Osmylopsychostoechus sogulensis, Khramov, Alexander V. & Makarkin, Vladimir N., 2015

Khramov, Alexander V. & Makarkin, Vladimir N., 2015, New fossil Osmylopsychopidae (Neuroptera) from the Early / Middle Jurassic of Kyrgyzstan, Central Asia, Zootaxa 4059 (1), pp. 115-132 : 123-126

Osmylopsychostoechus sogulensis sp. nov.

Figs 6–8

Description. Holotype PIN 2032/510 ( Fig. 6 ). Forewing ca. 23 mm long as preserved (estimated complete length ca. 24–25 mm), ca. 11 mm wide. Costal space broad, dilated towards wing base. All preserved subcostal veinlets once or twice forked. Sc, R fused distally. Veinlets of Sc+R1 numerous (≥ 10); humeral veinlet recurrent, branched. Subcostal space moderately broad, with two detected crossveins. Space between R1, Rs narrowed towards apex; crossveins scarce (four detected). Rs with 23 almost parallel branches, not forked before outer gradate series of crossveins (except one in apical portion of wing), rather regularly forked distad outer gradate series. MA deeply forked some distance from its origin. Distal nygma well visible, located in MA space. MP forked distad fork of MA; MP space markedly dilated proximally. MP1 strongly concave, dichotomously branched at outer gradate series. MP2 pectinate, with four long branches, one of these deeply forked. Basal crossvein 1m-cu (‘M5’) rather long, oblique. Cu dividing into CuA, CuP near wing base. CuA somewhat incurved, strongly convex before branching, pectinate, with five long branches, three of these deeply forked. CuP long, deeply dichotomously branched (with three long branches). Basally between Cu/CuP, A1 well-developed concave fold. A1 strongly convex proximally, dichotomously branched; basal-most fork of A1 proximad basal-most fork of CuP. Space between A1, A2 basally broad. A2 pectinate, with more than four branches (incompletely preserved); basal-most branch deeply forked. A3 long, pectinately branched, with three preserved branches; proximal-most branch deeply forked. Crossveins relatively numerous (compared with other specimens), arranged irregularly except for incomplete inner gradate series and complete outer gradate series. Trichosors prominent along proximal part of hind margin, indistinct along other preserved margins. Color pattern not preserved.

Paratype PIN 1526/116 ( Fig. 7 ). Forewing ca. 23 mm long as preserved (estimated complete length ca. 25–26 mm); 10.5 mm wide. Costal space broad, dilated towards wing base. All preserved subcostal veinlets forked, once to dichotomously. Sc, R fused distally. Sc+R1 with 10 veinlets, mainly dichotomously branched. Costal crossveins not detected. Subcostal space moderately broad, definite crossveins not detected but two or three possible (poorly discernable). Space between R1, Rs narrow; crossveins scarce (three or four possible but poorly discernable). Rs with 22 almost parallel branches. Rs2, Rs5 deeply forked (before inner gradate series), three other branches forked between inner, outer gradate series; other branches of Rs forked distad outer gradate series. Proximal part of MA (including its fork) not preserved; both branches not forked before outer gradate series. Distal nygma not detected, but probably present in dilation of MA space before inner gradate series. MP forked distad primary fork of CuP. MP1 strongly concave, dichotomously branched just after outer gradate series. MP2 few pectinate, with three long branches, one of these deeply forked. Intra-MP space slightly dilated proximally. Basal crossvein 1m-cu (‘M5’) rather long, oblique. Cu dividing into CuA, CuP near wing base. CuA somewhat incurved, strongly convex before branching, pectinate, with four long branches, two of these deeply forked. CuP long, deeply dichotomously branched (with two long branches). A1 probably dichotomously branched. A2 pectinate, probably with five branches. A3 pectinate. Proximal parts of anal veins not preserved. Crossveins scarce, mainly arranged in incomplete inner gradate series and almost complete outer gradate series. Trichosors clearly visible along costal margin, poorly preserved along other preserved margins. Color pattern not preserved.

Paratype PIN 2384/472 ( Fig. 8 ). Forewing ca. 25 mm long, 11 mm wide. Costal space broad, dilated towards wing base. Subcostal veinlets poorly preserved, probably forked. Sc, R1 fused distally. Sc+R1 with 12 forked veinlets. Subcostal space moderately broad in proximal portion, crossveins not detected. Space between R1, Rs moderately broad in proximal portion, narrowed apically; two crossveins in distal portion. Rs with 23 almost parallel branches (proximal portions incompletely preserved in some branches); three-four branches forked between inner, outer gradate series; all branches forked (mainly dichotomously) distad outer gradate series. MA deeply forked, its branches few dichotomously branched after outer gradate series. Distal nygma not preserved, but probably present in dilation of intra-MA space. MP forked slightly distad fork of MA. MP1 concave, pectinately branched, with four very oblique branches, proximal-most of these deeply forked. MP2 slightly sinuate, strongly convex, few pectinate, with three long very oblique branches, one of these deeply forked. Intra-MP space slightly dilated proximally. 1m-cu (‘M5’) rather long, oblique. Cu dividing into CuA, CuP near wing base. CuA somewhat incurved, strongly convex proximad branching, pectinate, with four long branches, two of these deeply forked. CuP deeply dichotomously branched. A1 strongly convex proximad branching, deeply dichotomously branched. A2 pectinate, with six branches. A3 not preserved. Crossveins very poorly visible; preserved crossveins mainly belong to outer gradate series. Trichosors clearly visible along almost entire costal margin, poorly presented along other preserved margins. Color pattern not preserved.

Hind wing fragmentarily preserved. MP forked far from wing base. CuA strongly concave, slightly incurved.

Type material. Holotype PIN 2032/510 (part, counterpart), a nearly complete well-preserved forewing. Paratype PIN 1526/116 (part, counterpart), a nearly complete well-preserved forewing. Paratype PIN 2384/472 (part only), a nearly complete well-preserved fore- and hind wing overlapping. All housed in PIN.

Type locality and horizon. Kyrgyzstan: Sai-Sagul; late Early Jurassic to the early Middle Jurassic.

Etymology. From the Sogul Formation.

Remarks. The venation of these three forewings is generally similar to each other differing, however, in some details. For example, the crossvenation varies rather greatly, but this might be due to differences in preservation.

Figures

FIGURE 6. Osmylopsychostoechus sogulensis gen. et sp. nov. Holotype PIN 2032 / 510. A, part. B, counterpart. C, forewing venation. Scale bars = 5 mm (B, C to scale).

FIGURE 7. Osmylopsychostoechus sogulensis gen. et sp. nov. Paratype PIN 1526 / 116. A, part. B, counterpart. C, forewing venation. Scale bar = 5 mm (all to scale).

FIGURE 8. Osmylopsychostoechus sogulensis gen. et sp. nov. Paratype PIN 2384 / 472, specimen as preserved. Scale bar = 5 mm.