Eusphalerum schatzmayri, Bernhauer, 1906

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

schatzmayri Bernhauer, 1906 , see: algidum Erichson, 1840 .

tectum Paykull, 1789: 68 ( Staphylinus ; Type locality: Suecia).

— Paykull, 1792: 147 ( Staphylinus ; characters).

— Olivier, 1795:(42): 36 ( Staphylinus ; characters).

— Paykull, 1800: 411 ( Staphylinus ; characters; Sweden).

— Gravenhorst, 1806: 213 ( Omalium ; characters; Sweden).

— Gyllenhal, 1810: 202 ( Omalium ; characters).

— Olivier, 1811: 478 ( Omalium ; characters; Sweden).

— Gyllenhal, 1827: 464 ( Omalium ).

— C. Sahlberg, 1827: 278 ( Omalium ; characters; Finland).

— Zetterstedt, 1828: 49 ( Omalium ; characters; Lapland).

— Curtis, 1829: 28 ( Anthobium ; catalog; Britain).

— Mannerheim, 1830: 52 ( Omalium ; Finland; Sweden).

— Mannerheim, 1831: 466 ( Omalium ; Finland; Sweden).

— Stephens, 1834: 340 ( Anthobium ; characters; England).

— Lacordaire, 1835: 487 ( Omalium ; characters; France).

— Zetterstedt, 1838: 50 ( Omalium ; characters; habitat; Lapland; Sweden).

— Erichson, 1839a: 627 ( Deliphrum ; characters; Germany).

— Heer, 1839: 187 ( Deliphrum ; characters; Switzerland).

— Stephens, 1839: 426 ( Deliphrum ; characters; England).

— Erichson, 1840: 872 ( Deliphrum ; characters; Germany; Sweden).

— Laporte, 1840: 191 ( Omalium ; characters; France).

— Redtenbacher, 1849: 750 ( Deliphrum ; characters; Austria).

— Hardy, 1851: 58 ( Deliphrum ; notes; England).

— Fairmaire and Laboulbène, 1856: 638 ( Deliphrum ; characters; France).

— Redtenbacher, 1857: 249 ( Deliphrum ; characters; Austria).

— Kraatz, 1857d: 952 ( Deliphrum ; characters; Germany).

— Thomson, 1859: 50 (type species of Deliphrum ).

— Thomson, 1861: 197 ( Deliphrum ; characters; Scandinavia).

— Hochhuth, 1862: 103 ( Deliphrum ; Russia).

— Fauvel, 1871a: 121 [= 1872: 95] ( Deliphrum ; characters; France; Scandinavia; Britain; Germany; Switzerland; Austria; Russia).

— Redtenbacher, 1874: 270 ( Deliphrum ; characters; Austria).

— Seidlitz, 1875: 236 ( Deliphrum ; characters; Baltic region).

— J. Sahlberg, 1876: 214 ( Deliphrum ; Finland).

— Mulsant and Rey, 1880: 117 ( Deliphrum ; characters; France).

— Fowler, 1888: 406 ( Deliphrum ; characters; Britain).

— Ganglbauer, 1895: 723 ( Deliphrum ; characters; north and middle Europe).

— Luze, 1905c: 245 ( Deliphrum ; characters; northern and central Europe; Siberia).

— Reitter, 1909: 188 ( Deliphrum ; characters; Germany).

— Bernhauer and Schubert, 1910: 64 ( Deliphrum ; catalog).

— Johansen, 1914: 597 ( Deliphrum ; characters; Denmark).

— Portevin, 1929: 442 ( Deliphrum ; characters; France).

— Scheerpeltz, 1933: 1055 ( Deliphrum ; catalog).

— Scheerpeltz and Höfler, 1948: 153 ( Deliphrum ; characters; fungus inhabitant).

— Palm, 1948: 98 ( Deliphrum ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 102 ( Deliphrum ; Britain; Denmark; Norway; Sweden; Finland; Russia; Baltic States; Poland; Czech Republic; Slovakia; Romania; Bosnia; Italy; France; Switzerland; Germany; Austria).

— Lohse, 1964: 51 ( Deliphrum ; characters; central Europe).

— Smetana, 1964d: 54 ( Deliphrum ; subgenus Deliphrum ; Czechoslovakia).

— Legner and Moore, 1977: 175 ( Anthobium ; found under bark).

— Pope, 1977: 22 ( Deliphrum ; Britain).

— Hammond, 1980: 136 ( Deliphrum ; Ireland).

— Filatova, 1981: 17 ( Deliphrum ; Russia).

— Zanetti, 1987: 286 ( Deliphrum ; subgenus Deliphrum ; characters; Italy).

— Lucht, 1987: 88 ( Deliphrum ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Denmark; Sweden).

— Boháč, 1993: 41 ( Xylostiba ; Slovakia; Czech Republic).

— Muona and Viramo, 1995: 9 ( Deliphrum ; collected from fungus and dung of dog, Reindeer, and Ursus arctos ; Finland).

— Zanetti, 1995: 8 ( Deliphrum ; Italy).

— Ryabukhin, 1999: 21 ( Deliphrum ; catalog; habitat; distribution; northeastern Russia).

— Cibuļskis, 1999: 35 ( Deliphrum ; list; Latvia).

— Kashcheev, 1999a: 57 ( Deliphrum ; Kazakhstan).

- DISTRIBUTION: Finland, Sweden, Ireland, Britain, Denmark, Germany, France, Switzerland, Italy, Austria, Hungary, Czechoslovakia, Poland, Russia.

laeve Gravenhorst, 1806: 211 ( Omalium ; Type locality: Borussia).

— Olivier, 1811: 477 ( Omalium ; characters; Prussia).

— Curtis, 1829: 29 ( Omalium ; catalog; Britain).

— Stephens, 1834: 352 ( Omalium ; characters; England).

— C. Sahlberg, 1830: 282 ( Omalium ; synonym of laeviusculum Gyllenhal ).

— Erichson, 1839a: 628 ( Deliphrum ; synonym of tectum ).

— Heer, 1839: 187 ( Deliphrum ; synonym of tectum ).

— Stephens, 1839: 429 ( Omalium ; characters; England).

— Erichson, 1840: 872 ( Deliphrum ; synonym of tectum ).

— Kraatz, 1857d: 952 ( Deliphrum ; synonym of tectum ).

— Mulsant and Rey, 1880: 117 ( Deliphrum ; synonym of tectum ).

— Ganglbauer, 1895: 723 ( Deliphrum ; synonym of tectum ).

— Luze, 1905c: 245 ( Deliphrum ; synonym of tectum ).

— Bernhauer and Schubert, 1910: 64 ( Deliphrum ; synonym of tectum ).

Emodensia

[1 species; Palaearctic Region]

Emodensia Coiffait, 1977: 262 (species included: nigra ). TYPE SPECIES: Emodensia nigra Coiffait , fixed by original designation.

nigra Coiffait, 1977: 263 ( Emodensia ; Type locality: Environs du Col Mahidacla, 5000 m, région de Jumla, Népal occidental).

- DISTRIBUTION: Nepal.

Eucnecosum

[4 species; Nearctic and Palaearctic Regions]

Eucnecosum Reitter, 1909: 186 (subgenus of Arpedium ; species included: brachypterum , gyllenhali ). TYPE SPECIES: Omalium brachypterum Gravenhorst , fixed by subsequent designation by Blackwelder, 1952: 154.

— Scheerpeltz, 1933: 1060 (subgenus of Arpedium ).

— Lohse, 1963: 170 (valid genus).

— Horion, 1963: 117 (subgenus of Arpedium ; list of central European species).

— Lohse, 1964: 56 (valid genus; characters; key to central European species).

— Scheerpeltz, 1968a: 15 (catalog of Austrian species).

— Steel, 1970a: 19 (cited as Arpedium ; larval characters; biological notes).

— Tikhomirova, 1973a: 140 (checklist of species of USSR).

— Burakowski, Mroczkowski, and Stefańska, 1979: 62 (catalog; Poland).

— Muona, 1979: 18 (list of Scandinavian species).

— Uhlig and Vogler, 1981: 87 (cited as Eunecosum; list of some species collected in Germany; habitat and collecting notes).

— Campbell, 1984: 489 (characters; revison of and key to North American species).

— Muona and Viramo, 1986: 15 (cited as valid genus; list of species of northeastern Finland).

— Zanetti, 1987: 311 (characters; habitus; key to species of Italy).

— Campbell and Davies, 1991: 90 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Zanetti, 1995: 8 (list of species of Italy).

— M. Hansen, 1996: 95 (list of Danish species).

— M. Hansen, Mahler, Palm, and Pedersen, 1996: 240 (additions to list of Danish species).

— Downie and Arnett, 1996: 431 (characters; key to species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 339 (3 Nearctic species; notes; characters in key). Revelstokea Hatch, 1957: 49 , 57 (species included: hoppingi ). TYPE SPECIES: Revelstokea hoppingi Hatch , fixed by original designation and monotypy.

— Steel, 1964a: 250 (synonym of Arpedium ).

— Campbell, 1984: 489 (synonym of Eucnecosum ).

beringense Van Dyke, 1921, see: brachypterum Gravenhorst, 1802 .

brachypterum Gravenhorst, 1802: 114 ( Omalium ; Type locality: Not cited, but presumably Brunsvigae; [Note: See lectotype designation by Campbell, 1984]).

— Latreille, 1804: 372 ( Omalium ; characters; Germany).

— Gravenhorst, 1806: 203 ( Omalium ).

— Olivier, 1811: 476 ( Omalium ; characters; Germany).

— Erichson, 1840: 859 ( Arpedium ; characters; Germany; Lapland; Finland).

— Redtenbacher, 1849: 828 ( Arpedium ; characters).

— Hardy, 1851: 56 ( Arpedium ; characters; notes; England).

— Redtenbacher, 1857: 246 ( Arpedium ; characters; Germany).

— Kraatz, 1857d: 959 ( Arpedium ; characters; Germany).

— Thomson, 1861: 189 ( Arpedium ; characters; Scandinavia).

— Fauvel, 1871a: 113 [= 1872: 87] ( Arpedium ; characters; Lapland; Scandinavia; Germany; Switzerland).

— Redtenbacher, 1874: 270 ( Arpedium ; characters).

— Seidlitz, 1875: 237 ( Arpedium ; characters; Baltic region).

— J. Sahlberg, 1876: 210 ( Arpedium ; Finland).

— Mulsant and Rey, 1880: 151 ( Arpedium ; characters; France).

— Fowler, 1888: 407 ( Arpedium ; characters; Britain).

— Poppius, 1909: 5 ( Arpedium ; Lena Valley, Siberia ).

— Reitter, 1909: 187 ( Arpedium ; subgenus Eucnecosum ; characters; Germany).

— Bernhauer and Schubert, 1910: 67 ( Arpedium ; catalog).

— Johansen, 1914: 601 ( Arpedium ; characters; Denmark).

— Munster, 1933: 259 ( Arpedium ; characters; Norway).

— Scheerpeltz, 1933: 1060 ( Arpedium ; catalog).

— West, 1937: 44 ( Arpedium ; Faroe Islands).

— Palm, 1948: 111 ( Arpedium ; characters; Sweden; Denmark; Norway; Finland).

— Smetana, 1963c: 292 ( Arpedium ; subgenus Eucnecosum ; Mongolia).

— Horion, 1963: 117 ( Arpedium ; subgenus Eucnecosum ; Britain; Norway; Sweden; Finland; Russia; Denmark; Estonia; Poland; Czechoslovakia; Romania; Bulgaria; Italy; Switzerland; Germany; Austria).

— Lohse, 1964: 56 ( Eucnecosum ; characters; central Europe).

— Smetana, 1968: 228 ( Arpedium ; subgenus Eucnecosum ; Mongolia).

— Szujecki, 1969a: 248 ( Arpedium ; notes; Poland).

— Steel, 1970a: 19 ( Arpedium ; larval characters; biological notes).

— Ashworth, 1972: 214 ( Arpedium ; late-glacial subfossil; Britain).

— Smetana 1975c: 159 ( Arpedium ; Mongolia).

— Pope, 1977: 22 ( Arpedium ; Britain).

— Hammond, 1980: 136 ( Arpedium ; Ireland).

— Tóth, 1982: 83 ( Arpedium ; characters).

— Campbell, 1984: 491 ( Eucnecosum ; lectotype designation; characters; Manitoba; Northwest Territories; Yukon Territory; Alaska; Maine; New Hampshire) .

— Zanetti, 1987: 312 ( Eucnecosum ; characters; Italy).

— Schülke and Uhlig, 1988: 3 ( Eucnecosum ; notes; Germany).

— Siitonen, 1993: 230 ( Eucnecosum ; Finland).

— Boháč, 1993: 41 ( Eucnecosum ; Slovakia; Czech Republic).

— Zanetti, 1995: 8 ( Eucnecosum ; Italy).

— Downie and Arnett, 1996: 431 ( Eucnecosum ; characters; USA).

— Cibuļskis, 1998: 67 ( Eucnecosum ; Latvia).

— Ryabukhin, 1999: 25 ( Eucnecosum ; catalog; habitat; distribution; northeastern Russia).

— Cibuļskis, 1999: 36 ( Eucnecosum ; list; Latvia).

- DISTRIBUTION: Faroe Islands, Europe, Russia, Mongolia, Canada, USA. impressum Zetterstedt, 1828: 52 ( Omalium ; Type locality: Lapponia Tornensis: alpi Tornensi; Kattilakoski; Juhonpietis; Wittangi).

— Zetterstedt, 1838: 51 ( Omalium ; characters; habitat; Lapland).

— Erichson, 1840: 859 ( Arpedium ; synonym of brachypterum ).

— Kraatz, 1857d: 959 ( Arpedium ; synonym of brachypterum ).

— Thomson, 1861: 190 ( Arpedium ; synonym of brachypterum ).

— Mulsant and Rey, 1880: 151 ( Arpedium ; synonym of brachypterum ).

