Drasteria caucasica caucasica ( Kolenati, 1846 )

Yu, Alexey & Korb, Stanislav K., 2019, A revision of the genus Drasteria of Central Asia and Kazakhstan with special attention to the adjacent areas (Lepidoptera: Erebidae), Zootaxa 4673 (1), pp. 1-104 : 37-38

Drasteria caucasica caucasica ( Kolenati, 1846)

(= D. astrida (Eversmann, 1857)) ( Map 12 ) ( Figs. 103–114 (imago), 234, 235 (male genitalia), 279 (female genitalia))

Kolenati, 1846: 104; Tab. 18, fig. 4 ( Euclidia Caucasica ). Type locality: “in Caucaso Lhesgico et Iberia ” (by the original de- scription). Type material. Not found. = astrida Eversmann, 1857: 399 ; pl. 3, fig. 2 ( Ophiusa Astrida ). Type locality: “l’Armenie russe” (by original description);

“ Armenia ” (by the lectotype designation) .

Type material. Lectotype male, (designated here in accordance with ICZN Art. 74) ( Figs. 112–114 ), labelled: “ Armenia ” (white paper, handwritten); “685a” (white paper, handwritten); “an Var ?” (white paper, handwritten); “coll Eversmann ” (white paper, printed) ( ZISP) . Paralectotype: 1 female ( ZISP) .

= aksuensis Fuchs, 1903: 80 (Leuc.[anitis] caucasica Koll. var. aksuensis ). Type locality: “aus Aksu ” (by original description).

Type material. Not found.

Alphéraky, 1882: 93 [Khorgosse]; Staudinger, Rebel, 1901: 244 [Ili]; Fuchs, 1903: 50 [Aksu]; John, 1910: 619 [Uralsk, Perovsk, Khorgos]; Uvarov, 1910: 168 [Kok-Dzhida in the confluence of the rivers Emba and Kuldenen-Temir]; Warren 1913:392 [Ili]; Kuznetzov, 1960: 87 [Askhabad]; Rafes, 1960: 157 [Narun sands]; Daricheva, 1965: 57 [Kara-Kala; Sumbar valley]; Shek, 1972: 143 [Chimkent Prov.]; Sukhareva, 1972: 62 [Ispaz]; Shekh, 1975: 414 [Chimkent, Dzhezkazgan, Alma-Ata, Taldy-Kurgan Prov]; Degtyareva, 1981: 130 [Karategin]; Ronkay, 1983: 238 [Chovd aimak: 10 km SSW von Somon Bulgan, 1200 m]; Daricheva et al., 1983: 38 [Lambe]; Radzhabova, 1985: 25 [Kuraminsky Mts.: Tashbulak valley, Cholata]; Ishkov, Sukhareva, 1986: 124 [Aksu-Dzhabagly Nature Reserve: Novonikolaevka, Dzhabagly]; Nekrasov et al., 1988: 51 [Arslanbob, Isfana, Naryn, Pokrovka, Dugoba, Dzhekendy (partim, D. hyblaeoides )]; Poole, 1989: 327, 328; Klyuchko, Kreuzberg, 1998: 26 [Nuratinsky Nature Reserve]; Radzhabova, 1992: 181 [North Tajikistan]; Beck, 1996: 25; Beskokotov, 1996: 162 [Aksu-Dzhabagly Nature Reserve]; Milko, 1996: 258 [Regions close to Fergana (including southern slopes of Chatkalsky and Fergansky Mts. and northern slopes of Alai and Turkestansky Mts.), Issyk-Kul depression (including northern slopes of Terskey Ala-Too Mts., southern slopes of Kungey Ala-Too Mts. and Tyup river valley), Inner Tian-Shan, Alai (partim, D. hyblaeoides )]; Weisert, 1997: 73 [Provinz Mary, Karakum-Wiiste, 12km N Zakhmet; Provinz Mary, 14km N Zakhmet; Provinz Chardzhou, 18km N Farap, Auwald, Amu -Dar’ya-FluB; Provinz Chardzhou, Repetek NSG; Provinz Mary, 35km S Yolatan]; Lehmann et al., 1998: 527 [33.37’N, 79.21’О, Sarytogai, Tal des Flusses Sharyn, ca 20 km vor der Mündung in den Ili, 666m; Ili area, Tschu river, Mujunkum, 200m; Ili river, Bakanas, 400m; Ili area, Charyn river, 400m; Alma-Ata area, Ili river, Taschkarasu, 450m; Sarikamysch Mts.. River Kjokjomeren. 5 km n Amok;Alai mts., Maidan vill.]; Poltavsky et al., 1998: 108 [Lambe, Kara-Kala, Tschumamed-kum; Iolotan, Tschardzhou]; Ivinskis, Miatleuski, 1999: 203 [Parchay, Ai-Dere, Sandykatschi]; Anikin et al., 2000: 332 [Astrakhan]; Lehmann, 2000: 95 [Jurievka und Umgebung bei Syn-Tasch, 42° 42’ N, 75°03’ E; Jurievka bei Syn-Tasch, 42°42’ N, 75°03’ E; Karadon südlich Jurievka, 42°36’ N, 75°02’ E]; Hacker, Miatleuski, 2001: 814 [Sajkhin; Urda; Kandagash; Dzhanibek]; Nupponen, Fibiger, 2002: 135 [50°40-45’ N 54°26 -28’E, 170-230 m, Pokrovka village 20 km S, Schibendy valley; 50°58’N 54°25’E, 100 m, near Burannoe village, Ilek river valley]; Poltavsky, Ilyina, 2002: 17 [Sumur]; Poltavsky, Nekrasov, 2002: 26 [Rostov-on-Don Region, Krasnodar Region, Ichkeria Republic, Kalmykia Republic, Daghestan Republic]; Goater et al., 2003: 78 [ Asia Minor and Central Asia, extending westwards into Europe, in Kazakhstan: Sajkhin, Urda, Kandagash and Dzhanibek and across the north of the Black Sea to the eastern parts of Rumania and Bulgaria]; Lehmann, Bergmann, 2005: 20 [Northern steppe plain, Issyk Kul depression, Susamyrtoo, Dzhumgaltoo, Sarykamish Mts., Kirgizsky Mts., Tshatkalsky Mts., Central and western part (Naryn), Toktogul area, Fergansky Mts., Turkestansky Mts., Alai Mts., Togous-Toraje (partim, D. hyblaeoides )]; Poltavsky et al., 2008: 227 [Bessergenevskaya, Kalinin, Konygin, Nizhnekundryuchenskaya, Razdorskaya]; Bálint, Benedek, 2009: 7 [ Mongolia, Hovd aimag, Mongol Altay Mts, 30 km N of Uyench, Uyench river valley, N46°22.978’, E92°07.856’, 1,916 m]; Benedek, Bálint, 2009: 163 [Prov. Almaty, Toraygir Mts, near Alasay pass, 1340 m, N43°18’ E078°56’]; Kononenko, 2010: 89 [ Kazakhstan]; Poltavsky et al., 2010: 95 [Botkul]; Gorbunov, 2011: 57 [West Kazakhstan without Mangyshlak and South Ustyurt]; Lehmann & Zahiri, 2011: 150 [ Iran: Gate Dehh, Aras]; Korb, 2012: 518 [ Tajikistan, Shugnansky Mts., 2,5 km SW of kishlak Vodzh, 2672 m (N 37 41.887, E 71 55.821); Tajikistan, Severo-Alichurskiy Mts., road to Murgab, 5,84 km SSW from Karasu, 3943 m (N 37 57.870, E 73 54.587)]; Benedek, Bálint, 2013: 139, 141 [Province Almaty, 15 km N of Masak, Ayak-Kalkan, 500 m, N 43° 45’, E 078° 30’; Province Almaty, Toraygir Mts, at the Charyn River, 1060 m, N 43° 16’, E 078° 58’ (partim, D. hyblaeoides ); Province Almaty, 9 km N of Kegen, near Kegen Pass, 1700 m, N 43° 10’, E 079° 13’]; Korb, 2013: 28 [Bridge through Ili River 22 km SE of Koktal, Ili River Valley, N 43 58.004 E 79 35.905, 600 m; 35 km N of Bakanas, Ili River Valley, N 44 53.940 E 75 53.479, 377 m; Singing Dune in Altyn Emel Nature Reserve, eastern border of Kapchagai Reservoir, Ili River Valley, N 43 50.273 E 78 35.759, 467 m]; Kazenas, 2014a: 148 [National Park “Altyn-Emel”]; Kazenas, 2014b: 80 [Charynsky National Park]; Poltavsky, 2016: 19 [Rostov-on-Don Province]; Shovkoon, Trofimova, 2016: 607 [Aktobe Prov., Bolshie Barsuki sands, N 46°58’; E 59°13’; Aktobe Prov., Shagyrai Plateau, N 46°45’; E 57°31’; Aktobe Prov., Shagyrai Plateau, N 46°44’; E 57°30’].

