Scale bar = 2 mm.

FIGURE 3. P. hirsuta sp. n. Line drawings of holotype, CNU-PSY-NN 2011008. A. Ventral view of body with wings; B. Dorsal view of body with wings. Scale bar = 2 mm.

FIGURE 4. P. hirsuta sp. n. Photographs, CNU-PSY-NN 2011008. A. Right forewing; B. Left fore leg and antennae; C. Rostrum; D. Right hind leg; E. Left middle leg. Scale bar = 1 mm.