Parisognoriste eocenica, Blagoderov, Vladimir, Hippa, Heikki & Nel, Andre, 2010

Blagoderov, Vladimir, Hippa, Heikki & Nel, Andre, 2010, Parisognoriste, a new genus of Lygistorrhinidae (Diptera: Sciaroidea) from the Oise amber with redescription of Palaeognoriste Meunier, ZooKeys 50, pp. 79-90 : 81-83

Parisognoriste eocenica sp. n.

Material examined.

Holotype: complete inclusion of female in transparent fossil resin, Oise amber, deposited in the Muséum national d’Histoire naturelle, Paris: MNHN A32914 (PA876).

Type locality: France: Oise department: region of Creil, Chevrière, farm Le Quesnoy, 49°19.533'N, 2°40.833'E. Geological horizon: the lowermost Eocene, in amber, c. -53 My, Sparnacian, level MP7 of the mammal fauna of Dormaal ( Nel et al. 2004).

Paratypes: two inclusions of females in the same piece of amber (during preparation the piece was cut into two), deposited in the Muséum national d’Histoire naturelle, Paris: MNHN A32914 (PA876).

Morphology.

Female. Measurements, mm: Length total 2.22-2.48 (holotype 2.48); wing 1.83-1.91 (holotype 1.91); antenna 0.44-0.49 (holotype 0.49); palpi 0.22.

Head (Fig. 1) globular with short evenly distributed setae on occiput, face non-setose. Eye bare, rounded, not emarginated at antennae base, facets round and equal in size. Ocelli almost in a straight transverse line, the middle slightly smaller than the laterals, distance between lateral ocellus and middle ocellus 2.0x the distance between lateral ocellus and eye margin. Four palpomeres visible: ultimate one elongated, its length 3x the width; penultimate palpomere subglobular, slightly longer than wide; antepenultimate palpomere (the sensilliferous palpomere) about 2x longer than wide, much wider than the more apical ones; the palpomere basally from the penultimate one short, shorter than the penultimate one. Other mouthparts very short, ~ 1/3 of head height, labrum dark, triangular; labellum and hypopharynx light, narrow, sharply pointed. Antenna with 2+14 segments, apical segment secondarily divided into two. Scape and pedicel 2x wider than flagellomeres, globular. Flagellomeres subcylindrical, ca. 1.5x broader than long.

Thorax. Scutum uniformly setose with lateral and posterior setae being longer. Scutellum with 10 marginal setae, which are as long and strong as the setae on the posterior part of scutum. Anteprontum with five setae, proepisternum with three setae. Pleural pit inconspicuous. Anepimerone pointed ventrally, touching katepisternum in single point. Laterotergite not produced strongly laterally, bare. Mediotergite non-setose, not convex posteriorly. Metepisternum trapezoidal, with height almost equal to width.

Legs. Procoxa longer than the others. Metacoxa with a row of long posterolateral setae. Protibia about the length of profemur. Tibial organ with a small lobe, but without any distinct setation. Metatibia not conspicuously expanded apically but steadily widening from base to apex, its apical width ca. 2x the basal width. The vestiture on the apical third of tibia in rows. Spur formula 1:2:2, mesotibia with inner spur 2x the outer, metatibia with inner spur 2.5x the outer. Claws 1-3 pointed.

Wing (Fig. 2). Costa extending beyond R5 tip at 3/5 distance between tips of R5 and M1. Sc joining C. Rs distinct, oblique. M stem and base of M1+2 fork inconspicuous. M3+4 reduced at base. Transverse vein connecting Rs and CuA with a distinct kink at the base of M3+4. R1 with 18-20 dorsal setae, Sc, R5, M, and CuA bare.

Abdomen. Cercus (Fig. 3) two-segmented, basal segment rounded, very short, apical segment oval, its length 2x the width. Sternite 8 wide, rounded apically.

Etymology.

The species name is an adjective in the nominative singular, derived from the Eocene period, referring to the temporal distribution of the species.

Discussion.

See under Parisognoriste above.