Habitat ( Figs. 297–304 , 306, 309, 311–314, 319 ). Inhabitant of lowlands, living in steppes, semideserts and dry meadows. Elevation: up to 1800 m.

Distribution. South-East part of Europe (South Ukraine, South Russia (Krasnodar, Rostov-on-Don, Stavropol, Astrakhan, southern part of Volgograd Prov., Chechnya, Daghestan, Crimea), South Moldavia, South-East Romania, East Bulgaria), Transcaucasus, Kazakhstan (excl. Central), Asia Minor, deserts and semideserts of Central Asia.

Material examined (in total: 554 specimens). Armenia. 1 ♂, Armenia, 685a, slide Matov 0522 (lectotype of astrida ) ( ZISP) ; 1 ♀, Armenia , 685b, ex coll. Eversmann, ex coll. Grand Duke Nikolai Mikhailovich (paralectotype of astrida ) ( ZISP) ; 1 ♀, 5.05.1925, Armenia, pr. Eriwan, Parakar , leg. A. Schelkovnikov ( ZISP) ; 1 ♀, 5.09.1932, Araks river , 10 km upper Mindzhevan, [leg. M. Ryabov] ( ZISP) ; 1 ♂, 18.05.1957, Erevan, zoo, leg. A. Zaguljaev ( ZISP) ; 1 ♀, 9.07.1970, Nakhichevan, Pazmara in Ordubad distr. , 1500 m, leg. A.V. Zvetaev ( ZMMU) ; 2 ♂, 1 ♀, 22.06.1973, Arzni , leg. Isakov ( ZISP) ; 1 ♂, 22.06.1985, Nurnus , leg. V.Ambarzumyan, ex coll. A. Nekrasov ( ZISP) ; 1 ♀, 20.06.2002, Gegamsky range, Khosrov res., leg. M. Kalashian, ex coll. A. Nekrasov ( ZISP) ; 1 ♂, 10.06.2010, Tavusksk Prov., Dilizhansky Nature Reserve , leg. A.V. Krupitsky ( CKO) . Azerbaijan. 1 ♂, 7 ♀, 16–28.04.[18]84, Warvara , leg. Chr [istoph], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 1 ♀, Helenendorf , leg. Leder, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 2 ♀, Helenendorf , [leg. Leder], ex coll. Erschov ( ZISP) ; 3 ♂, 1 ♀, Elisabethpol , leg. unknown, ex coll. O. John ( ZISP) ; 2 ♀, 26.06– 17.07.1901, Elisavetpol Prov., Geoktepe , leg. R. Shmidt ( ZISP) ; 1 ♂, 1 ♀, 11.05.1902, Kr. Aresch , leg. E. Koenig ( ZISP) ; 1 ♀, 08.1914, Elizavetpol Prov., Aresh distr., Geoktapa , leg. Mordvilko ( ZISP) ; 1 ♀, 2.05.1916, Kreis Nucha , leg. E. Koenig ( ZISP) ; 1 ♂, 8.05.1926, Trans- cauc., Grünfeld , leg. I. Kozhantschikov ( ZISP) ; 1 ♀, 10.05.1926, Salyanovsky distr., Dzhafarkhan , leg. Aleksandrov ( ZISP) ; 2 ♂, 19.07.1933, Nakhichevan distr., Disar near Ordubad, leg. Znoiko ( ZISP) ; 2 ♂, 10.05.1955, Nakhiche- van ASSR, Dzhagry , leg. A. Zagulyaev ; 1 ♀, 30.05.1979, Talysh Mts., Gosmolyan , h= 1300 m, leg. A. Dantchenko, ex coll. A. Nekrasov ( ZISP) . China. 2 ♂, Ost-Turkestan, Aksu , Rückbeil ( ZMHU) ; 2 ♀, Kuldja , 3- 16.04.1879, leg. Alph [eraky], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 16.04.[18]79, Kuldja , [leg. Alpheraky], ex coll. Erschov ( ZISP) ; 2 ♂, 26.04.[?], Culdscha , leg. unknown ( ZISP) ; 2 ♂, Aksu , leg. M. Bartel, ex coll. O. John ( ZISP) . Georgia. 1 ♀, 29.08.[?], Batum , leg. unknown ( ZISP) ; 1 ♀, 14.06.[18]95, Tiflis, leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 2 ♂, 18–20.05.[18]96, Tiflis Prov., Chachuny, leg. Mlokosevich ( ZISP) ; 1 ♀, 16– 25.06.1910, Tiflis, leg. P. Satunin ( ZISP) ; 1 ♀, 24.05.1915, distr. Tiflis, loc. Mtshet , leg. L. Bankovskiy, ex coll. Tshetverikov ( ZISP) ; 2 ♀, 13.09.1934, 26.06.1947, Sukhum , [leg. E. Milyanovsky] ( ZISP) ; 2 ♂, 4 ♀, 30.05– 18.07.1945, 12.06.1946, 8.06.1947, Gagry, leg. E. Miljanovskij ( ZISP) . Kazakhstan. 2 ♂, 2 ♀, Uralsk , leg. un- known, ex coll. O. John ( ZISP) ; 1 ♂, 1 ♀, Aulie Ata , leg. unknown, ex coll. O. John ( ZISP) ; 1 ♂, 7.05.