Payraudeautia ermesi, Pedriali & Sosso & Dell’Angelo, 2019

Pedriali, Luca, Sosso, Maurizio & Dell’Angelo, Bruno, 2019, Naticid gastropods from the middle Miocene of western Ukraine, Zootaxa 4700 (2), pp. 151-195 : 177-180

Payraudeautia ermesi sp. nov.

(Fig. 29.1−3)

Type material. Holotype and 9 paratypes ( Table 14).

Type locality. Varovtsi , Ukraine .

Type stage. Middle Miocene (Upper Badenian) .

Etymology. The species is named for Ermes Pedriali, father of one of the present authors (LP).

Other material. Varovtsi: NP 10030 (8 shells); BD 168 (9 shells); MS 107 (10 shells). Horodok: NP 10038 (2 shells). Zalistsi: NP 10037 (3 shells); BD 169 (6 shells).

Diagnosis. Globose-oval shell with rather elevated spire and inflated last whorl. Umbilicus deep, narrow, with broad, depressed funicle and slightly prominent abapical spiral ridge. Parietal callus large, with small adapical knob and semielliptical anterior lobe; umbilical callus thick, semicircular, demarcated from anterior lobe of parietal callus by slight transverse groove. Colour pattern of uneven whitish-brown collabral stripes, darker subsuturally and on lower base, over uniformly pale brown background; apex, parietal and umbilical calluses whitish.

Description. Protoconch small, turbiniform, of 2.20−2.25 convex, smooth whorls, tip small. Teleoconch glo- bose-oval, solid. Spire conical, rather elevated, whorls convex; suture thin, adpressed. Last whorl inflated, slightly expanded toward aperture. Subsutural shelf indistinct, periphery above midline. Aperture D-shaped in slightly prosocline plane. Parietal callus large, with concave abapertural outline, moderately thick, with small adapical knob and semielliptical anterior lobe, not reaching basal fasciole. Umbilicus deep, narrow; umbilical border rounded; umbilical wall concave, incised by weak, shallow groove and bearing scarcely prominent, narrowly round-topped spiral ridge bent toward interior of umbilicus and terminating in subtriangular, asymmetric plug on lowermost part of inner lip. Funicle broad, rather depressed, separated from abapical spiral ridge by broad, shallow groove. Umbilical callus thick, semicircular, demarcated from anterior lobe of parietal callus by slight transverse groove. Basal fasciole poorly defined. Surface with very fine growth lines. One well preserved specimen retains uniform pale brown background with pattern of uneven whitish-brown collabral stripes, darker subsuturally and on lower base; apex, parietal and umbilical calluses whitish.

Remarks. Payraudeautia ermesi sp. nov. is closely similar to P. fasciolata (Figs 29.4, 29.5), a species from late Miocene to late Pliocene deposits of Northern and Central Italy, but differs from it in that it has: 1) a protoconch with smaller diameter (averages 0.746 mm and 1.037 mm, respectively); 2) a more elevated spire (see Fig. 11 ), with more convex whorls; 3) an adapical knob on parietal callus; 4) a greater aperture width (see Figs 14, 15 ); 5) a broader umbilical callus (see Fig. 20 ); and 6) a smaller umbilicus with a thinner inner spiral ridge, separated from the umbilical wall by a narrower and shallower groove.

The late Miocene to Recent Mediterranean species Payraudeautia intricata (Fig. 29.6−8) is also similar, but can be easily differentiated from P. ermesi sp. nov. by having: 1) a sculptured protoconch of 0.5 fewer whorls, with greater diameter of the initial half-whorl (average values 0.320 mm and 0.103 mm, respectively); 2) a less elevated spire with less convex whorls; 3) a parietal callus devoid of knob; 4) a smaller aperture width (see Figs 14 and 15 ); 5) a larger umbilicus; and 6) a more excavated umbilical wall bearing a more prominent spiral ridge. It is also worth noting that P. intricata attains a larger size.

Payraudeautia ermesi sp. nov. resembles the Tortonian northwestern Italian species P. bituberculata (Figs 30.7, 30.8), but is readily distinguished from it because of its protoconch of 0.7 fewer whorls, with significantly smaller diameter (averages 0.746 mm and 1.082 mm, respectively) and greater diameter of the first half-whorl, its higher spire (see Fig. 11 ), with more convex whorls, its parietal callus with an anterior lobe instead of a knob,and its broad- er umbilical callus (see Fig. 20 ), of different shape (semicircular and not subtriangular as in P. bituberculata ).

