Pachamama speciosa Owen & Pinto

Owen, Albert K. & Pinto, John D., 2004, Pachamama, an uncommon and distinctive new genus of Trichogrammatidae (Hymenoptera: Chalcidoidea) from tropical America, Zootaxa 664, pp. 1-8 : 3-7

Pachamama speciosa Owen & Pinto , new species

Type. Holotype ɗ: ECUADOR. Napo: Tiputini Biodiversity Sta.; nr. Yasuni National Park; 220–250m; 00°37’55”S, 76°08’39”W; Lot # 1940; fogging terre firme forest; 26.x.1998; T. Erwin et al. Deposited in USNM.

Etymology. speciosus: Latin for beautiful, handsome, imposing.

Description. Body length 0.6mm (N=1; measurement of single card­mounted female); color golden­brown, with distinctly darker mesonotum; legs colored similarly to body, tending to be darker towards base; head light brown, with or without a near­white band immediately below eye; antennal scape and pedicel transversely banded with offwhite and varying shades of brown; club uniformly brown; forewing fumate except apical tenth. Antenna with BPS elongate ( Fig. 7 ), ca. 5 x as long as wide, slightly bulbous apically. Maxillary palp ( Fig. 9 ) with a well­developed apical seta and a smaller medial seta; apical peg­like sensillum absent. Forewing ca. 1.8 x as long as wide; setation sparse, restricted to apical.35 of wing; only R, RS2 and r­m setal tracks discernible; setation posterior to r­m not clearly arranged in tracks; fringe elongate, with longest setae on anteroapical margin of wing, ca. 0.3 greatest wing width; venation elongate, extending ca. 0.7 x wing length; MV with 2­3 elongate setae arising from anterodorsal margin; costal cell lacking setae. Hind wing ca. 5 x as long as wide; disk with two complete setal tracks closely spaced in the anterior 1/3 and a 3rd complete setal track in the posterior ¼, setae between 2nd and 3rd tracks scattered, not linearly arranged; fringe stout and elongate, with maximum length along posterior edge, and extending 0.9 hind wing width.

Male (N = 3, except for antennal characteristics N=2)

Antenna with scape extremely enlarged, suboval, ca. 1.5 x as long as wide, subequal to length of remainder of antenna; club conical and symmetrical; relative length of scape: pedicel: club = 3:1:2; C3 2– 3 x as long as C1 and C2 combined. C1 with 1 APB and 0–1 BPS; C2 with 1 APB, 0–2 APA, and 2 BPS; C3 with 3 APB in basal half, 1–2 APA, 3 BPS, 4–5 FS, and 3 PLS parallel to longitudinal axis of segment. Forewing with relative length of PM: MV: SV = 4:5:1. Genital capsule widest at middle, sides tapering gradually to apex, and more abruptly and unevenly to base; genital capsule and aedeagus including apodemes subequal, together ca. 0.6–0.7 x hind tibial length; apodemes ca. 0.2 x genitalia length and restricted to basal 1/3 of capsule.

Female (N= 3)

Antennal scape unmodified, elongate, ca. 3 x as long as wide; club apically attenuate and slightly asymmetrical, tapering evenly on dorsal surface and more abruptly so on ventral surface; with more sensilla than male; relative length of scape: pedicel: club = 2:1:3; C3 7­ 8 x as long as C1 + C2 combined. C1 with 1 APB and 0­1 BPS; C2 with 1 APB, 0­ 2 APA, and 2 BPS; C3 with 2­3 APB near base and 2­4 short APB near apex, 0­1 APA, 3­ 4 BPS, 10­13 FS, 1 elongate and setiform UPP at apex, 5 PLS including 1 PLS transverse at base of segment, but curving longitudinally in apical half, and 1 PLS on dorsal surface extending beyond apex. Forewing with relative length of PM: MV: SV = 2:2:1. Ovipositor ca. 0.6 x length of hind tibia.

Geographic distribution. Costa Rica, Ecuador, and Honduras.

Material examined. 4Ψ (1 on card), 3ɗ. COSTA RICA. Puntarenas: R. F. Golfo Dulce; 24km. west of Piedras Blancas; 200m; ii.1992; 1Ψ; P. Hanson [ UCRC]. ECUA­ DOR. Napo: Reserva Etnica Waorani; 1km S. Onkone Gare Camp; 220m; 00°39’10”S, 76°26’00”W; 3.vii.1995; fogging terre firme forest; Lot # 1091; 1Ψ; T. Erwin et al.

[ NMNH]; As previous except collected 21.vi.1994 and Lot# 697; 1Ψ (card mounted) [ UCRC]. Napo: Tiputini Biodiversity Sta. 1ɗ (See type) [ NMNH]; 1ɗ, same data as holotype (missing antennae) [ NMNH]; 1ɗ, same data as holotype except collected 5.vii.1998 and Lot # 1889 [ UCRC]. HONDURAS. Olancho: La Muralla National Park; ~ 1480m; 15°05’49”N, 86°44’17”W; 4–7.vii.2002; 1 Ψ; D. Yanega [ UCRC].

Remarks. Specimens are deposited as indicated above in brackets. The specimen from La Muralla National Park, Honduras, is a primary molecular voucher (D# 776; Gen­ Bank accession number AY623495 for 28S D2 & D3; Owen et al. in prep.). It is represented on a slide by an individual forewing, hind wing, and antenna.

Figures

FIGURES 7 – 9. Pachamama speciosa. 7, antenna (medial). 8, mandible showing two strong posterior teeth, and socketed anterior tooth (at arrow). 9, maxillary palp.