Paleocader rovnensis, HEISS & GOLUB, 2021

HEISS, ERNST & GOLUB, VIKTOR B., 2021, Three new species of the fossil Cantacaderinae genus Paleocader Froeschner, 1996 from the Baltic and Rovno amber (Hemiptera, Heteroptera, Tingidae), Palaeoentomology 4 (4), pp. 360-366 : 364-365

Paleocader rovnensis sp. nov.

( Fig. 3 )

Holotype. Male in an uncut piece of Rovno amber 57 × 37 × 20 mm. The specimen is complete, head with antennal segments I+II partly obscured on dorsal and ventral side by impurities. This specimen JDC 9234 (20/02) is designated as holotype and deposited at CEHI as ROVTING-01.

Etymology. The name refers to Rovno amber where this interesting fossil tingid was discovered.

Diagnosis. P. rovnensis sp. nov. is recognized and differs by following characters from other congeners described herein: from P. balticus sp. nov. and P. pulchellus sp. nov. by smaller size (3.70 vs. 3.85 and 3.90 mm); number of transverse veinlets in subcostal area (2 vs. 3 and 3); shorter antennae, ratio antennal segments I–IV / width of head (3.09 vs. 4.90 and 3.54).

Description. Macropterous male.

Head. About as long as wide, antennae about 3.1 times as long as width of head, segments I+II short and thicker, III longest and thinner, about 2.3 times as long as width of head, IV short thin and fusiform; approximate length of antennal segments I / II / III / IV = 0.1 / 0.1 / 1.25 / 0.25 mm; eyes globular semi-inserted in head; rostral groove laterally laminate along head, pro-, meso- and metasternum, rostrum short, reaching middle of sternite 2.0.

Pronotum. Wider than long, lateral margins straightly converging anteriorly from rounded humeral angles to narrowly rounded anterolateral angles; paranota with three rows of areolae at its widest part, one or two elsewhere; surface of disk areolate with five rows of carinae, the two humeral ones shorter than the three median ones extending along full length of pronotum; posterior lobe posteromedially raised, higher than anterior lobe; posterior margin convex without prolongation.

Scutellum. Triangular, small with acute apex.

Hemelytra. Elongate oval, distinctly produced over abdomen, 1.62 times as long as width, lateral margins evenly rounded; clavus with rows of areolae, commissura slightly longer than pronotum; sutural area with one row of areolae along clavus and few additional ones at insinuation at apex of clavus; discoidal area with six rows of areolae at its widest and two transverse veinlets; subcostal area with six rows and two (left side) and three veinlets (right side); costal area with four rows of areolae which are larger than those of all other parts of hemelytra; stenocostal area with one row; membrane completely areolate, their areoles increasing in size posteriorly.

Legs. Femora and tibiae straight, long and slender, tarsi two-segmented with long claws.

Measurements. Length 3.7 mm, head length / width 0.55 / 0.55 mm; pronotum length / width 0.8 / 1.1 mm; hemelytra length / width 2.75 / 1.7 mm, ratio 1.62; length of antennal segment III / width of head 1.25 / 0.55 mm, ratio 2.3.

Key to species of Paleocader . (Measurements of previously described species were taken from their descriptions or illustrations).

1 Pronotum subrectangular, antennae shorter, about 2.8 times as long as width of head, antennal segment III about twice as long as width of head, Baltic amber.... P. avitus (Drake)

- Pronotum converging anteriorly, antennae longer, at least 3 times as long as width of head, antennal segment III at least 2.2 times as long as width of head ................................... 2

2 Lateral margins of pronotum posterolaterally angular at humeri, antennae 5.6 times as long as width of head, Baltic amber ........................................ P. gusenleitnerorum Heiss

- Lateral margins of pronotum posterolaterally rounded, antennae shorter................................................................ 3

3 Costal area of hemelytra with six rows of areolae ........... 4

- Costal area of hemelytra with four rows of areolae ......... 5

4 Larger species (4.5 mm), abdomen wider, ratio length to with of hemelytra 1.49, antennal segment III about 3.2 times as long as width of head, Baltic amber ..................... .................................................................. P. serafini Heiss

- Smaller species (3.9 mm), abdomen narrower, ratio length to with of hemelytra 1.68, antennal segment III about 3.8 times as long as width of head, Baltic amber ........................................ P. strictus Golub & Popov

5 Antennae longer, about 4.9 times as long as width of head, antennal segment III 4 times as long as width of head, Baltic amber ................................................ P. pulchellus sp. nov.

- Antennae shorter, at most 3.5 times as long as width of head, antennal segment III at most 2.7 times as long as width of head................................................................................... 6

6 Larger species (3.9 mm) abdomen wider, ratio length to with of hemelytra 1.53, antennae longer, about 3.5 times as long as width of head, Baltic amber .... P. balticus sp. nov.

- Smaller species (3.7 mm), abdomen slender, ratio length to with of hemelytra 1.62, antennae shorter, 3.1 times as long as width of head, Rovno amber ......... P. rovnensis sp. nov.

Figures

FIGURE 3. Paleocader rovnensis sp. nov., holotype male. A, Dorsal view. B, Ventral view. C, Anterior body, ventral. D, Amber piece with inclusion, arrow indicates the position of specimen. E, Terminal abdomen, ventral. Scale bars = 1 mm in A and B; 500 μm in C and E. ©J. Damzen