Paraphloeostiba pilosa, Shavrin, 2024

Shavrin, Alexey V., 2024, Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960, Zootaxa 5424 (3), pp. 251-307 : 287-289

Paraphloeostiba pilosa sp. n.

( Figs 77 , 104–110 )

Type material. Holotype ♂ ( Fig. 77 ; left antennomeres 3–11 glued on the same card under the specimen): ‘ IRIAN JAYA: Paniai Prov. | Sinak, trail to Ilaga | 2100-2500m, 15. 12. 1995 | leg. A. Riedel’ <printed>, ‘ HOLOTYPE | Paraphloeostiba | pilosa sp. n. | Shavrin A. V. des. 2024’ <red, printed> ( NMW).

Paratypes: 1 ♂ (dissected): same data as the holotype ( NMW); 4 ♀♀ (one specimen without left antennomeres 3–11; one specimen dissected): ‘ NEW GUINEA | Okasa, | 7.II.1966 ’ <printed>, ‘ R. Hornabrook | BM 1970-232.’ <printed> ( BMNH). All paratypes with additional red printed label: ‘ PARATYPE | Paraphloeostiba | pilosa sp. n. | Shavrin A. V. des. 2024’.

Description. Measurements (n=6): HW: 0.42–0.49; HL: 0.25–0.27; OL: 0.12; TL: 0.02–0.03; AL (holotype): 0.63; PL: 0.35–0.40; PWmax: 0.57–0.60; PWmin: 0.53–0.56; ESL: 0.66–0.74; EW: 0.73–0.88; MTbL (holotype): 0.40; MTrL (holotype): 0.24 (MTrL 1–4: 0.08; MTrL 5: 0.16); AW: 0.73–0.90; AedL: 0.47; BL: 1.95–2.65 (holotype).

Habitus as in Fig. 77 . Body dark-brown, with indistinctly paler elytra; antennomeres 6–11 or 7–11 brown; mouthparts and antennomeres 1–5 or 1–6 yellowish; legs reddish, with slightly paler tarsi. Head, neck, pronotum and scutellum without punctation; elytra with very dense, moderartely large and deep punctation, denser around scutellum and sparser along suture; abdominal tergites with indistinct, sparse and fine punctation, more visible on tergites III–V. Head with very dense, coarse, small microsculpture; pronotum with very dense isodiametric microsculpture, slightly coarser than that in middle portion of head; scutellum with dense and coarse transverse meshes; microsculpture of elytra moderartely fine, longitudinal or isodiametric, indistinct or absent in middle portions of each elytron, denser and caorser in mediobasal and medioapical portions.

Head slightly elevated in middle, with distinct, shallow or moderately deep oval anteriomedian depressions, 1.6–1.8 times as broad as long, with narrow and short anteocellar foveae, diagonally stretching anteriad; postocular carina distinct, acute. Ocelli large, located slightly below level of postocular carina; distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Apical maxillary palpomere 1.3–1.5 times as long as preapical segment. Antennomeres 7–10 slightly transverse; antennomere 3 disitnctly longer and narrower than 2, 5 slightly broader than 4, 6 slightly narrower than 5, 7 slightly longer and distinctly broader than 6, 8 distinctly longer and broader than 7, 9–10 slightly longer and broader than 8.

Pronotum distinctly convex, 1.5–1.6 times as broad as long, 1.2–1.3 times as broad as head, from widest middle distinctly more narrowed posteriad than anteriad.

Elytra 1.1 times as broad as long; surface of each elytron with longitudinal elevations between punctures and diagonal elevations in apical portion.

Metatarsi slightly less than twice as long as metatibia.

Abdomen about as broad as elytra.

Male. Posterior margin of abdominal tergite VIII truncate ( Fig. 106 ). Posterior margin of abdominal sternite VIII widely concave ( Fig. 107 ). Aedeagus with very wide basal portion, gradually narrowed toward rounded apex of median lobe; parameres significantly exceeding apex of median lobe, from widest preapical portions slightly narrowed toward rounded apices, with numerous very long apical and preapical setae; internal sac moderately narrow and long, with two elongate and sclerotized structures in basal part ( Fig. 104 ). Lateral aspect of the aedeagus as in Fig. 105 .

Female. Posterior margin of abdominal tergite VIII truncate ( Fig. 108 ). Posterior margin of abdominal sternite VIII rounded ( Fig. 109 ). Accessory sclerite short, from widest basal portion gradually narrowed toward widely rounded apex ( Fig. 110 ).

Comparative notes. Based on the general shape of the aedeagus, with the parameres distinctly longer than apex of the median lobe with very long apical and preapical setation, P. pilosa sp. n. is similar to P. sulcicollis (Fauvel, 1904) from Sumatra and Hong Kong (see Shavrin 2020). It can be distinguished from P. sulcicollis by the presence of the punctation of the head, more convex pronotum without distinct longitudinal impressions, the broader aedeagus with narrower apical portions of the median lobe and parameres, and the different shape of the female accessory sclerite.

Distribution. Paraphloeostiba pilosa sp. n. is known from two localities in Western New Guinea ( Indonesia) and central part of Papua New Guinea.

Etymology. The specific epithet is the Latin pilosus, - a, - um (hairy). It alludes to the presence of very long setation of the parameres.

Bionomics. Specimens were collected at elevations from 2100 to 2200 m a.s.l. The detailed bionomical data are unknown.

Figures

FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm.

FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm.