Patellapis (Patellapis) rubricata, Timmermann & Kuhlmann, 2009

Timmermann, Kim & Kuhlmann, Michael, 2009, Variable Selection and Inference for Multi-period Forecasting Problems, Zootaxa 2099, pp. 1-188 : 20-24

Patellapis (Patellapis) rubricata sp. n. Timmermann

( Figs. 8a–g ; 9a–h )

Diagnosis. The females differ from those of P. depressa and P. gessorum in lacking a median longitudinal clypeal depression. Further, the inner hind tibial spurs of P. rubricata are more coarsely serrate than in P. depressa and P. gessorum . The males of P. rubricata can be easily distinguished from those of P. depressa by the size and structure of the gonostylus and flagellomeres A3 and A4. The gonostylus of P. rubricata has a large, dorso-lateral projection (simple in P. depressa ) and A4 is as long as A3 (A4 about twice as long as A 3 in P. depressa ). Both sexes of P. rubricata differ markedly from P. bifurcata in their larger body length. The males of P. rubricata are closely related to those of P. pseudorubricata . The males of P. pseudorubricata differ from those of P. rubricata in having a larger body length, a broader apical plate and a scopa-like vestiture on the hind tibia. (long, dense, and branched hairs). Further, the penis valves of P. pseudorubricata are simple in size, whereas those of P. rubricata are dorsally enlarged with a hyaline membrane.

Description

Female. Bl = 7,2–7,7mm. General habitus ( Fig. 8e ). Head. L = 1,8–1,9mm; W = 2,2–2,3mm. Head wider than long ( Fig. 8a ). Integument black except mandibles sometimes partly reddish-brown. Face shiny and with loose, long, greyish, erect hairs. Mandibles bidentate. Malar area very short, linear. Clypeus and supraclypeal area convex in profile (sometimes supraclypeal area weakly convex). Clypeus strongly and sparsely punctate (i = 1,5–2,0d). Clypeoantennal distance 0,4mm. Supraclypeal area sparsely punctate (i = 1,5–2,0d); surface partly sculptured. Paraocular area strongly and densely punctate; surface partly sculptured. Antennae mostly brownish. Mesosoma. L = 1,4–1,5mm; W (ITS) = 1,7–1,8mm. Integument black. Scutum polished; surface partly reticulate anteriorly; disc strongly and sparsely punctate (i = 1,5–2,0d) ( Fig. 8b ). Scutellum as illustrated in Fig. 8c . Scutum, scutellum and metanotum with long, greyish to brownish, erect hairs; mesepisternum and propodeum with long, greyish, erect hairs. Basal area of propodeum as illustrated in Fig. 8c ; propodeum shiny and densely punctate; surface finely sculptured. Marginal region of posterior truncation shiny; finely reticulate. Wings. Hyaline. Legs. Integument mostly brownish. Vestiture greyish to brownish. Ts moderately finely serrate with about 10 oblique projections ( Fig. 8f ). Bp broad and pointed at apex ( Fig. 8g ).

Metasoma. L = 3,6–4,2mm; W = 2,5–2,6mm. Integument mostly reddish-brown except the margins of T pallid reddish-brown to pallid yellowish (some specimens have the metasomal terga completely reddish to pale reddish); margins extensively translucent. Apical hair bands on T1–T4 hardly existent (at most weakly developed). Prepygidial fimbria yellowish to brownish ( Fig. 8d ).

Male. Bl = 7,2–7,6mm. General habitus ( Fig. 9e ). Head. L = 1,8–1,9mm; W = 2,2–2,4mm. Head wider than long. Integument black except mandibles sometimes partly reddish-brown. Clypeus, supraclypeal area, paraocular area and vertex covered with erect, greyish to yellowish hairs; the lower half of paraocular area, supraclypeal area and clypeus also covered with appressed, whitish hairs. Mandibles simple. Malar area very short, linear. Antennae brownish-black. Flagellomeres of normal structure, not knotty. A4 about as long as A3.

Mesosoma. L = 1,5–1,6mm; W (ITS) = 1,7–1,8mm. Integument black. Scutum polished; strongly and sparsely punctate (i = 1,5–2,0d) ( Fig. 9a ). Scutellum as illustrated in Fig. 9b . Scutum, scutellum,metanotum, mesepisternum and propodeum with long, greyish to yellowish, erect hairs. Basal area of propodeum as illustrated in Fig. 9b ; propodeum shiny, strongly and densely punctate; surface finely sculptured. Marginal region of posterior truncation shiny and reticulate. Wings. Hyaline. Legs. Integument brownish to black. Vestiture greyish to yellowish. Metasoma. L = 4,0– 4,5mm; W = 2,4–2,5mm. Integument brownish-black except the margins of T pallid brownish; margins extensively translucent ( Fig. 9c ). Apical plate broad and pointed apically. Posterior margin of S4 with a row of strong, erect bristles; medial bristles nearly appressed (superficially it looks like a medial interruption of the row of bristles); metasomal sternum S5 with small, erect hair patches laterally ( Fig. 9d ). S7 and S8 as illustrated in Fig. 9f . Genitalia. Genitalia as illustrated in Fig. 9g –h ; gonocoxa dull and gonostylus with dorso-lateral projection.