[18]85, Shuran , leg. unknown, ex coll. N. Filipyev ( ZISP) ; 2 ♂, 5– 15.05.1905, Perovsk , leg. Tell ( ZISP) ; 2 ♀, 24.04.1907, prov. Semiretshensk, prope Dzharkent, ad fl. Ussek , leg. M. Divnogorsky ( ZISP) ; 3 ♀, 3.05.1907, prov. Semiretsh- ensk, prope Dzharkent , leg. M. Divnogorsky ( ZISP) ; 1 ♀, 9.05.1907, prov. Semiretshensk, prope Dzharkent, ad fl. Ili , leg. M. Divnogorsky ( ZISP) ; 1 ♀, 25.05.1907, Bolshie Barsuki, Chelkar lake , leg. Volman ( ZISP) ; 1 ♂, 5 ♀, 1908, Syr-Darja, Perovsk , leg. Nikolsky, ex coll. O. John ( ZISP) ; 2 ♂, 23.05.1908, Emba, Kok-Dzhida valley , leg. D. Gorodin ( ZISP) ; 5 ♂, 2 ♀, 24.04– 11.06.1909, prov. Syr-Darjensis, Perovsk circ., leg. E. Miller ( ZISP) ; 10 ♂, 1 ♀, 30.07– 7.08.1909, Syr-Darja, Aj-Darle , leg. Koshantschikoff, ex coll. O. John ( ZISP) ; 1 ♂, 4 ♀, 21.05.1910, Semirechye Prov., Kopal distr. , village Iliyskaya, leg. N. Tshetverikov ; 6 ♂, 2 ♀, 3.07– 5.08.1910, Syr-Darja, Dshulek , leg. Koshantschikoff, ex coll. O. John ( ZISP) ; 1 ♀, 7.07.1910, Cherny Irtysh , leg. А. Yakobson, ex coll. O. John ( ZISP) ; 8 ♂, 14 ♀, 20.07– 15.08.1910, 7.06– 9.08.1911, Syr-Daria, Baigacum , leg. Koshantschikoff, ex coll. O. John ( ZISP) ; 1 ♀, 15.06.1911, Burly-Tau Mt. , leg. Molchanov ( ZISP) ; 1 ♂, 10.07.1911, Perovsk , leg. Zarudny ( ZISP) ; 1 ♂, 12.04.1912, Golodnaya Steppe , leg. unknown ( ZISP) ; 6 ♂, 7 ♀, 1915, Semiretshye Prov., Dzharkent , leg. unknown ( ZISP) ; 3 ♂, 04.1915, Dzhar-Khuduk on the southern Dzharkent , leg. V. Kavrigin ( ZISP) ; 2 ♂, end 04.1915, Semirechye, Chulokai on the south of Dzharkent , leg. V. Kavrigin ( ZISP) ; 2 ♂, 06.1916, Perovsk Prov., Kara-Uzyak , leg. unknown ( ZISP) ; 1 ♀, 27.06.1930, Chelkar , leg. А. Gozhev ( ZISP) ; 2 ♀, 25.07– 28.08.1933, Bol- shie Barsuki, Chelkar , leg. Luppova ( ZISP) ; 1 ♂, 6.07.1951, Chelkar , leg. unknown ( ZISP) ; 2 ♂, 31.05.1952, Ka- zachenkov settl., Bogdy forest , leg. Levin ( ZISP) ; 1 ♂, 18.06.1952, Urda , leg. Levin ( ZISP) ; 1 ♂, 9.07.1954, Turke- stan, leg. A. Bundel ( ZISP) ; 1 ♂, 29.05.1966, 80 km NW Turkestan, Akkum , leg. Pastukhov ( ZISP) ; 1 ♂, 31.05.1966, 30 km NW Kyzyl-Orda , leg. Pastukhov ( ZISP) ; 1 ♂, 6.06.1966, 40 km W Kyzyl-Orda , leg. Pastukhov ( ZISP) ; 1 ♀, 8.07.1976, Aksu-Dzhabagly Nature Reserve, Dzhabagly , leg. Ishkov ( ZISP) ; 1 ♂, 2.06. 1988, 150 km NW Alma- Ata, right shore of Ili, Mynbulak valley , leg. M. Falkovich ( ZISP) ; 1 ♀, 24.05.1989, Moyunkum , leg. Amirguzhin ( ZISP) ; 1 ♂, 2.05.1990, Zhel-Turanga , leg. S. Toropov, ex coll. A. Nekrasov, microvial N 962g Nekrasov ( ZISP) ; 1 ♂, 4.07.2010, Ili river valley, bridge near Koktal , N 43 58.004 E 79 35.905, 600 m (leg. S.K. Korb) ; 1 ♂, 2011, Transili Alataua Mts., Chingeldy valley , leg. P.Egorov ( SKK) ; 2 ♂, 1 ♀, 25– 27.04.2011, Almaty Prov., 32 km NW of Bakanas, Ili river , leg. S.A. Knyazev ( CKO) ; 1 ♂, 28.04.2011, Altyn-Emel Nature Reserve, Singing Dunes , 400 m, leg. S.K. Korb ( SKK) ; 1 ♀, 17.05.2011, Kazaly distr., Basykara env., near r. Syr Daria , h= 67 m, 45.755° N 62.303° E, leg. A. Rokhletzova ( ZISP) ; 8 ♂, 2 ♀, 30.06– 1.07.2012, Alma-Ata Prov., Panfilov distr., 25 km SW of Koktal, Ili river , leg. S. Rybalkin ( CKO) ; 1 ♀, 7– 8.05.2013, Altyn-Emel Nature Reserve, Singing Dunes , 400 m, leg S.A. Knyazev ( CKO) ; 1 ♂, 30.06.2015, Transilian Alatau Mts., near Koram , N43° 29.307’ E78° 10.240’, 786 m, leg. S.K. Korb ( SKK) ; 4 ♂, 2 ♀, 1– 3.07.2015, Ili river valley near Kok-Tal bridge, N43° 57.818’ E79° 36.322’, 500 m, leg. S.K. Korb ( SKK) ; 1 ♀, 1– 3.07.2015, Ili river, valley near Kok-Tal bridge, h= 500 m, 43°57.818’ N 79°36.322’ E, leg. A. Nikolaev ( ZISP) ; 1 ♀, 4– 5.05.2017, Kyzylorda Prov. , 13 km N of Aiteke Bi, sands, leg. S.A. Knyazev ( CKO) ; 1 ♂, 27.05.2018, Zaisan district, 30 km NW of Maikapchagaj, Bozaigyrkum Sands , h= 530 m, 47°41’ N 85°20’ E, leg. P. Gorbunov ( ZISP) ; 2 ♂, 12– 14.06.2018, Aidarly env., 43.993779 N, 79.483105 E, leg. A. Litovtsev ( SKK) . Kyrgyzstan. 2 ♀, Tura, Kudara, leg. Gr [umm]. Gr [zhimajlo], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 1 ♀, Osch, leg. unknown, ex coll. N. Filipyev ( ZISP) ; 1 ♀, 6.05.[?], Toguz-Bulak , leg. Zarudny ( ZISP) ; 2 ♂, 2 ♀, 12.04–29.06.[18]84, Osh, leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, 30.07.1951, Alai valley, Daraut-Kurgan , leg. A.V. Zvetaev ( ZMMU) ; 1 ♂, 1 ♀, 30.07.1951, Alai, Daraut Kurgan , [leg. A. Bundel] ( ZISP) ; 4 ♂, 1 ♀, 31.07.1951, Alai valley, Dzhekendy , leg. A.V. Zvetaev ( ZMMU) ; 1 ♂, 8.08.1952, Alai valley, Dzhekaindy river , h= 2500 m, leg. A. Bundel ( ZISP) ; 1 ♂, 1 ♀, 12.07.1955, Osch, leg. A. Bundel ( ZISP) ; 1 ♂, 3.07.1957, Terskey Ala-Too Mts., Pokrovka , leg. A.V. Zvetaev ( ZMMU) ; 1 ♀, 1961, Osh, leg. Zherdenko ( ZISP) ; 1 ♀, 16.07.1986, Issyk-Kul, Toru-Aygyr , leg. A. Kondratyev, ex coll. A. Nekrasov ( ZISP) ; 1 ♀, 18.07.2014, Kyrgyzstan , Bishkek, 900 m, leg. S.K. Korb ( SKK) ; 4 ♂, 21.07.2014, Kirghizsky Mts., Bishkek environs, Sarban village , 42°47’55.47”N, 74°30’45.70”E, leg. S.K. Korb ( SKK) ; 1 ♂, 12.07.2015, Kirghizsky Mts., Ala-Too settl. near Bishkek, N42° 45.296’ E74° 40.329’, 1254 m, leg. S.K. Korb ( SKK) ; 2 ♂, 13.07.2015, Fergansky Mts., 5,5 km S of Imeni Chkalova (S shore of Toktogul reservoir), N41° 43.223’ E72° 57.165’, 1768 m, leg. S.K. Korb ( SKK) ; 1 ♂, 14.07.2015, Alai Mts., small valley between Tashkoro and Karabulak , N40° 14.119’ E73° 24.484’, 1805 m, leg. S.K. Korb ( SKK) . Romania. 1 ♂, 12.07.1991, Delta. C.A. Rasetti , leg. L. Szekely ( HNHM) . Russia. 1 ♂, 30.05.1926, Stavropol Prov., Praskovea , [leg. M. Ryabov] ( ZISP) ; 1 ♂, 08.1935, North Caucasus, Prikumsk , leg. A.V. Zvetaev ( ZMMU) ; 1 ♂, 18.08.1947, Makhachkala , DSHI, lake, leg. M. Ryabov ( ZISP) ; 1 ♀, 21.07.1949, Kapchugai , [leg. N. Egorov] ( ZISP) ; 1 ♀, 31.07.1953, Volgograd Prov., Tingutinsk forest , leg. Mazurkevich ( ZISP) ; 1 ♂, 19.08.1968, Astrakhan Prov., Osypnoi Bugor , leg. Penchukovskaya ( ZISP) ; 1 ♂, 23.06.1970, Astrakhan Prov., Dosang , leg. А. Lvovsky ( ZISP) ; 1 ♂, 19.08.1999, Krasnodar area, Anapa, Dzhemsta , leg. V. Stshurov ( ZISP) ; 1 ♂, 10.09.2000, Rostov Prov., 40 km SE Millerovo, Efremovo-Stepanovka , leg. A. Poltavsky ( ZISP) ; 2 ♂, 1 ♀, 27.07.2001, 25 km SW Buynaksk, Balakhani vill., leg. A. Magomedova ( ZISP) ; 1 ♂, 08.2001, Volgograd Prov., Bolshaja Golubaya riv. bas., leg. S. Kunitzin, ex coll. А. Nekrasov ( ZISP) ; 1 ♂, 4.05.[20]02, Astrakhan Prov., Akhtubinsk distr., Bolshoe Bogdo Mt. , Kordonnaya Balka, leg. D. Komarov ( ZISP) ; 1 ♂, 8– 18.06.2003, Astrakhan Prov., Akhtubinsky distr., Bogdinsko-Baskunchaksky Nature Reserve, Bogdo vic., dry steppe, leg. V. Anikin ( ZISP) ; 1 ♂, 29.06.2003, 18 km W Makhachkala, Barkhan Sarykum Nature Reserve , leg. D. Magomedova ( ZISP) ; 5 ♂, 1 ♀, 2.08.2004, 11.07.2005, Tsumadinsky distr. , Agvali, leg. E. Ilyina ( ZISP) ; 2 ♂, 10– 18.07.2007, Daghestan, Kizlar distr., Kirova settlement, leg. unknown ( ZISP) ; 1 ♂, 30.04.2009, Astrakhan Prov., Dosang , leg. S.K. Korb ( SKK) ; 1 ♀, 3.05.2010, Astrakhan Prov., Dosang , leg. E. Tsvetkov ( ZISP) ; 1 ♂, Astrakhan Prov., Bogdinsko-Baskunchaksky Nature Reserve ., near camping, 48°09’ N 46°49’ E, leg. S. Nedoshivina. Tajikistan. 1 ♂, Kalininabad , leg. O. Legezin, ex coll. A. Nekra- sov ( ZISP) ; 1 ♂, 1 ♀, end 07.[?], Parkhar , Kyzylsu, leg. unknown ( ZISP) ; 19 ♂, 5 ♀, 08.1909, 28.05– 15.06.1910, 28.05– 1.07.1911, Pamir , Post Chorog, leg. A. Bereskij ( ZISP) ; 1 ♀, 1912, Pamir , leg. A. Berezsky ( ZISP) ; 1 ♂, 2 ♀, 16– 22.07.1915, Khorog , leg. Lazdin ( ZISP) ; 1 ♀, 13.06.1941, Dzhilikul on Vakhsh river, leg. Gussakovsky ( ZISP) ; 1 ♀, 4.04.1948, Peschany Pass , 8 km N Bura-Tau, leg. Stshetkin ( ZISP) ; 1 ♂, 3 ♀, 9.05.1948, 23.04– 15.05.1949, Vakhsh river , Staraya Pristan, leg. Stshetkin ( ZISP) ; 1 ♀, 18.05.1951, Stalinabad vic., Kurgan-Tyube Pass , [leg. A. Bundel] ( ZISP) ; 1 ♂, 1 ♀, 1.08.1951, Karamuk valley , h= 2450 m, leg. A. Bundel ( ZISP) ; 1 ♂, 3 ♀, 30.07.1955, distr. Wantsch, loc. Djowid , h= 2100 m, leg. A. Bundel ( ZISP) ; 1 ♀, 5.08.1955, Wantsch, Garm-Tscaschma , h= 2300 m, leg. A. Bundel ( ZISP) ; 1 ♂, 6.08.1956, Pamir merid. occ., mts. Schachdarensis, fl. Badom-Dara , h= 3400 m, leg. A. Bundel ( ZISP) ; 1 ♀, 8.08.1956, Pamir mer. occ., fl. Schach-dara, loc. Barwoz , h= 3000 m, leg. A. Bundel ( ZISP) ; 21 ♂, 11 ♀, 10.08.1956, 15.07– 18.08.1957, 20.07.1960, 9.08.1961, Pamir mer. occ., Chorog , h= 2300-2340 m, leg. A. Bundel ( ZISP) ; 1 ♂, 7 ♀, 9.07.1957, SW Tajikistan, Tigrovaya Balka Nature reserve , leg. A.V. Zvetaev ( ZMMU) ; 1 ♀, 10.07.1957, Pamir merid. occ., mts. Schugnanensis, cl. sept., fl. Akmamed-Dara , h= 3300 m, leg. A. Bundel ( ZISP) ; 1 ♂, 1 ♀, 25.07.1957, Pamir merid. occ., mts. Schachdarensis , cliv. sept., fl. Seidj-Dara, h= 3200 m, leg. A. Bundel ( ZISP) ; 37 ♂, 25 ♀, 26.05.1958, 14.05– 1.09.1959, 26.05– 13.08.1960, 3– 18.05.1961, 26.04– 29.05.1962, 17.06.1963, 14– 26.05.1965 ,2.05– 6.09.1966, 29.08.1967, 12.05– 26.06.1968, 2– 19.05.1969, 25.08.1971, 16.04.1974, 27.05.1975, 7.06.1976, 6.05.1978, 30.04– 24.05.1982, 27.07.1983, 20.05.1988, Pamir, Khorog, botanical garden, h= 2300 m, leg. М. Zapryagaev, slide Matov 0177 (♂), microvial N 962а Nekrasov ( ZISP) ; 1 ♂, 24.06.1965, Iskan- der-Darya valley, h= 1700 m, leg. Falkovich ( ZISP) ; 5 ♂, 7 ♀, 25– 26.04.1966, Gissar Mts., Tigrovaya Balka , leg. A. Danilevsky ( ZISP) ; 1 ♂, 2 ♀, 31.05– 7.06.1969, Khorog , botanical garden, leg. Martynova ( ZISP) ; 2 ♂, 1 ♀, 6.06.1973, W Pamir, Khorog , near botanical garden, leg. Isakov ( ZISP) ; 5 ♂, 2 ♀, 17– 19.05.1974, Tigrovaya Balka Nature Reserve , leg. V. Prasolov ( ZISP) ; 3 ♂, 2 ♀, 28.08.1974, W. Pamir, Vanch river, kishlak Dzhoud , h= 2000 m, leg. Zapryagaev ( ZISP) ; 4 ♂, 2 ♀, 24.06.1976, Tigrovaya Balka Nature Reserve , leg. A. Lvovsky ( ZISP) ; 1 ♀, 27.08.1977, Vakhanskij Chr. , 2200 m, Coll. J. Wojtusiak ( HNHM) ; 1 ♂, 7.07.1979, Peter the Great Mts., Muksu valley , kishlak Muk , leg. V. Prasolov ( ZISP) ; 1 ♂, 1 ♀, 28.07– 1.08.1980, Pamir, Khorog , botanical garden, leg. V. Prasolov ( ZISP) ; 1 ♂, 1 ♀, 29.07.1980, Pamir, Khorog, Shakhdara valley , leg. V. Prasolov ( ZISP) ; 1 ♀, 13.07.1983, Khorog env., kishlak Shod , leg. V. Prasolov ( ZISP) ; 1 ♂, 15.07.1984, Garm , leg. A. Ivanov ( ZISP) ; 1 ♂, 10.06.1988, West Pamir, kishlak Shod, leg. M. Zapryagaev, ex coll. A. Nekrasov, microvial N 962v Nekrasov ( ZISP) ; 1 ♀, 12.07.2001, Hindukush, Gunis, leg. M. Markhasev, microvial N 3344 Nekrasov , ex coll. A. Nekrasov ( ZISP) ; 1 ♀, 14– 29.07.2011, 71 km N of Khorog, Shipad kishlak, leg. A. Zubov ( CKO) . Turkey. 2 ♂, 30.06.[?], Kagyzman , leg. unknown, ex coll. Dyakonov ( ZISP) ; 4 ♂, 2 ♀, 3.05– 30.06.1911, Kars Prov., Kagyzman , h= 4500 feet, leg. Е. Miller, ex coll. Tshetverikov ( ZISP) ; 1 ♂, 9.07.1977, Göreme, Törökország , leg. Podlussány ( HNHM) ; 1 ♀, 8.08.1988, Prov. Erzurum-Erzincan , 10 km W of Aþkale, 1700 m, 40º34´E, 39º50´N, leg. Gyulai, Hreblay, Ronkay and Ronkay ( HNHM) ; 1 ♂, 28.06.1996, Prov. Erzincan, 46 km W of Tercan 1200 m, leg. Podlussány ( HNHM) . Turkmenistan. 2 ♂, 23–25.04.[?], Chardzhui , leg. Zarudny ( ZISP) ; 4 ♂, 18.05– 18.07.1894, Transcaspien, Sumbar , leg. Herz, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 2 ♂, 1 ♀, 20.04– 25.06.1913, Chardzhui, leg. Samok- ish; 2 ♂, 1 ♀, 25.05.1952, 5.06.1953, Kara-Kala , institute garden, leg. V. Kuznetzov ( ZISP) ; 1 ♀, 13.09.1952, Iolo- tani settlement environs, leg. Vydkova ( ZMMU) ; 1 ♀, 2.05.1962, Kara-Kala , leg. Krasilnikova ( ZISP) ; 2 ♂, 3 ♀, 1– 2.06.1965, 70 km NW Chardzhou, Ispas , leg. М. Falkovitch ( ZISP) ; 2 ♀, 9.04.1966, Kulatau Tugai on Amudarya river, leg. I. Sukhareva ( ZISP) ; 1 ♂, 5.06.1986, W Kopet Dagh, Aidere , h= 850 m, leg. M. Falkovitch ( ZISP) ; 1 ♀, 16.08.1990, Takhta vic., Kuibyshevo Sovkhoz , leg. V. Mironov ( ZISP) . Ukraine. 1 ♀, 7– 10.07.1961, Kherson Prov., Skadovsk distr. , Novaya Alekseevka, leg. unknown ( ZISP) ; 1 ♂, 1 ♀, 10.08.1964, Dnepropetrovsk Prov., Dnepropetrovsk, leg. unknown ( ZISP) ; 1 ♀, 27.08.1987, Lugansk Prov., Sverdlovsk distr., Provalye , leg. P. Ply- ustsh, ex coll. A. Nekrasov ( ZISP) ; 1 ♂, 1.05.1994, Odessa Prov., Odessa vic., Khadzhibeisky Liman , leg. R. Prokhorov ( ZISP) ; 1 ♂, 26.06.1995, Tavriisk , leg. A. Khalin ( ZISP) ; 1 ♀, 19.07.1995, Kherson Prov., Novaya Kak- hovka, leg. A. Khalin ( ZISP) ; 1 ♂, 1– 13.06.2003, N Arabatskaja strelka, 5 km S vill. Stchastlivzevo , leg. I. Solodovnikov ( ZISP) ; 1 ♀, 15.05.2010, Zaporozhye Prov., Melitopol distr., Troizkoe , leg. I. Msyanikov, I. Kovalev ( SKK) . Uzbekistan. 1 ♂, Russian Turkestan , [leg. Fedtchenko], ex coll. Erschov ( ZISP) ; 1 ♂, 1 ♀, Margelan , leg. Alph [eraky], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 2 ♂, 5.05.1903, Khodzhent Prov., Begovat , leg. Valnev ( ZISP) ; 1 ♂, 05.1904, Namangan distr. , leg. Yankovsky ( ZISP) ; 1 ♂, 18.06.[19]08, Fergana sept., Naman- gan, Fl. Padscha-ata, Toste , leg. B. Grigoriev, ex coll. O. John ( ZISP) ; 1 ♀, 24.08.1910, Tashkent , leg. Zarudny ( ZISP) ; 5 ♀, 4– 19.04.1911, Buchara, stc. Farab , leg. A. Golbeck ( ZISP) ; 1 ♂, 2 ♀, 21.04– 14.07.1912, Termez , leg. А. Kirichenko, ex coll. Dyakonov ( ZISP) ; 1 ♀, 10.07.1916, Tashkent , leg. Gerkhner ( ZISP) ; 1 ♂, 1 ♀, 2.05– 2.07.1927, Khiva , leg. L. Zimin ( ZISP) ; 1 ♀, 14.05.1928, Buchara, Adisabad , leg. A. Gerasimov ( ZISP) ; 1 ♀, 27.05.1928, Buchara, Chargush , leg. A. Gerasimov ( ZISP) ; 1 ♂, 2.06.1928, Buchara, Yargak pr. Chatyrtshy , leg. L. Zimin ( ZISP) ; 1 ♂, 4 ♀, 24.07– 2.08.1929, Chiva, Urgenč , leg. A. Gerasimov ( ZISP) ; 1 ♂, 20.07.1935, Golodnaya Step’ , leg. N.Fursov ( ZMMU) ; 5 ♂, 3 ♀, 22.06.1938, Samarkand Prov., Aman-Kutan valley , leg. A.V. Zvetaev ( ZMMU) ; 1 ♀, 4.08.1958, Ferghana Prov., Kokand, Buivada , leg. A.V. Zvetaev ( ZMMU) ; 1 ♂, 1 ♀, 2– 10.05.1977, Baisuntau Mts., Vakhshivar , h= 1450 m, leg. A. Nekrasov, ex coll. A. Nekrasov ( ZISP) ; 1 ♀, 27.08.1982, Kash- kadarya Prov., Mubarek , leg. А. Kondratyev, ex coll. A. Nekrasov ( ZISP) .