— Ganglbauer, 1895: 718 ( Arpedium ; synonym of brachypterum ).

— Bernhauer and Schubert, 1910: 68 ( Arpedium ; synonym of brachypterum ).

— Munster, 1933: 259 ( Arpedium ; synonym of brachypterum ).

— Campbell, 1984: 491 ( Eucnecosum ; synonym of brachypterum ).

gyllenhalii C. Sahlberg, 1830: 286 ( Omalium ; Type locality: Lapponia: Yläne).

— Mannerheim, 1830: 54 ( Anthobium ; Finland).

— Mannerheim, 1831: 468 ( Anthobium ; Finland).

— Erichson, 1840: 859 ( Arpedium ; synonym of brachypterum ).

— Redtenbacher, 1849: 828 ( Arpedium ; synonym of brachypterum ).

— Kraatz, 1857d: 959 ( Arpedium ; synonym of brachypterum ).

— Thomson, 1861: 190 ( Arpedium ; synonym of brachypterum ).

— Seidlitz, 1875: 237 ( Arpedium ; characters; Baltic region).

— J. Sahlberg, 1876: 210 ( Arpedium ; synonym of brachypterum ).

— Ganglbauer, 1895: 718 ( Arpedium ; synonym of brachypterum ).

— Poppius, 1909: 6 ( Arpedium ; Lena Valley, Siberia ).

— Reitter, 1909: 187 ( Arpedium ; subgenus Eucnecosum ; characters; Germany).

— Bernhauer and Schubert, 1910: 68 ( Arpedium ; synonym of brachypterum ).

— Munster, 1933: 259 ( Arpedium ; variety of brachypterum ).

— Campbell, 1984: 491 ( Eucnecosum ; synonym of brachypterum ).

troglodytes Kiesenwetter, 1847: 77 ( Arpedium ; Type locality: Riesengebirge).

— Redtenbacher, 1849: 828 ( Arpedium ; characters).

— Redtenbacher, 1857: 246 ( Arpedium ; characters; Germany).

— Kraatz, 1857d: 959 ( Arpedium ; characters; Germany).

— Fauvel, 1871a: 113 [= 1872: 87] ( Arpedium ; characters; France).

— Redtenbacher, 1874: 271 ( Arpedium ; characters).

— Seidlitz, 1875: 237 ( Arpedium ; characters; Baltic region).

— Mulsant and Rey, 1880: 152 ( Arpedium ; characters; France).

— Ganglbauer, 1895: 718 ( Arpedium ; characters; Alps; Scotland; Caucasus).

— Bernhauer and Schubert, 1910: 68 ( Arpedium ; synonym of brachypterum ).

— Portevin, 1929: 441 ( Arpedium ; probably a race of brachypterum ; characters; France).

— Munster, 1933: 259 ( Arpedium ; synonym of brachypterum ).

— Scheerpeltz, 1933: 1060 ( Arpedium ; synonym of brachypterum ).

— Lohse, 1964: 56 (Eucnecsoum; synonym of brachypterum ).

— Campbell, 1984: 491 ( Eucnecosum ; synonym of brachypterum ).

— Zanetti, 1987: 312 ( Eucnecosum ; synonym of brachypterum ).

engadinicum Heyden, 1863: 21 ( Arpedium ; Type locality: Bernina).

— Bernhauer and Schubert, 1910: 68 ( Arpedium ; synonym of brachypterum ).

— Campbell, 1984: 491 ( Eucnecosum ; synonym of brachypterum ). mixtum Bernhauer, 1902d: 702 ( Arpedium ; Type locality: Tartola, Samarovo, Fl. Konda, Fl. Irtisch, Leusch).

— Poppius, 1909: 6 ( Arpedium ; synonym of tenue ).

— Bernhauer and Schubert, 1910: 69 ( Arpedium ; synonym of tenue ).

— Campbell, 1984: 491 ( Eucnecosum ; synonym of brachypterum ).

beringense Van Dyke, 1921: 162 ( Arpedium ; Type locality: St. George Island, Pribilof Islands , Alaska) .

— Scheerpeltz, 1933: 1060 ( Arpedium ; catalog).

— Steel, 1964: 249 ( Arpedium ; synonym of beringense).

— Campbell, 1984: 491 ( Eucnecosum ; synonym of brachypterum ).

brunnescens J. Sahlberg, 1871 a: 423 ( Arpedium ; Type locality: Muonioniska).

— Zetterstedt, 1838: 52 ( Omalium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 68 cited gyllenhali Zetterstedt as a synonym of brunnescens J. Sahlberg ; Zetterstedt did not describe the species as new, he attributed it to Sahlberg]; characters; habitat; Finland).

— J. Sahlberg, 1876: 210 ( Arpedium ; characters; Finland).

— Fauvel, 1878d: 227 [= 1878e: 63] ( Arpedium ; variety of tenue ; characters; Lake Superior; Lapland).

— Bernhauer and Schubert, 1910: 68 ( Arpedium ; catalog).

— Fall, 1926: 145 ( Arpedium ; Alaska).

— Munster, 1933: 264 ( Arpedium ; characters; Norway).

— Scheerpeltz, 1933: 1060 ( Arpedium ; catalog).

— Palm, 1948: 113 ( Arpedium ; characters; Sweden; Norway; Finland).

— Steel, 1964a: 249 ( Arpedium ; Canada; Alaska).

— Campbell, 1984: 501 ( Eucnecosum ; characters; Alberta; British Columbia ; Manitoba; Northwest Territories; Labrador; Ontario; Quebec; Yukon Territory; Alaska; Colorado; Michigan; New York).

— Elias, 1991: 556 ( Eucnecosum ; Quaternary fossil collection; Colorado).

— Siitonen, 1993: 230 ( Eucnecosum ; Finland).

— Downie and Arnett, 1996: 432 ( Eucnecosum ; characters; Canada; USA).

— Lavoie, Elias, and Payette, 1997: 231 ( Eucnecosum ; Holocene fossil; subarctic Quebec).

— Ryabukhin, 1999: 27 ( Eucnecosum ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Norway, Finland, Canada, USA.

hoppingi Hatch, 1957: 49 , 57 ( Revelstokea ; Type locality: Revelstoke Mt., B. C. [ British Columbia ]).

— Steel, 1964: 249 ( Arpedium ; synonym of brunnescens ).

— Campbell, 1984: 501 ( Eucnecosum ; synonym of brunnescens ).

engadinicum Heyden, 1863 , see: brachypterum Gravenhorst, 1802 .

gyllenhalii C. Sahlberg, 1830 , see: brachypterum Gravenhorst, 1802 .

hoppingi Hatch, 1957 , see: brunnescens J. Sahlberg, 1871 .

impressum Zetterstedt, 1828 , see: brachypterum Gravenhorst, 1802 .

kittilense Munster, 1933 , see: tenue LeConte, 1863 .

meybohmi Lohse, 1963 , see: tenue LeConte, 1863 .

mixtum Bernhauer, 1902 , see: brachypterum Gravenhorst, 1802 .

norvegicum Munster, 1933 , see: tenue LeConte, 1863 .

puncticolle J. Sahlberg, 1880: 106 ( Arpedium ; Type locality: Spirina; Tschemaschevo; Tobolsk; Acinovo; Kurejka; Turuchansk).

— Poppius, 1909: 6 ( Arpedium ; Lena Valley, Siberia ).

— Bernhauer and Schubert, 1910: 68 ( Arpedium ; catalog).

— Munster, 1933: 265 ( Arpedium ; characters; Russia).

— Palm, 1948: 111 ( Arpedium ; Siberia; Kola Peninsula).

— Tikhomirova, 1973a: 140 ( Eucnecosum ; checklist; northeastern Europe; Siberia).

— Silfverberg, 1992: 22 ( Eucnecosum ; Finland; Russia). - DISTRIBUTION: Finland, Russia.

salaasi Munster, 1933, see: tenue LeConte, 1863 .

tenue LeConte, 1863 a: 55 ( Acidota ; Type locality: north shore of Lake Superior). — Fauvel, 1878d: 226 [= 1878e: 62] ( Arpedium ; characters; Michigan; Lake Superior; British Columbia ; Lapland; Siberia). — Ganglbauer, 1895: 718 ( Arpedium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 69 cited brachypterum Ganglbauer as a synonym of tenue ; Ganglbauer did not describe the species as new, he attributed it to Gravenhorst]; characters; northern Europe; Switzerland; North America ). — Poppius, 1909: 6 ( Arpedium ; Lena Valley, Siberia ). — Bernhauer and Schubert, 1910: 69 ( Arpedium ; catalog). — Bernhauer, 1926g: 2 ( Arpedium ; Kamchatka region). — Scheerpeltz, 1933: 1061 ( Arpedium ; catalog). — Hatch, 1957: 54 ( Arpedium ; characters; British Columbia ). — Campbell, 1984: 496 ( Eucnecosum ; characters; Alberta; British Columbia ; Manitoba; Northwest Territories; Labrador; Ontario; Quebec; Saskatchewan; Yukon Territory; Alaska; Colorado; Michigan; Utah). — Zanetti, 1987: 314 ( Eucnecosum ; characters; Italy). — Elias, 1991: 556 ( Eucnecosum ; Quaternary fossil collection; Colorado). — Siitonen, 1993: 230 ( Eucnecosum ; Finland). — Zanetti, 1995: 8 ( Eucnecosum ; Italy). — Downie and Arnett, 1996: 431 ( Eucnecosum ; characters; Canada; USA). — Ryabukhin, 1999: 26 ( Eucnecosum ; catalog; habitat; distribution; northeastern Russia). - DISTRIBUTION: Norway, Sweden, Finland, Russia, Austria, Italy, Canada, USA.

norvegicum Munster, 1933: 261 ( Arpedium ; Type locality: Norway [Note: No specific locality cited; on page 264 a paragraph gave a general distribution]). — Palm, 1948: 112 ( Arpedium ; characters; Sweden; Norway; Finland). — Lohse, 1964: 57 ( Eucnecosum ; characters; central Europe). — Steel, 1964: 249 ( Arpedium ; synonym of tenue ). — Campbell, 1984: 496 ( Eucnecosum ; synonym of tenue ). — Lucht, 1987: 89 ( Eucnecosum ; checklist; Austria). — Zanetti, 1987: 314 ( Eucnecosum ; synonym of tenue ).

kittilense Munster, 1933: 261 ( Arpedium ; variety of norvegicum ; Type locality: Norway). — Palm, 1948: 113 ( Arpedium ; variety of norvegicum ; characters; Sweden). — Steel, 1964: 249 ( Arpedium ; synonym of tenue ). — Campbell, 1984: 496 ( Eucnecosum ; synonym of tenue ).

saalasi Munster, 1933: 261 , 263 ( Arpedium ; variety of norvegicum ; [Note: Cited as saalasi on page 261 and as salaasi on page 263; the correct spelling is saalasi since the species was named for Uunio Saalas]; Type locality: Kittilä in Finnish Lapland). — Campbell, 1984: 496 ( Eucnecosum ; synonym of tenue ).

meybohmi Lohse, 1963: 171 ( Eucnecosum ; Type locality: Rätische Alpen, Heidelberger Hütte in etwa 2000 m). — Lohse, 1964: 57 ( Eucnecosum ; synonym of norvegicum ). — Campbell, 1984: 496 ( Eucnecosum ; synonym of tenue ). — Zanetti, 1987: 314 ( Eucnecosum ; synonym of tenue ).

troglodytes Kiesenwetter, 1847 , see: brachypterum Gravenhorst, 1802 .

Geodromicus

[123 species; Nearctic, Palaearctic , and Oriental Regions]

Geodromicus Redtenbacher, 1857: 244 (replacement name for Geodromus Heer ; [Note: Redtenbacher cited it again in 1874 as replacement name]). TYPE SPECIES: Staphy-

— Jacquelin du Val, 1857: 70 (characters).

— Kraatz, 1857d: 925 (group of Anthophagus ).

— G. Waterhouse, 1858: 31 (catalog of British species).

— Thomson, 1859: 48 (characters; type species: plagiatus ).

— Schaum, 1859: 31 (subgroup of Anthophagus ; catalog; European species).

— Thomson, 1861: 180 (characters).

— Gredler, 1863: 121 (list of species from Tirol).

— Gemminger and Harold, 1868: 657 (synonym of Anthophagus ).

— Fauvel, 1871a: 132 [= 1872: 106] (characters; key to species).

— Sharp, 1871d: 14 (list of British species).

— Redtenbacher, 1874: 266 (characters).

— Fauvel, 1875a:VII [= 1875b: 209] (catalog).

— Fauvel, 1878d: 252 [= 1878e: 88] (key to North American species).

— Mulsant and Rey, 1880: 53 (characters; key to French species).

— Heyden, 1880: 81 (list of species of Siberian region).

— Fowler, 1888: 399 (notes; key to British species).

— Casey, 1894: 406 (notes; key to North American species).

— Seidlitz, 1891: 347 (key to some European species).

— Seidlitz, 1891a: 370 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 118 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 710 (characters; key to middle European species).

— Luze, 1903: 103 (characters; key to species).

— Heyden, Reitter, and Weise, 1906: 135 (list of species of Europe and Caucasus).

— Reitter, 1909: 183 (characters; key to German species).

— Bernhauer and Schubert, 1910: 75 (world catalog; 26 species).

— Blatchley, 1910: 475 (characters).

— Petri, 1912: 55 (list of species of Siebenbürgen).

— Johansen, 1914: 587 (characters).

— Leng, 1920: 95 (catalog of North American species).

— Winkler, 1925: 335 (catalog for Palaearctic region).

— Cameron, 1925: 12, 104 (catalog of Indian species).

— Porta, 1926: 30 (characters; key to species of Italy).

— Portevin, 1929: 436 (characters; key to French species).

— Cameron, 1930: 160, 455 (characters; key to species of British India).

— Roubal, 1930: 316 (catalog; Slovakia).

— Joy, 1932: 98 (key to British species).

— Scheerpeltz, 1933: 1065 (world catalog supplement).

— Kloet and Hincks, 1945: 161 (list of British species).

— Palm, 1948: 121 (characters; species of Sweden).

— Tottenham, 1949: 358, 408 (type species: plagiatus Fabricius ; list of British species). — Blackwelder, 1952: 169 (type species).

— Tottenham, 1954: 35 (characters; key to British species).

— Hatch, 1957: 51, 69 (characters; key to species of Pacific Northwest).

— Fagel, 1958b (notes; key to european species).

— Smetana, 1959c (notes; key to species of Czechoslovakia).

— Palm, 1961: 155 (key to Swedish species).

— Horion, 1963: 133 (list of central European species).

— Lohse, 1964: 61 (characters; key to central European species).

— Moore, 1966: 48, 50 (characters; notes).

— Scheerpeltz, 1968a: 17 (catalog of Austrian species).

— Steel, 1970a: 21 (larval characters; biological notes).

— Tikhomirova, 1973a: 140 (checklist of species of USSR).

— Fagel, 1976: 382 (notes on type species; key to species of Turkey).

— Pope, 1977: 22 (list of British species).

— Topp, 1978: 309 (larval characters).

— Bordoni, 1984a: 19 (characters; discussions; species of Palaearctic region).

— Zerche, 1987a (valid genus; senior generic name).

— Zanetti, 1987: 370 (synonym of Psephidonus ).

— Iablokov-Khnzorian, 1989: 137 (key to species of Caucasus).

— Lohse and Lucht, 1989: 127 (notes).

— Watanabe, 1990: 265 (synonym of Psephidonus ).

— Bordoni, 1990a: 100 (characters).

— Campbell and Davies, 1991: 91 (checklist of species; Alaska and Canada).

— Zerche, 1992a: 110, 139 (senior generic name, published September 24, 1856 not 1857; phylogenetic relationships).

— Silfverberg, 1992: 22 (synonym of Psephidonus ).

— Zanetti, 1995: 9 (list of species of Italy).

— M. Hansen, 1996: 95 (list of Danish species).

— Newton, Thayer, Ashe, and Chandler, 2000: 339 (12 Nearctic species; notes; characters in key).

— Herman, 2001: 57 (discussion; date of publication after that of Psephidonus ; evidence supports date of publication in 1857; request to Commission to place Geodromicus on Official List of Generic Names in Zoology).

Geobius Heer, 1839: 193 (species included: plagiatus , kunzei ; preoccupied by Dejean, 1831, and Brullé, 1832). TYPE SPECIES: Staphylinus plagiatus Fabricius , fixed by subsequent designation (see Blackwelder, 1952: 169).

— Heer, 1841: 572 (notes).

— Jacquelin du Val, 1857: 70 (synonym of Geodromicus ).

— Kraatz, 1857d: 925 (synonym of Geodromicus ).

— Thomson, 1858: 37 (characters).

— Schaum, 1859: 31 (synonym of Geodromicus ).

— Ganglbauer, 1895: 710 (synonym of Geodromicus ).

— Luze, 1903: 103 (synonym of Geodromicus ).

— Bernhauer and Schubert, 1910: 75 (synonym of Geodromicus ).

— Cameron, 1930: 160 (synonym of Geodromicus ).

— Zerche, 1987a (synonym of Geodromicus ).

— Watanabe, 1990: 265 (synonym of Psephidonus ).

— Zerche, 1992a: 110 (synonym of Geodromicus ).

Geodromus Heer, 1841: 572 (preoccupied by Dejean, 1829; replacement name for Geobius Heer ). TYPE SPECIES: Staphylinus plagiatus Fabricius , fixed by objective synonymy with Geobius .

— Redtenbacher, 1849: 747 (characters).

— Schaum, 1852: 30 (list of species; Europe).

— Fairmaire and Laboulbène, 1856: 629 (characters).

— Jacquelin du Val, 1857: 70 (synonym of Geodromicus ).

— Kraatz, 1857d: 925 (synonym of Geodromicus ).

— G. Waterhouse, 1858: 31 (synonym of Geodromicus ).

— Jacquelin du Val, 1868: 79 (catalog; European species).

— Gemminger and Harold, 1868: 657 (synonym of Anthophagus ).

— Ganglbauer, 1895: 710 (synonym of Geodromicus ).

— Luze, 1903: 103 (synonym of Geodromicus ).

— Bernhauer and Schubert, 1910: 75 (synonym of Geodromicus ).

— Cameron, 1930: 160 (synonym of Geodromicus ).

— Scheerpeltz, 1933: 1065 (synonym of Geodromicus ).

— Blackwelder, 1952: 169 (type species).

— Zerche, 1987a (synonym of Geodromicus ).

— Watanabe, 1990: 265 (synonym of Psephidonus ).

— Zerche, 1992a: 110 (synonym of Geodromicus ).

Psephidonus Gistel, 1856: 29 (species included: kunzei ). TYPE SPECIES: Geobius kunzei Heer , fixed by monotypy. — Székessy, 1963: 22 (characters).

— Moore and Legner, 1975: 211 (catalog; North America north of Mexico).

— Shibata, 1976: 116 (checklist of species of Japan).

— Burakowski, Mroczkowski, and Stefańska, 1979: 68 (catalog; Poland).

— Moore and Legner, 1979: 194 (characters; notes).

— Muona, 1979: 18 (list of Scandinavian species).

— Tóth, 1982: 100 (characters; habitus; key to species of Hungary).

— Yuh, Paik, Kwon, and Lee, 1985: 225 (checklist of species of Korea).

— Muona and Viramo, 1986: 15 (list of species of northeastern Finland).

— Zerche, 1987a (synonym of Geodromicus ).

— Zanetti, 1987: 370 (characters; habitus; key to species of Italy).

— Watanabe, 1990: 265 (characters; key to species of Japan).

— Angelini, 1991: 193 (list of species from high elevations of Calabria, Italy).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Zerche, 1992a: 110 (synonym of Geodromicus ; date of publication December 31, 1856).

— Siitonen, 1993: 230 (list of Finnish species).

— Boháč, 1993: 41 (cited as valid name; list of species of Czech Republic and Slovakia).

— Downie and Arnett, 1996: 434 (characters; key to species of Northeastern North America ).

— Herman, 2001: 57, 83 (discussion; date of publication, February 18, 1856, preceded that of Geodromicus ; request will be made to Commission to place Psephidonus on Official List of Rejected Names).

Brachydromicus Bordoni, 1993: 157 (subgenus of Geodromicus ; species included: latiusculus , brevicollis , constrictus ). TYPE SPECIES: Geodromicus latiusculus Eppelsheim , fixed by original designation.

— Bordoni, 1984a: 23, 24, 54 (subgenus of Geodromicus ; [Note: Cited as a new subgenus but was an unavailable name because it lacked a type species designation (Article 13.3)]; characters).

— Zanetti, 1987: 370 (subgenus of Psephidonus ; characters).

— Zerche, 1992a: 107, 110 (nomen nudum; synonym of Geodromicus ).

abditus † Scudder, 1900: 77 ( Geodromicus ; fossil; Type locality: Florissant , Colorado) .

— Handlirsch, 1907: 730 ( Geodromicus ; catalog; Miocene; Colorado).

- DISTRIBUTION: USA.

affinis Cameron, 1934: 17 ( Geodromicus ; Type locality: Indian Tibet: Kargil ).

- DISTRIBUTION: India.

afghanicus Scheerpeltz, 1961b: 36 ( Geodromicus ; subgenus Geodromicus ; Type locality: Afganistan: Panjao).

— Scheerpeltz, 1960b: 67 ( Geodromicus ; subgenus Geodromicus ; Afghanistan).

— Scheerpeltz, 1963: 5 ( Geodromicus ; Afghanistan).

— Coiffait, 1982b: 83 ( Geodromicus ; note).

— Kashcheev, 1999c: 16 ( Geodromicus ; characters in key).

- DISTRIBUTION: Afghanistan.

alticola Cameron, 1945: 65 ( Geodromicus ; Type locality: Kashmir: Lianmarg, altitude

10,000 feet).

- DISTRIBUTION: India.

amamianus Watanabe, 1990: 284 ( Psephidonus ; Type locality: Japan: Kagoshima Pref.: Shinmura, Amami-Ohshima Is.).

- DISTRIBUTION: Japan.

amnicola Champion, 1920: 246 ( Geodromicus ; Type locality: Sunderhunga in Kumaon ).

— Cameron, 1924b: 173 ( Geodromicus ; characters in key; India).

— Cameron, 1930: 163 ( Geodromicus ; characters; India).

— Scheerpeltz, 1933: 1065 ( Geodromicus ; catalog).

- DISTRIBUTION: India.

anatolicus Fagel, 1976: 383 ( Geodromicus ; Type locality: S.O. Anatolien: Ost Taurus bei Sürgü).

— Bordoni, 1984a: 38 ( Geodromicus ; Turkey).

- DISTRIBUTION: Turkey. antennatus Reitter, 1890 , see: constricticollis Eppelsheim, 1890 . anthracinus Brisout, 1863 , see: plagiatus Fabricius, 1798 . aokii Watanabe, 1972: 111 ( Psephidonus ; Type locality: north cirque, alt. 1800 m, on

Mt. Poroshiri-dake in the Hidaka mountains, Hokkaido).

— Watanabe, 1990: 291 ( Psephidonus ; characters; Japan).

— J. Li, 1993: 19 ( Psephidonus ; China).

- DISTRIBUTION: Japan, China. armeniacus Iablokov-Khnzorian, 1989: 134 ( Geodromicus ; Type locality: ArmSSR:

o. Sevlich. 3000 m nad ur. m. byshe Gorisa, u gory Mets Ishkhan).

- DISTRIBUTION: Armenia. arvernus Sainte-Claire Deville, 1935 , see: kunzei Heer, 1839 . asiaticus Bernhauer, 1902 a: 249 ( Geodromicus ; Type locality: Kleinasien, Bulghar Maaden).

— Luze, 1903: 110 ( Geodromicus ; characters; Kleinasien).

— Bernhauer and Schubert, 1910: 75 ( Geodromicus ; catalog).

— Tikhomirova, 1973a: 140 ( Geodromicus ; checklist; Asia Minor).

— Bordoni, 1984a: 36 ( Geodromicus ; characters; Turkey).

- DISTRIBUTION: Turkey. baerti Bordoni, 1984 a: 52 ( Geodromicus ; subgenus Geodromicus ; Type locality: Francia centrale: Auvergne, Cantal, Mont Dore).

- DISTRIBUTION: France. beibienkoi Tikhomirova, 1973: 156 ( Geodromicus ; Type locality: Suputinskoii zapovednik;

Kangauze; okrestnostiakh Vladivostoka; Kedrovaia pad).

— Kashcheev, 1999c: 16 ( Geodromicus ; characters in key).

- DISTRIBUTION: Russia. bodemeyeri Bernhauer, 1902 a: 250 ( Geodromicus ; Type locality: Kleinasien, Bulghar

Maaden).

— Luze, 1903: 113 ( Geodromicus ; characters; Kleinsasien).

— Bernhauer and Schubert, 1910: 75 ( Geodromicus ; catalog).

— Tikhomirova, 1973a: 140 ( Geodromicus ; checklist; Asia Minor).

— Bordoni, 1984a: 42 ( Geodromicus ; characters; Turkey).

- DISTRIBUTION: Turkey. brancuccii Coiffait, 1984: 142 ( Geodromicus ; Type locality: Pakistan: Sharan, 2400–2700 m).

- DISTRIBUTION: Pakistan. brevicollis Fauvel, 1875 a:vii [= 1875a: 209], note 7 ( Geodromicus ; proposed for Hochhuth’s misidentification of kunzei ; Type locality: Caucase).

— Hochhuth, 1849: 202 ( Anthophagus ; [Note: Misidentification: Bernhauer and Schubert, 1910:

76 cited kunzei Hochhuth as a synonym of globulicollis ; Hochhuth did not describe the species as new, he attributed it to Heer. However Fauvel, 1875a:vii proposed a name for Hochhuth’s misidentification]; characters; Caucasus; Akhaltzik).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of globulicollis ).

— Bordoni, 1984a: 55 ( Geodromicus ; valid species; characters; Caucasus).

— Iablokov-Khnzorian, 1989: 138 ( Geodromicus ; characters).

— Bordoni, 1993: 157 ( Geodromicus ; subgenus Brachydromicus ).

- DISTRIBUTION: Caucasus. brevitarsis Champion, 1925: 104 ( Geodromicus ; Type locality: Shelshel, N. Kumaon,

alt. 15,750 ft.).

— Cameron, 1930: 162 ( Geodromicus ; characters; India).

— Scheerpeltz, 1933: 1065 ( Geodromicus ; catalog).

brunneus Say, 1823: 158 ( Anthophagus ; Type locality: on the banks of the Missouri above the confluence of the Platte river ).

— Erichson, 1840: 854 ( Anthophagus ; characters; Missouri).

— Provancher, 1877: 259 ( Anthophagus ; characters; Quebec).

— Fauvel, 1878d: 254 [= 1878e: 90] ( Geodromicus ; characters; Missouri).

— Casey, 1894: 407 ( Geodromicus ; notes).

— Bernhauer and Schubert, 1910: 75 ( Geodromicus ; catalog).

— Blatchley, 1910: 475 ( Geodromicus ; characters; Indiana).

— Notman, 1919a: 95 ( Geodromicus ; Ontario).

— Notman, 1920a: 180 ( Geodromicus ; New York).

— Downie and Arnett, 1996: 435 ( Psephidonus ; characters; USA).

- DISTRIBUTION: Canada, USA.

caesus Erichson, 1840: 853 ( Anthophagus ; Type locality: America septentrionalis ).

— Fauvel, 1878d: 252 [= 1878e: 88] ( Geodromicus ; characters; Pennsylvania; Illinois; Missouri; Maryland; West Virginia).

— Casey, 1894: 407 ( Geodromicus ; synonym of brunneus ).

— Bernhauer and Schubert, 1910: 75 ( Geodromicus ; synonym of brunneus ).

caesus Erichson, 1840 , see: brunneus Say, 1823 .

caliginosus Sharp, 1889: 470 ( Anthophagus ; Type locality: Hiogo).

— Bernhauer and Schubert, 1910: 79 ( Anthophagus ; catalog).

— Scheerpeltz, 1933: 1065 ( Geodromicus ; catalog).

— Nakane, 1963a: 83 ( Psephidonus ; characters; habitus photograph; Japan).

— Yuh, Paik, Kwon, and Lee, 1985: 225 ( Psephidonus ; Korea).

— Watanabe, 1990: 288 ( Psephidonus ; characters; Japan).

- DISTRIBUTION: Japan.

lewisi Cameron, 1930c: 183 ( Geodromicus ; Type locality: Kobe: Shinowara).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog).

— Watanabe, 1990: 289 ( Psephidonus ; synonym of caliginosus ).

canaliculatus Stephens, 1834 , see: globulicollis Mannerheim, 1830 .

caucasianus Bordoni, 1984 a: 34 ( Geodromicus ; subgenus Geodromicus ; Type locality: Caucaso occ.: Krasnaja Poljana).

- DISTRIBUTION: Russia.

championi Cameron, 1924b: 171 , 174 ( Geodromicus ; Type locality: Chakrata District; Sainj Khud and Binal Gad, 6500 to 8000 feet).

— Cameron, 1930: 163 ( Geodromicus ; characters; India).

— Scheerpeltz, 1933: 1065 ( Geodromicus ; catalog).

- DISTRIBUTION: India.

chinensis Bernhauer, 1938: 19 ( Geodromicus ; Type locality: China: Provinz Fukien , Foochow).

- DISTRIBUTION: China.

cimicoides Cameron, 1941: 144 ( Geodromicus ; Type locality: Kashmir: Gulmarg).

- DISTRIBUTION: India.

concolor Luze, 1903 [Note: Nomen nudum], see: suturalis Lacordaire, 1835 .

consors Cameron, 1945: 65 ( Geodromicus ; Type locality: Kashmir: Gulmarg).

- DISTRIBUTION: India.

constricticollis Eppelsheim, 1890: 172 ( Geodromicus ; Type locality: westliches Kaukasusgebiet am Fischt ).

— Luze, 1903: 116 ( Geodromicus ; characters; Caucasus).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

— Khnzorian, 1962: 110 ( Geodromicus ; subspecies of globulicollis ; [Note: This change of status seems not to have been followed by others]; characters).

— Bordoni, 1984a: 57 ( Geodromicus ; characters; Caucasus).

— Bordoni, 1993: 157 ( Geodromicus ; subgenus Brachydromicus ).

- DISTRIBUTION: Russia.

antennatus Reitter, 1890: 189 ( Geodromicus ; Type locality: Cirkassien: Montes Aibgenses). — Luze, 1903: 116 ( Geodromicus ; synonym of constricticollis ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of constricticollis ).

convexicollis Luze, 1903: 112 ( Geodromicus ; Type locality: Russisches Zentralasien: Turkestan: Taschkent).

— Luze, 1904: 78 ( Geodromicus ; Turkestan).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

— Coiffait, 1970d: 144 ( Geodromicus ; Uzbekistan).

— Tikhomirova, 1973a: 141 ( Geodromicus ; checklist; Central Asia).

— Zerche, 1992a: 111 ( Geodromicus ; characters; Uzbekistan; Tadzhikistan).

- DISTRIBUTION: Uzbekistan, Tajikistan.

hauserianus Bordoni, 1984 a: 42 ( Geodromicus ; subgenus Geodromicus ; Type locality:

Ost. Buchara: Tschiftschantan, Nufswald).

— Zerche, 1992a: 107, 111 ( Geodromicus ; synonym of convexicollis ).

convexus Khnzorian, 1962: 106 ( Geodromicus ; Type locality: ArmSSR: Sevan).

— Tikhomirova, 1973a: 141 ( Geodromicus ; checklist; Armenia).

- DISTRIBUTION: Armenia.

cordicollis Petri, 1891 , see: puncticollis Weise, 1875 .

coriaceus Cameron, 1924 , see: Hygrodromicus .

corsicus Fagel, 1958b: 223 ( Geodromicus ; Type locality: Corse: forêt de Vizzavona, 850 m). — Bordoni, 1984a: 27 ( Geodromicus ; notes).

— Zanetti, 1987: 380 ( Psephidonus ; subgenus Psephidonus ; characters; Italy uncertain).

— Zanetti, 1995: 9 ( Geodromicus ; Sardinia?).

- DISTRIBUTION: Corsica, Sardinia?.

crassipalpis Champion, 1920: 245 ( Geodromicus ; Type locality: Sunderhunga in Kumaon ,

alt. 12,000 ft.).

— Cameron, 1924b: 173 ( Geodromicus ; characters in key; India).

— Cameron, 1930: 164 ( Geodromicus ; characters; India).

— Scheerpeltz, 1933: 1065 ( Geodromicus ; catalog).

- DISTRIBUTION: India.

curtipennis Fauvel, 1871 a: 136 [= 1872: 110] ( Geodromicus ; variety of plagiatus ; Type locality: Savoie, Alpes de la Vanoise; vallée de la Reuve).

— Ganglbauer, 1895: 711 ( Geodromicus ; synonym of globulicollis ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; variety of globulicollis ).

— Portevin, 1929: 436 ( Geodromicus ; subspecies of globulicollis ; characters; France).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; variety of globulicollis ).

— Jarrige, 1968b: 76 ( Geodromicus ; subspecies of kunzei ; France).

— Bordoni, 1984a: 51 ( Geodromicus ; valid species; characters; France).

- DISTRIBUTION: France.

curvipes Cameron, 1924b: 173 , 174 ( Geodromicus ; Type locality: Simla Hills; Gahan,

7000 feet).

— Cameron, 1930: 164 ( Geodromicus ; characters; India).

— Scheerpeltz, 1933: 1065 ( Geodromicus ; catalog).

- DISTRIBUTION: India.

cylindricus Tronquet, 1981: 74 ( Geodromicus ; Type locality: Afghanistan: Band-I-Amir).

- DISTRIBUTION: Afghanistan.

danieli Smetana, 1959c: 361 ( Geodromicus ; Type locality: Slowakei, Vysoké Tatry , [Hohe Tatra]: Koprová dolina).

— Tóth, 1982: 104 ( Psephidonus ; characters).

— Bordoni, 1984a: 51 ( Geodromicus ; characters; Slovakia).

— Lucht, 1987: 90 ( Geodromicus ; subspecies of kunzei ; checklist; Poland; Czechoslovakia).

— Lohse and Lucht, 1989: 127 ( Geodromicus ; valid species).

— Boháč, 1993: 41 ( Psephidonus ; Slovakia).

