Paraphloeostiba Steel, 1960

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

Paraphloeostiba Steel, 1960 a: 141 (species included: alutacea , specularis , rhopalocera ,

coriacea , opacicollis , similis , assamensis , impressicollis , secreta , sonani , sulcicollis , impressa , djawensis , papuana , obesa , solomonensis , gayndahense , hebridensis , cheesmani , seychellensis , subopaca , singularis , norfolcensis , fruhstorferi , sumatrensis , marianicola , apicalis , tonkinensis , celebensis ; key to species). TYPE SPECIES: Paraphloeostiba marianicola Steel , fixed by original designation .

— Shibata, 1973c: 27 (checklist of species of Taiwan).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Shibata, 1976: 106 (checklist of species of Japan).

— Zanetti, 1995: 7 (list of species of Italy).

— Assing, 1996a: 180, 181 (characters; notes).

— Assing, 1998f: 124 (characters).

— Newton, Thayer, Ashe, and Chandler, 2000: 337 (1 Nearctic species; notes; characters in key). alutacea Cameron, 1941b: 431 ( Phloeonomus ; Type locality: Philippines: Luzon, Imugan). — Steel, 1960a: 154 ( Paraphloeostiba ; characters; Philippines).

- DISTRIBUTION: Philippines.

angulata Motschulsky, 1861 , see: singulare Kraatz, 1859 .

apicalis Cameron, 1925 a: 175 ( Phloeonomus ; Type locality: Sumatra: Si Rambé ).

— Cameron, 1928d: 98 ( Phloeonomus ; Sumatra).

— Scheerpeltz, 1933: 1049 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 163 ( Paraphloeostiba ; characters; Sumatra; Java; Lombok; Philippines Islands). - DISTRIBUTION: Indonesia, Philippines.

assamensis Cameron, 1930: 145 ( Phloeonomus ; Type locality: Assam: Naga Hills ,

alt. 4000 feet).

— Scheerpeltz, 1933: 1049 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 149 ( Paraphloeostiba ; characters; Assam).

- DISTRIBUTION: India.

celebensis Steel, 1960 a: 169 ( Paraphloeostiba ; Type locality: Celebes: Kandari ).

- DISTRIBUTION: Indonesia.

cheesmani Cameron, 1934 a: 20 ( Phloeonomus ; Type locality: New Hebrides: Tanna ).

— Steel, 1960a: 157 ( Paraphloeostiba ; characters; New Hebrides).

- DISTRIBUTION: New Hebrides.

coriacea Cameron, 1952: 243 ( Phloeonomus ; Type locality: New Guinea: Snow Mountains: Sigi Camp, 1500 m).

— Steel, 1960a: 146 ( Paraphloeostiba ; characters; New Guinea).

- DISTRIBUTION: New Guinea.

densepunctatus Bernhauer, 1915 , see: obesa Fauvel, 1904 .

djawensis Steel, 1960 a: 154 ( Paraphloeostiba ; Type locality: Java: Tjibodas , 4000 feet).

- DISTRIBUTION: Indonesia.

fruhstorferi Steel, 1960 a: 163 ( Paraphloeostiba ; Type locality: Java: Tengger Mts. ,

2000 feet).

- DISTRIBUTION: Indonesia.

gayndahense W. J. MacLeay, 1873: 151 ( Omalium ; Type locality: Gayndah).

— Fauvel, 1877: 190 ( Omalium , cited as Homalium ; characters; Australia).

— Bernhauer and Schubert, 1910: 53 ( Omalium ; subgenus Omalium ; catalog).

— Bernhauer, 1926f: 312 ( Phloeonomus ; synonym of singularis ).

— Scheerpeltz, 1933: 1051 ( Phloeonomus ; synonym of singulare ).

— Steel, 1960a: 161 ( Paraphloeostiba ; valid species; characters; Queensland; New South Wales; Lord Howe Island ) .

— Kuschel, 1990: 25 ( Paraphloeostiba ; introduced, Australian species; winged; habitat; New Zealand).

— Zanetti, 1995: 7 ( Paraphloeostiba ; Italy).

— Assing, 1996a ( Paraphloeostiba ; characters; collecting notes; Italy; Sicily; Sardinia; Corsica; Canary Islands; Madeira; France; Germany).

— Klimaszewski, Newton, and Thayer, 1996: 146 ( Paraphloeostiba ; adventive in New Zealand). — Rogé, 1997 ( Paraphloeostiba ; characters; notes; France).

— Tronquet, 1998: 9 ( Paraphloeostiba ; characters; notes; France).

— Mineau, 1998: 272 ( Paraphloeostiba ; notes; France).

— Assing, Frisch, Kahlen, et al., 1998: 124 ( Paraphloeostiba ; characters).

— Assing and Wunderle, 1999: 435 ( Paraphloeostiba ; collecting notes; Canary Islands).

— Tronquet, 1999: 234 ( Paraphloeostiba ; notes; France).

— Letellier and Dauphin, 1999: 80 ( Paraphloeostiba ; notes; France).

— Newton, Thayer, Ashe, and Chandler, 2000: 337 ( Paraphloeostiba ; introduced; notes; California).

- DISTRIBUTION: Australia, New Zealand, Europe, Canary Islands, Madeira, USA. hebridensis Bernhauer, 1934 a: 18 ( Phloeonomus ; Type locality: Neue Hebriden: N.E. Malekula).

— Cameron, 1934a: 20 ( Phloeonomus ; New Hebrides).

— Steel, 1960a: 155 ( Paraphloeostiba ; characters; New Hebrides).

- DISTRIBUTION: New Hebrides.

impressicollis Cameron, 1924c: 312 ( Phloeonomus ; Type locality: Sumatra: Fort de Kock ).

— Cameron, 1928d: 98 ( Phloeonomus ; Sumatra).

— Scheerpeltz, 1933: 1050 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 149 ( Paraphloeostiba ; characters; Sumatra).

- DISTRIBUTION: Indonesia.

impressa Cameron, 1928d: 98 ( Phloeonomus ; Type locality: Sumatra: Fort de Kock ).

— Scheerpeltz, 1933: 1050 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 154 ( Paraphloeostiba ; characters; Sumatra).

- DISTRIBUTION: Indonesia.

kanalensis Steel, 1960: 4 ( Paraphloeostiba ; Type locality: New Caledonia: Kanala ).

- DISTRIBUTION: New Caledonia.

kyushuensis Watanabe, 1962 , see: Phloeonomus .

marianicola Steel, 1960 a: 165 ( Paraphloeostiba ; Type locality: Mariana Is.: Saipan, Sadog Talofofo).

- DISTRIBUTION: Mariana Islands.

norfolcensis Steel, 1960 a: 161 ( Paraphloeostiba ; Type locality: Norfolk Island).

- DISTRIBUTION: Norfolk Island.

obesa Fauvel, 1904b: 89 ( Omalium , cited as Homalium ; Type locality: Sumatra: region de Deli).

— Bernhauer and Schubert, 1910: 49 ( Phyllodrepa ; subgenus Hapalaraea ; catalog).

— Cameron, 1928d: 98 ( Phyllodrepa ; subgenus Hapalaraea ; Sumatra).

— Steel, 1960a: 152 ( Paraphloeostiba ; characters; Sumatra; Java; Sarawak; Philippine Islands). — Hammond, 1984: 195 ( Paraphloeostiba ; Borneo).

- DISTRIBUTION: Borneo, Malaysia, Indonesia, Philippines.

densepunctata Bernhauer, 1915f: 135 ( Phloeonomus ; Type locality: Sarawak: Mt. Matang , 1000 Fuss).

— Scheerpeltz, 1933: 1049 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 152 ( Paraphloeostiba ; synonym of obesa ).

opacicollis Steel, 1960 a: 150 ( Paraphloeostiba ; Type locality: Tonkin: Hoa Binh).

