Ormosia (Oreophila) confluenta Alexander, 1922

Kato, Daichi, Watanabe, Kozo & Kolcsar, Levente-Peter, 2022, Japanese species of Ormosia Rondani (Diptera, Limoniidae): revision of the subgenera Oreophila Lackschewitz and Parormosia Alexander, ZooKeys 1132, pp. 127-162 : 127

publication ID	https://dx.doi.org/10.3897/zookeys.1132.86022
publication LSID	lsid:zoobank.org:pub:46E68453-A2FA-4D22-A0D7-4509DFFB7C1B
persistent identifier	https://treatment.plazi.org/id/12777ED8-864B-5DC6-BD1A-68DC255C9E78
treatment provided by	ZooKeys by Pensoft
scientific name	Ormosia (Oreophila) confluenta Alexander, 1922
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Ormosia (Oreophila) confluenta Alexander, 1922 View in CoL

Figs 1A View Figure 1 , 2 View Figure 2 , 3 View Figure 3 , 4 View Figure 4 , 5A View Figure 5

Ormosia confluenta in Alexander 1922: 183: original description (type locality: Japan, Honshu, Ôsaka, Mt. Minomo).

Ormosia (Ormosia) confluenta in Alexander 1953a: 71: faunistic record.

Ormosia (Oreophila) confluent in Nakamura 2002: 169: faunistic record; Nakamura 2014: 31: distribution; Oosterbroek 2022: distribution.

Type material examined.

Paratype. Japan • ♀; Honshu, Mt. Minomo; 4 May 1921; K. Takeuchi leg.; USNM.

Non-type material examined.

Japan • 2 ♂; Honshu, Nagano, Ueda-shi, Sanada-machi-Osa, Kakuma Valley; 36.45378°N, 138.36592°E; alt. 1050 m; 16 May 2012; D. Kato leg.; BLKU. • 2 ♂; Honshu, Tokyo, near Tokyo, Mt. Mitake; 10 May 1931; B. Oda leg.; USNM. • 1 ♂; Honshu, Okayama, Maniwa-shi, Hiruzen-Shimotokuyama; 35.32931°N, 133.59725°E; alt. 780 m; 1 May 2016; D. Kato leg.; BLKU. • 1 ♀; Honshu, Hiroshima, Akiôta-chô, Yokogô; 34.59419°N, 132.14497°E; alt. 890 m; 18 May 2015; D. Kato leg.; BLKU. • 1 ♂; Shikoku, Tokushima, Mt. Tsurugi; alt. 1400-1950 m; 31 May 1950; Issiki and Ito leg.; USNM. • 1 ♂; Shikoku, Tokushima, Miyoshi-shi, Higashiiya-Ochiai, near Matsuogawa Dam; 33.96478°N, 133.93908°E; alt. 900 m; 15 May 2015; • 1 ♂, 1 ♀; same locality; 30 Apr. 2016; D. Kato leg., BLKU. • 6 ♂, 1 ♀; Shikoku, Tokushima, Miyoshi-shi, Higashiiya-Sugeoi, near Nagoro Dam; 33.85182°N, 134.0234°E; alt. 920 m; 29 Apr. 2016; • 3 ♂; same locality; 30 Apr. 2016; D. Kato leg. BLKU. • 4 ♂, 1 ♀; Shikoku, Ehime, Kumakogen, River Myogadani, springs; 33.56701°N, 132.9344 °E; 1420 m; 8 May 2022; L.-P. Kolcsár leg.; CKLP. • 1 ♂; Kyushu, Saga, Karatsu-shi, Kyuragi-Hirano, Mt. Sakurei-zan; 33.35701°N, 130.07038°E; alt. 860 m; 26 Apr. 2015; D. Kato leg.; BLKU.

Diagnosis.

General coloration yellow to pale brown (Fig. 1A View Figure 1 ). Vertex often pale brownish grey. Antenna dark brown except scape. Wing yellowish tinged, unpatterned. Legs distal to mid-tibiae gradually becoming dark brown towards tips. Male terminalia: tergite 9 bearing pair of triangular lobes at caudal margin. Gonocoxite slightly produced beyond base of clasper of gonostylus. Clasper of gonostylus wider apically, 3/4 length of lobe of gonostylus, distal part 2 × as wide as that of lobe of gonostylus. Interbases fused medially into roundish sac-like plate, without mesal-apical lobe. Female terminalia with cercus almost straight, slightly upcurved on distal part. Genital frame with lateral arm of genital fork roughly triangular, situated at posterior 1/3 of genital fork, with finger-shaped lobe on posterior end. Sternite 9 very small, fan-shaped.

Redescription.

Male. Body length 2.9-4.2 mm, wing length 4.1-5.9 mm.

Head: covered with yellow to brown setae. Vertex dusky yellow to pale brownish grey, paler grey on anterior part, often widely dark brown on center of posterior part. Eyes small and widely separated, ~ 1/2 as wide as narrowest point of vertex, ~ 1/3 length of head including rostrum in dorsal view. Rostrum dusky yellow to pale brown, ~ 1/2 length of eye in lateral view. Palpus 5-segmented, 2/3 length of head, dusky yellow on basal two segments, dark brown on succeeding segments, palpomere 1 small, globular, palpomere 2 cylindrical, palpomeres 3-5 globular. Labellum dark brown. Antenna 15 to 16-segmented, relatively short, ~ 2 × length of head; scape dusky yellow to pale brown, 2 × as long as wide; pedicel dark brown, oval, 2/3 length of scape; flagellomeres dark brown, barrel-shaped, gradually decreasing in size toward apical segment; each flagellomere with ca. six verticils, longest one except in apical segment ~ 1.5 × as long as each segment.