Type material ( 422 specimens). Holotype, male, South Africa, Northern Cape, Nieuwoudtville, road to farm Glen Lyon , slope and side of the road, S31°23'25" / E19°08'28", 740m, leg. K. Timmermann, 18.viii.2007, SANC.

Paratypes: 221♀, 200♂. South Africa. Western Cape : 1♀, Montagu , 09.x.1966, leg. M.H. and C.D. Michener, SEMC ; 7♀, Worcester , 30.ix.1966, leg. C.D. Michener, SEMC ; 5♀, idem., 08.x.1966, SEMC ; 3♀, 10♂, Citrusdal , 08.ix.1966, leg. C.D. Michener, SEMC ; 1♀, Roggeveld Escarpment , 07.ix.1995, leg. P. Goldblatt and J. Manning, SEMC ; 2♀, 1♂, near Redelinghuys , ix.2001, leg. P. Goldblatt, SEMC ; 1♀, 4♂, Clanwilliam , 08.ix.1966, leg. M. and C.D. Michener, SEMC ; 5♀, 11km W Clanwilliam , 03.x.1990, leg. C.D. Eradley, HYMA07639 , SANC ; 1♀, Graafwater , 14.ix.1987, leg. C.D. Eardley, Database No. HYMA 07280 , SANC ; 1♀, Ratelfontein / Olaf Bergfontein , 08.x.1995, leg. F.W., S.K. and R . W. Gess, No. 95/96/291, AMGS ; 1♀, near Die Berg , 20.ix.2005, leg. C.D. Eardley, SANC ; 1♀, Vanrhynsdorp near Water Fall , 15.ix.2005, leg. C.D. Eardley, SANC ; Northern Cape: 1♂, Leliefontein , pan trap, 05.ix.2002, leg. C. Mayer, KTPC ; 1♀, idem., 02.x.2002, KTPC ; 1♂, idem., 31.viii.2003, KTPC ; 7♂, idem., 05.ix.2003, KTPC ; 5♂, idem., 07.ix.2003, KTPC ; 18♀, 18♂, idem., 11.ix.2003, KTPC ; 3♂, idem., 15.ix.2003, KTPC ; 1♂, idem., 17.ix.2003, KTPC ; 3♀, 2♂, idem., 23.ix.2003, KTPC ; 2♂, idem., 04.x.2003, KTPC ; 2♂, idem., 06.x.2003 ,