- DISTRIBUTION: Vietnam.

papuana Cameron, 1931: 356 ( Phloeonomus ; Type locality: Mount Lamington, New Guinea). — Scheerpeltz, 1933: 1050 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 153 ( Paraphloeostiba ; characters; New Guinea).

- DISTRIBUTION: New Guinea.

rhopalocera Cameron, 1928d: 99 ( Phloeonomus ; Type locality: Sumatra: Fort de Kock ).

— Scheerpeltz, 1933: 1050 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 146 ( Paraphloeostiba ; characters; Sumatra; New Britain).

secreta Cameron, 1928d: 98 ( Phloeonomus ; Type locality: Sumatra: Gunung Singgalang ).

— Scheerpeltz, 1933: 1050 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 152 ( Paraphloeostiba ; characters; Sumatra).

- DISTRIBUTION: Indonesia.

seychellensis Steel, 1960 a: 159 ( Paraphloeostiba ; Type locality: Seychelles: Silhouette , high forest over 1000 feet).

- DISTRIBUTION: Seychelles.

similis Steel, 1960 a: 150 ( Paraphloeostiba ; Type locality: Sumatra: Palembang ).

- DISTRIBUTION: Indonesia.

singularis Kraatz, 1859: 181 ( Omalium ; Type locality: Ceylan).

— Fauvel, 1889b: 246 ( Omalium , cited as Homalium ; New Caledonia).

— Bernhauer, 1902b: 45 ( Omalium ; Ceylon).

— Fauvel, 1903d: 258 ( Omalium , cited as Homalium ; New Caledonia [Note: According to Steel, 1960: 4 the New Caledonia specimens are an undescribed species of Paraphloeostiba ]; Celebes; Lombock; Java; Sumatra; Borneo; Ceylon).

— Fauvel, 1904a: 45 ( Omalium , cited as Homalium ; Nilghiris; Ceylan; Sikkim; Sumatra; Borneo; Java; Lombok; Celebes; New Caledonia).

— Bernhauer and Schubert, 1910: 59 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Bernhauer, 1920: 4 ( Phloeonomus ; Australia).

— Bernhauer, 1922b: 167 ( Phloeonomus ; Seychelles).

— Bernhauer, 1926f: 312 ( Phloeonomus ; Buru).

— Cameron, 1928d: 98 ( Phloeonomus ; Sumatra).

— Cameron, 1930: 145 ( Phloeonomus ; characters; Ceylon; India).

— Scheerpeltz, 1933: 1050 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960: 4 ( Omalium , cited as Homalium ; notes).

— Steel, 1960a: 159 ( Paraphloeostiba ; characters; all of distributional records except Ceylon and India are based on misidentifications).

— Frank, 1982: 9 ( Phloeonomus ; list of parasites; Sumatra).

- DISTRIBUTION: Sri Lanka, India.

angulata Motschulsky, 1861: 142 ( Phloeonomus ; Type locality: Colombo).

— Bernhauer and Schubert, 1910: 51 ( Omalium ; subgenus Omalium ; catalog).

— Cameron, 1930: 146 ( Phloeonomus ; characters; Ceylon).

— Scheerpeltz, 1933: 1049 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Cameron, 1934d: 78 ( Phloeonomus ; notes on types one of which is singulare ).

— Steel, 1960a: 159 ( Paraphloeostiba ; synonym of singulare ).

— Scheerpeltz, 1966e: 393 ( Phloeonomus ; cited as valid species; [Note: Although cited by Scheerpeltz as a valid species, I have followed the work of Steel since Scheerpeltz may have overlooked that article]; New Caledonia).

solomonensis Steel, 1960 a: 155 ( Paraphloeostiba ; Type locality: Solomon Islands: Guadalcanal, Kukum ).

- DISTRIBUTION: Solomon Islands.

sonani Bernhauer, 1943 a: 171 ( Phloeonomus ; Type locality: Formosa: Arisan ).

— Steel, 1960a: 149 ( Paraphloeostiba ; characters; Formosa).