Payraudeautia pigeonblancensis Van Dingenen, Ceulemans & Landau, 2016 , described from the early Pliocene (Zanclean) of North West France (protoconch measures given by the authors: DHW 0.085 mm, PD 1.000 mm), is easily differentiated from Payraudeautia ermesi sp. nov. by its protoconch with greater diameter, its incised suture, its larger umbilicus, and its more excavated umbilical wall bearing a more prominent spiral ridge. Payraudeautia sabrinae sp. nov. and Payraudeautia varovtsiana sp. nov. (see below), can be separated from P. pigeonblancensis on the basis of the same teleoconch characters; moreover, these latter species exibit a protoconch with fewer whorls (respectively of 1.5 to 1.6 for P. sabrinae sp. nov. and 1.65 for P. varovtsiana sp. nov. instead of 2.60 as in P. pigeonblancensis ), and with smaller diameter (average 0.742 mm, 0.706 mm and 1.000 mm, respectively) and greater diameter (average 0.306 mm, 0.191 mm and 0.085 mm, respectively) of the first half-whorl.

Distribution. Middle Miocene: Central Paratethys (upper Badenian) in Ukraine (this paper).

Figures

FIGURES 4–11. FIGURE 4. Relationship between the diameter of the first half-whorl of the protoconch and the protoconch diameter in the naticine taxa studied. FIGURE 5. Relationship between the diameter of the first half-whorl of the protoconch and the protoconch diameter in the poliniceine and sinine taxa studied. FIGURE 6. Relationship between the diameter of the first half-whorl of the protoconch and the protoconch whorls in the naticine taxa studied. FIGURE 7. Relationship between the diameter of the first half-whorl of the protoconch and the protoconch whorls in the poliniceine and sinine taxa studied. FIG- URE 8. Relationship between the protoconch diameter/diameter of the first half-whorl ratio and the protoconch whorls in the naticine taxa studied. FIGURE 9. Relationship between the protoconch diameter/diameter of the first half-whorl ratio and the protoconch whorls in the poliniceine and sinine taxa studied. FIGURE 10. Relationship between the spire height and the shell height for species of Euspira Agassiz in J. Sowerby, 1837 studied herein. FIGURE 11. Relationship between the spire height and the shell height for species of Payraudeautia Bucquoy, Dautzenberg & Dolfuss, 1883 studied herein.

FIGURES 12–19. FIGURE 12. Relationship between the spire height and the shell height for species of Polinices Monfort, 1810 studied herein. FIGURE 13. Relationship between the aperture width and the aperture height for species of Cochlis Röding, 1798 studied herein. FIGURE 14. Relationship between the aperture width and the aperture height for species of Payraudeautia Bucquoy, Dautzenberg & Dolfuss, 1883 studied herein. FIGURE 15. Relationship between the aperture width and the maximum diameter for species of Payraudeautia Bucquoy, Dautzenberg & Dolfuss, 1883 studied herein. FIGURE 16. Relationship between the umbilicus width and the maximum diameter for species of Euspira Agassiz in J. Sowerby, 1837 studied herein. FIGURE 17. Relationship between the umbilicus width and the maximum diameter for species of Payraudeautia Bucquoy, Dautzenberg & Dolfuss, 1883 studied herein. FIGURE 18. Relationship between the width of the umbilical callus and the maximum diameter for species of Cochlis Röding, 1798 studied herein. FIGURE 19. Relationship between the width of the umbilical callus and the maximum diameter for species of Tectonatica Sacco, 1890 studied herein.

FIGURES 20–22. FIGURE 20. Relationship between the width of the umbilical callus and the maximum diameter for species of Payraudeautia Bucquoy, Dautzenberg & Dolfuss, 1883 studied herein. FIGURE 21. Relationship between the width of the umbilical callus and the umbilicus width for species of Cochlis Röding, 1798 studied herein. FIGURE 22. Relationship between the width of the abapical sulcus and the width of the adapical sulcus for species of Cochlis Röding, 1798 studied herein.