KTPC; 18♀, 11♂, Witwater , pan trap, 17.ix.2003 , leg. C. Mayer, KTPC; 31♀, 5♂, idem., 23.ix.2003 , KTPC; 2♀, 2♂, idem., 25.ix.2003 , KTPC; 3♀, idem., 01.x.2003 , KTPC; 3♀, idem., 04.x.2003 , KTPC; 7♀, Remhoogte , pan trap, 21.ix.2002 , leg. C. Mayer, KTPC; 1♀, idem., 02.x.2002 , KTPC; 3♂, idem., 31.viii.2003 , KTPC; 8♂, idem., 05.ix.2003 , KTPC; 2♀, 24♂, idem., 07.ix.2003 , KTPC; 3♀, 17♂, idem., 11.ix.2003 , KTPC; 1♂, idem., 13.ix.2003 , KTPC; 6♂, idem., 15.ix.2003 , KTPC; 1♂, idem., 17.ix.2003 , KTPC; 2♀, 1♂, idem., 23.ix.2003 , KTPC; 2♂, idem., 25.ix.2003 , 1♂, idem., 01.x.2003 , KTPC; 1♀, 15 km S Nieuwoudtville , pan trap, 09.x.2002 , leg. F.D. Parker and M.E. Irwin, BLCU; 7♂, Nieuwoudtville , pad to Farm Glenlyon (road to R27 ), slope, 18.viii.2007 , leg. K. Timmermann, KTPC; 1♂, Nieuwoudtville , pad to Farm Glenlyon (road to R27 ), 19.viii.2007 , leg. K. Timmermann, KTPC; 12♀, 33♂, idem., 03.ix.2007 , KTPC; 32♀, 11♂, idem., 07.ix.2007 , KTPC; 35♀, 5♂, idem., 17.ix.2007 , leg. K. Timmermann and M. Kuhlmann, KTPC; 1♀, Nieuwoudtville , Caravan Park, 26.–30.ix.1990 , leg. F.W. and S.K. Gess, No. 90/91/ 56, AMGS; 1♀, Farm Papkuilsfontein , 12.–19.x.2000 , leg. S. van Noort and H.G. Robertson, SANC; 4♀, Farm Dassiefontein near Kamieskroon, 01.x.1990 , leg. C.D. Eardley, SANC; 9♀, 2♂, Groenekloof , 7,5 km SE of Leliefontein, 13.ix.1992 , leg. F.W. and S.K. Gess, No. 92/93/83, AMGS; 2♀, 1♂, Springbok , Goegap Nature Reserve (hills), 8.–10.ix.1992 , leg. F.W. and S.K. Gess, No. 92/93/30, No. 92/93/28, AMGS; 2♂, Steinkopf , leg. L. Schultze, Am. Mus. Nat. Hist. Dept. Invert. Zool. No. 24828 , AMNH and ZMHB; 1♂, Leliefontein , pan trap, 05.ix.2002 , leg. C. Mayer, KTPC; 1♀, idem., 02.x.2002 , KTPC; 1♂, idem., 31.viii.2003 , KTPC; 7♂, idem., 05.ix.2003 , KTPC; 5♂, idem., 07.ix.2003 , KTPC; 18♀, 18♂, idem., 11.ix.2003 , KTPC; 3♂, idem., 15.ix.2003 , KTPC; 1♂, idem., 17.ix.2003 , KTPC; 3♀, 2♂, idem., 23.ix.2003 , KTPC; 2♂, idem., 04.x.2003 , KTPC; 2♂, idem., 06.x.2003 , KTPC; 18♀, 11♂, Witwater , pan trap, 17.ix.2003 , leg. C. Mayer, KTPC; 31♀, 5♂, idem., 23.ix.2003 , KTPC; 2♀, 2♂, idem., 25.ix.2003 , KTPC; 3♀, idem., 01.x.2003 , KTPC; 3♀, idem., 04.x.2003 , KTPC; 7♀, Remhoogte , pan trap, 21.ix.2002 , leg. C. Mayer, KTPC; 1♀, idem., 02.x.2002 , KTPC; 3♂, idem., 31.viii.2003 , KTPC; 8♂, idem., 05.ix.2003 , KTPC; 2♀, 24♂, idem., 07.ix.2003 , KTPC; 3♀, 17♂, idem., 11.ix.2003 , KTPC; 1♂, idem., 13.ix.2003 , KTPC; 6♂, idem., 15.ix.2003 , KTPC; 1♂, idem., 17.ix.2003 , KTPC; 2♀, 1♂, idem., 23.ix.2003 , KTPC; 2♂, idem., 25.ix.2003 , 1♂, idem., 01.x.2003 , KTPC.

Additional material (30 specimens). 10♀, 20♂. South Africa. Western Cape : 6♀, 13♂, Worcester, ix.1928, leg. R . E. Turner, BMNH ; 1♀, 1♂, idem., 03.–04.x.1928, BMNH ; 3♀, 4♂, idem., ix.–x.1931, BMNH ; 2♂, Capetown, x.1937, leg. R . E. Turner, BMNH .

Etymology. Named after the reddish metasomal terga.

Distribution ( Fig. 10 ). Widely distributed in the winter rainfall area.

Floral visitation. Moraea fugax , Romulea subfistulosa (Iridaceae) , Cheiridopsis cigarettifera , Cheiridopsis denticulata , Herrea spec. , Ruschia spec. , Ruschia unca (Mesembryanthemaceae) .

Seasonal activity (first–last observations). viii–x.

Remarks. This species is treated as Patellapis new species 8 in Timmermann and Kuhlmann (2008b).

Figures

FIGURE 8. Patellapis (P.) rubricata, female: a. Head (scale=1mm); b. Scutum (scale=1mm); c. Scutellum and propodeum (scale=0,5mm); d. Metasomal terga (scale=1mm); e. Body (scale=1mm); f. Inner hind tibial spur (scale=100µm); g. Basitibial plate (scale=100µm).

FIGURE 9. Patellapis (P.) rubricata, male: a. Scutum (scale=1mm); b. Scutellum and propodeum (scale=0,5mm); c. Metasomal terga (scale=1mm); d. Metasomal sterna (scale=1mm); e. Body (scale=1mm); f. Metasomal sterna S7 and S8, ventral view (scale=0,5mm); g. Genitalia, dorsal view (scale=0,5mm); h. Genitalia, lateral view (scale=0,5mm).

FIGURE 10. Distributions of P. gessorum, P. pseudorubricata and P.rubricata.