— Frank, 1982: 9 ( Paraphloeostiba ; list of parasites).

- DISTRIBUTION: Taiwan.

specularis Bernhauer, 1915 g: 181 ( Phloeonomus ; Type locality: Not cited).

— Scheerpeltz, 1933: 1051 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 146 ( Paraphloeostiba ; characters; New Britain).

- DISTRIBUTION: New Britain.

subopaca Cameron, 1924b: 168 ( Phloeonomus ; Type locality: Simla Hills; Gahan, 7000 feet).

— Cameron, 1930: 144 ( Phloeonomus ; characters; India).

— Scheerpeltz, 1933: 1051 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

sulcicollis Fauvel, 1904b: 89 ( Omalium , cited as Homalium ; Type locality: Sumatra: Palembang).

— Bernhauer and Schubert, 1910: 59 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Cameron, 1928d: 98 ( Phloeonomus ; Sumatra).

— Steel, 1960a: 148 ( Paraphloeostiba ; characters; Sumatra).

- DISTRIBUTION: Indonesia.

sumatrensis Bernhauer, 1905: 11 ( Phloeonomus ; Type locality: Sumatra: Balighe ).

— Bernhauer and Schubert, 1910: 59 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Cameron, 1928d: 98 ( Phloeonomus ; Sumatra).

— Steel, 1960a: 163 ( Paraphloeostiba ; characters; Sumatra; Lombok).

- DISTRIBUTION: Indonesia.

tonkinensis Cameron, 1940b: 181 ( Phloeonomus ; Type locality: Tonkin: Hoa Binh).

— Steel, 1960a: 167 ( Paraphloeostiba ; characters; Tonkin; Singapore).

- DISTRIBUTION: Vietnam, Singapore.

Phloeonomus

[57 species; Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan ,

Oriental, Australian, and Oceanic Regions]

Phloeonomus Heer, 1839: 184 (species included: monilicornis , inflatus , lucidus , pusillus , minimus ). TYPE SPECIES: Omalium pusillus Gravenhorst , fixed by subsequent designation by Thomson, 1859: 51.

— Lacordaire, 1854: 143 (synonym of Omalium ).

— Jacquelin du Val, 1857: 76 (synonym of Omalium ).

— Kraatz, 1857d: 974 (synonym of Omalium ).

— Thomson, 1859: 51 (characters; type species: pusillus ).

— Thomson, 1861: 210 (characters).

— Fauvel, 1875a: IV [= 1875b: 206] (catalog; group of Omalium , cited as Homalium ).

— Mulsant and Rey, 1880: 179 (characters; key to French species).

— Seidlitz, 1891: 339 (key to some European species).

— Seidlitz, 1891a: 361 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 120 (subgenus of Omalium ; list of species of Europe and Caucasus).

— Ganglbauer, 1895: 731 (characters; key to middle European species).

— Luze, 1906: 589 (characters; key to species).

— Heyden, Reitter, and Weise, 1906: 131 (list of species of Europe and Caucasus).

— Sainte-Claire Deville, 1907: 36 (key to and annotated list of species of Seine Basin, France).

— Bernhauer and Schubert, 1910: 57 (world catalog; 28 species).

— Petri, 1912: 53 (list of species of Siebenbürgen).

— Johansen, 1914: 606 (characters; key to Danish species).

— Leng, 1920: 94 (catalog of North American species).

— Cameron, 1921a: 350, 399 (characters; key to Singapore species; catalog).

— Cameron, 1924b: 169 (key to Indian species).

— Winkler, 1925: 331 (catalog for Palaearctic region).

— Cameron, 1925: 11, 104 (subgenus of Omalium ; catalog of Indian species).

— Porta, 1926: 22 (characters; key to species of Italy).

— Portevin, 1929: 445 (characters; key to French species).

— Cameron, 1930: 141 (characters; key to species of British India).

— Roubal, 1930: 310 (catalog; Slovakia).

— Joy, 1932: 100 (key to British species).