Thorax: covered with yellow setae. Antepronotum dusky yellow to pale brown; postpronotum pale yellow. Mesonotum subnitidous, pale brown to brown, lateral margin yellow, postero-outer corner of scutal lobe often yellow. Scutellum or mediotergite sometimes yellowish. Prescutal pit indistinctly present, oval to bacilliform. Tuberculate pit indistinctly present, situated slightly anterior to level of prescutal pit. Pleuron dusky yellow to yellow. Wing (Fig. 2 View Figure 2 ) yellowish tinged, narrow, 3.5-4 × as long as wide; stigma absent; Sc ending between level of forks of Rs and R2+3+4; crossvein sc-r indistinct or absent, situated on level of basal 1/3 of Rs if present; R2+3+4 1/7-1/4 length of R3; R2 situated between fork of R2+3+4 and length of itself distal to it; M4 1-1.5 × as long as M3+4; wing margin between tips of CuP and A1 1.3-2 × as long as that between tips of CuP and CuA; A1 almost straight. Halter yellow, ~ 2/3 length of thorax. Legs yellow on coxae to femora; tibiae yellow basally, gradually turning to dark brown toward tips; tarsi dark brown (Fig. 1A View Figure 1 ).

Abdomen: yellow to yellowish ochreous, densely covered with yellow setae.

Male terminalia (Fig. 3 View Figure 3 ): Tergite 9 with pair of triangular lobes at caudal margin, ~ 1/3 length of middle of tergite 9; anterior margin of tergite 9 deeply and widely notched; tergite 9 slightly wider than long including caudal lobe (Fig. 3A View Figure 3 ). Sternite 9 almost straight at caudal margin (Fig. 3B View Figure 3 ). Gonocoxite almost same width in whole length, ca. as long as tergite 9, posteroventral margin slightly produced beyond base of clasper of gonostylus (Fig. 3A View Figure 3 ). Gonocoxal apodeme short, connected to anterolateral part of interbase (Fig. 3D View Figure 3 ). Clasper of gonostylus scabrous, darkened apically, 3/4 length of lobe of gonostylus, gradually wide distally, rounded at tip, distal part 2 × as wide as that of lobe of gonostylus in apical view (Fig. 3C View Figure 3 ). Lobe of gonostylus long finger-shaped, ~ 1/2 length of gonocoxite, slightly curved, distal part flattened in apical view (Fig. 3C View Figure 3 ). Interbases fused medially into roundish sac-like plate, ca. as long as wide in dorsal view, posterior margin slightly concave, anterolateral part with short arm (Fig. 3D View Figure 3 ). Paramere roughly blade-shaped, ca. as long as interbase, anterior end curved inward (Fig. 3D View Figure 3 ). Aedeagus dorso-ventrally flattened, ~ 1/3 width of interbase, tip slightly beyond apex of interbase (Fig. 3D View Figure 3 ). Sperm pump long oval, anterior end reaching at anterior 1/3 of paramere (Fig. 3D, E View Figure 3 ). Ejaculatory apodeme poorly developed (Fig. 3D, E View Figure 3 ).

Female. Body length 5.0-6.2 mm, wing length 5.6-6.7 mm. Generally resembling male.

Female terminalia (Fig. 4 View Figure 4 ): yellow to yellowish ochreous, cercus and hypogynial valve amber-colored. Tergites 8 and 9 fused. Cercus almost straight, slightly upcurved on distal part, 1.5 × as long as tergite 10; hypogynial valve ~ 1.7 × as long as sternite 8, basal part distinctly wider than that of cercus, gradually narrowed on distal 1/2 toward tip, tip acute, ending near level of middle of cercus (Fig. 4A View Figure 4 ). Genital frame with genital fork, constricted at anterior 1/3, roughly heart-shaped on anterior end; lateral arm of genital fork roughly triangular distally, situated at anterior 2/3 of genital fork, with small finger-shaped lobe at posterior end; sternite 9 very small, narrower than posterior end of genital fork, fan-shaped distally, largely desclerotized on middle of posterior part. Three spermathecal ducts present, middle one very wide; spermathecae indistinct (Fig. 4B View Figure 4 ).

Distribution.

Japan (Honshu, Shikoku, and Kyushu) (Fig. 5A View Figure 5 ), Russia (Far East), and Kuril Is.

Remarks.

This species resembles a Nearctic species, Ormosia (Oreophila) flaveola (Coquillett, 1900) ( Coquillett 1900), and an Eastern Palaearctic species known from North Korea and China, Ormosia (Oreophila) yankovskyi Alexander, 1940 ( Alexander 1940a), but is differentiated from them by the following characters: femora entirely yellow (yellow basally with darker apical parts in Ormosia (Oreophila) flaveola and Ormosia (Oreophila) yankovskyi ).