Figures

MAP 12. Distribution map of Drasteria caucasica (Kolenati, 1846).

FIGURES 85–105. Drasteria specimens. 85–87. D. rada (Boisduval, 1848), Altyn-Emel Nature Reserve, Kazakhstan (85), Charyn Canyon, Kazakhstan (86), Ili river valley, Kazakhstan (87). 88–90. D. rada sibirica (Kozhantschikov, 1925), holotype (88: upperside; 89: underside; 90: labels). 91–96. D. christophi (Alphéraky, 1895), lectotype (91: upperside; 92: underside; 93: labels), holotype of dubia Bang-Haas in litt. (94: upperside; 95: underside; 96: labels). 97–99. D. kabylaria (Bang-Haas, 1906), lectotype (97: upperside; 98: underside; 99: labels). 100–102. D. kabylaria columbina (Brandt, 1941), paratype (100: upperside; 101: underside; 102: labels). 103–105. D. caucasica (Kolenati, 1848), Arzni, Armenia (103, 104: male; 105: female).

FIGURES 106–126. Drasteria specimens. 106–111. D. caucasica (Kolenati, 1848), Bishkek environs, Kyrgyzstan (106, 107), Usek valley, Dzhungar Ala-Too, Kazakhstan (phenotype hyblaeoides) (108), Uch-Turfan, Kashgaria, China (phenotype hyblaeoides) (109, 110), Usek valley, Dzhungar Alatau Mts., Kazakhstan (111). 112–114: Drasteria caucasica astrida (Eversmann, 1857), lectotype (112: upperside; 113: underside; 114: labels). 115–117. D. hyblaeoides (Moore, 1878), holotype, upperside (115), underside (116), labels (117). 118–120. D. cashmirensis (Hampson, 1894), holotype (118: upperside; 119: labels), Naltar, Pakistan (120). 121–123. D. saisani saisani (Staudinger, 1882), lectotype (121: upperside, 122: underside; 123: labels). 124–126. D. saisani clara (Staudinger, 1894), lectotype (124: upperside; 125: underside; 126: labels).

FIGURES 297–304. Drasteria habitats. 297. 25 km N of Panfilov, Dzhungarian Alatau, Kazakhstan (one of the authors in the collecting trip 2010). 298. Ili river valley near Koktal bridge, Almaty Prov., Kazakhstan (Russian Entomological Society and Institute of Zoology of Kazakhstan collecting trip, 2010). 299. Sands near Dosang environs, Astrakhan Prov., Russia. 300. Saksaul forest near Bakanas, Almaty Prov., Kazakhstan. 301. Chulak gorge, foothills of Dzhungarian Alatau Mts. in the Altyn-Emel Nature Reserve, Kazakhstan. 302. Ili river shore near Singing Dunes, Altyn-Emel Nature Reserve, Kazakhstan. 303. Katutau mountains, Altyn-Emel Nature Reserve, Kazakhstan. 304. Koram environs, northern slopes of Transilian Alatau Mts., Kazakhstan. Photos: S.K. Korb.

FIGURES 305–312. Drasteria habitats. 305. Sary-Kaiky gorge, Dzhumgaltoo Mts., Kyrgyzstan. 306. Arashan environs near Bishkek, Kirghiz Mts., Kyrgyzstan. 307. Kok-Moynok environs, Terskey Ala-Too Mts., Kyrgyzstan. 308. Aksu environs, Terskey Ala-Too Mts., Kyrgyzstan. 309. Koro-Goo Pass, Moldo-Too Mts., Kyrgyzstan. 310. Alai valley, Kyrgyzstan. 311. Archaty environs, Alai Mts., Kyrgyzstan. 312. The same locality, light trapping. Photos: 305–309: S.K. Korb; 310–312: E. Komarov.

FIGURES 313–320. Drasteria habitats. 313. Kulyab environs, Tajikistan. 314. Pyandzh river near kishlak Padkhinov, Tajikistan. 315. Dunkeldyk lake, Tajikistan. 316. Vezdara river valley near Kishlak Vezd, Tajikistan. 317. Khaburobot Pass, Darvaz Mts., Tajikistan. 318. Shore of the Lake Karakul, Tajikistan. 319. Kalai-Khumb environs, Tajikistan. 320. Khorog, botanical garden environs, Tajikistan. Photos: S.K. Korb.