— Scheerpeltz, 1933: 1048 (world catalog supplement).

— Blackwelder, 1943: 51 (characters; type species: pusillus ).

— Blackwelder, 1944: 102 (checklist of species from Mexico, Central and South America , and the West Indies).

— Kloet and Hincks, 1945: 161 (list of British species).

— Tottenham, 1949: 355, 406 (type species: pusillus Gravenhorst ; list of British species). — Blackwelder, 1952: 305 (type species: pusillus ). — Jarrige, 1950a: 93 (cited as Phlaeonomus; key to French species). — Blackwelder, 1952: 305 (type species). — Tottenham, 1954: 25 (characters; key to British species). — Hatch, 1957: 52, 81 (characters; key to species of Pacific Northwest). — Smetana, 1959a: 196 (list of species collected in Albania). — Székessy, 1963: 15 (characters). — Lohse, 1964: 46 (characters; key to central European species). — Smetana, 1964c: 169 (key to European species). — Pototskaia, 1967: 20 (cited as Pholoeonomus; larval characters). — Scheerpeltz, 1968a: 11 (catalog of Austrian species). — Steel, 1970a: 15 (larval characters; biological notes). — Tikhomirova, 1973a: 138 (subgenus of Phloeonomus ; checklist of species of USSR). — Moore and Legner, 1975: 209 (catalog; North America north of Mexico). — Shibata, 1976: 104 (checklist of species of Japan). — Pototskaia, 1976: 162 (larval characters). — Nikitskii, 1976: 183 (larval characters). — Pope, 1977: 23 (list of British species). — Burakowski, Mroczkowski, and Stefańska, 1979: 54 (catalog; Poland). — Moore and Legner, 1979: 209 (characters; notes). — Muona, 1979: 18 (list of Scandinavian species). — Uhlig, Vogel, and Sieber, 1980: 240 (some species collected in Germany). — Uhlig and Vogler, 1981: 86 (list of some species collected in Germany; habitat and collecting notes). — Tóth, 1982: 65 (characters; habitus; key to species of Hungary). — Muona and Viramo, 1986: 15 (list of species of northeastern Finland). — Segers, 1986: 20 (checklist of species; Belgium). — Zanetti, 1987: 247 (characters; habitus; key to species of Italy). — Watanabe, 1990: 99 (characters; key to species of Japan). — Angelini, 1991: 192 (list of species from high elevations of Calabria, Italy). — Campbell and Davies, 1991: 88 (checklist of species; Alaska and Canada). — Silfverberg, 1992: 21 (list of species of Fennoscandia, Denmark, and the Baltic States). — Welch, 1993 (ovariole structure and development). — Boháč, 1993: 41 (list of species of Czech Republic and Slovakia). — Zanetti, 1995: 7 (list of species of Italy). — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species). — M. Hansen, 1996: 94 (list of Danish species). — Assing, 1996a: 180, 181 (characters; notes). — Downie and Arnett, 1996: 438 (characters; species of Northeastern North America ). — Assing, 1998f: 123, 124 (characters; notes). — Newton, Thayer, Ashe, and Chandler, 2000: 337 (2 Nearctic species; notes; characters in key).

Phloeonomodes Smetana, 1964: 163 , 168 (subgenus of Phloeonomus ; species included: minimus , quadrifossulatus ). TYPE SPECIES: Omalium minimus Erichson , fixed by original designation. — Horion, 1963: 92 (subgenus of Phloeonomus ; list of central European species). — Tikhomirova, 1973a: 138 (cited as Phloenomodes; subgenus of Phloeonomus ; checklist of species of USSR). — Zanetti, 1987: 248, 252 (subgenus of Phloeonomus ; characters; habitus; species of Italy).

abietinus Thomson, 1867 , see: pusillus Gravenhorst, 1806 .

affinis Cameron, 1933b: 339 ( Phloeonomus ; Type locality: British North Borneo: Kenokok, 3300 feet). — Hammond, 1984: 195 ( Phloeonomus ; Borneo).