- DISTRIBUTION: Poland, Slovakia.

darahensis Coiffait, 1979d: 565 ( Geodromicus ; Type locality: province de Nangarhar, Darah i Nour , 3000 m).

- DISTRIBUTION: Afghanistan.

debilis Casey, 1894: 409 ( Geodromicus ; Type locality: Colorado).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

- DISTRIBUTION: USA.

depressus Cameron, 1941: 143 ( Geodromicus ; Type locality: Kashmir: Gulmarg).

- DISTRIBUTION: India.

dolpoensis Coiffait, 1982: 155 ( Geodromicus ; Type locality: Nepal: Dolpo, Aufstieg zum Namu La, 4300–4450).

- DISTRIBUTION: Nepal.

dubiosus Fagel, 1976: 384 ( Geodromicus ; Type locality: S.O. Anatolien: Ost Taurus bei Sürgü).

— Bordoni, 1984a: 38 ( Geodromicus ; Turkey).

- DISTRIBUTION: Turkey.

fauveli Casey, 1894: 408 ( Geodromicus ; Type locality: Oregon: The Dalles ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

— Scheerpeltz, 1933: 1065 ( Geodromicus ; catalog).

— Hatch, 1957: 70 ( Geodromicus ; characters; British Columbia ; Idaho; Washington; Oregon).

- DISTRIBUTION: Canada, USA.

flavipes Motschulsky, 1860 , see: Trichodromeus .

formosanus Hayashi, 1992 a: 110 ( Psephidonus ; Type locality: Taiwan: Mt. Yushan , Chiai Hsien).

- DISTRIBUTION: Taiwan.

frigidus Smetana, 1968: 229 ( Geodromicus ; Type locality: Mongolei: Uburchangaj aimak, Changaj Gebirge , Ongijn gol, 10 km, ONO Arbajcher, 1800 m).

— Tikhomirova, 1973a: 141 ( Geodromicus ; checklist; Mongolia).

— Kashcheev, 1999c: 16 ( Geodromicus ; characters in key).

- DISTRIBUTION: Mongolia.

globulicollis Mannerheim, 1830 , see: plagiatus Fabricius, 1798 .

gracilicornis Luze, 1903: 111 ( Geodromicus ; Type locality: Kaukasus: Hochgebirge östlich von Kasbek).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

— Khnzorian, 1962: 108 ( Geodromicus ; characters).

— Tikhomirova, 1973a: 141 ( Geodromicus ; checklist; Caucasus).

— Bordoni, 1984a: 36 ( Geodromicus ; characters; Caucasus).

- DISTRIBUTION: Georgia.

haarlovi Scheerpeltz, 1961b: 39 ( Geodromicus ; subgenus Geodromicus ; Type locality: Afghanistan: Afghanistan: Surta).

— Kashcheev, 1999c: 15 ( Geodromicus ; characters in key).

- DISTRIBUTION: Afghanistan.

hammondi Watanabe, 1990: 269 ( Psephidonus ; Type locality: Japan: Honshu: Gunma Pref.: near Marunuma, Oku-Nikkô ).

- DISTRIBUTION: Japan.

hermani Watanabe, 1991 a: 43 ( Psephidonus ; replacement name for pusillus Watanabe ).

- DISTRIBUTION: Japan.

pusillus Watanabe, 1990: 279 ( Psephidonus ; [preoccupied]; Type locality: Japan: Honshu: Shizuoka Pref.: Mt. Amagi-san, Izu ).

hoejeri Palm, 1961: 157 ( Geodromicus ; Type locality: S. Åberg bei Hoburgen auf Gotland). — Muona, 1979: 18 ( Psephidonus ; checklist; Sweden).

— Bordoni, 1984a: 48 ( Geodromicus ; characters).

— Silfverberg, 1992: 22 ( Psephidonus ; checklist; Sweden).

- DISTRIBUTION: Sweden.

humboldtianus Casey, 1894: 411 ( Geodromicus ; Type locality: California: Humboldt Co.). — Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

- DISTRIBUTION: USA.

iburinus Watanabe, 1990: 278 ( Psephidonus ; Type locality: Japan: Hokkaido: Kita-Yuzawa, Iburi ).

- DISTRIBUTION: Japan.

integer Casey, 1894: 412 ( Geodromicus ; Type locality: Washington State).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

— Hatch, 1957: 70 ( Geodromicus ; characters; British Columbia ; Washington; Oregon; Idaho).

- DISTRIBUTION: Canada, USA.

iranicus Coiffait, 1981: 20 ( Geodromicus ; Type locality: Iran: Col. de Schalous , dans le Massif de l’Elbrouz, 2600–2900 m).

- DISTRIBUTION: Iran.

japonicus Watanabe, 1990: 280 ( Psephidonus ; Type locality: Japan: Honshu: Tochigi Pref.: near Lake Sainoko, Oku-Nikkô ).

- DISTRIBUTION: Japan.

jarrigeanus Fagel, 1976: 390 ( Geodromicus ; Type locality: Col d’Ovit , Rize, 2500 m).

— Bordoni, 1984a: 37 ( Geodromicus ; Turkey).

- DISTRIBUTION: Turkey.

ziganensis Coiffait, 1978d: 171 ( Geodromicus ; Type locality: Col de Zigana , 2000 m).

— Coiffait, 1978f: 278 ( Geodromicus ; variety of jarrigeanus ; characters; Turkey).

karakorus Coiffait, 1977: 269 ( Geodromicus ; Type locality: Vallée Nang Mah , Karakorum). - DISTRIBUTION: Nepal.

kashmirensis Cameron, 1930: 455 ( Geodromicus ; Type locality: Kashmir: Gulmarg, Ferozepur Nala, alt. 6500 feet).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog).

- DISTRIBUTION: India.

kirschenblati Tikhomirova, 1973 , see: sibiricus Bernhauer, 1915 .

klimai Bordoni, 1984 a: 32 ( Geodromicus ; subgenus Geodromicus ; Type locality: Caucaso). — Iablokov-Khnzorian, 1989: 137, 138 ( Geodromicus ; characters).

- DISTRIBUTION: Caucasus.

kunduzii Tronquet, 1981: 72 ( Geodromicus ; Type locality: Afghanistan: versant nord du col de Salang, 2500 m, Hindu-Kush).

- DISTRIBUTION: Afghanistan.

kunzei Heer, 1839: 193 ( Geobius ; Type locality: Glarneralpen; Engadin; Avers; Stelvio; Samnaun; auch auf der Gemmi beim Daubensee).

— Heer, 1841: 572 ( Geodromus ).

— Märkel and Kiesenwetter, 1848a: 328 ( Anthophagus ; notes; Carpathian Alps).

— Kraatz, 1857d: 927 ( Anthophagus ; synonym of globulicollis ).

— Kraatz, 1857d: 927 ( Anthophagus ; [Note: Misidentified as globulicollis according to Zanetti, 1987: 381]; characters; Germany).

— Mulsant and Rey, 1880: 59 ( Geodromicus ; [Note: Misidentified as globulicollis according to Zanetti, 1987: 381]; characters; France). — Ganglbauer, 1895: 711 ( Geodromicus ; synonym of globulicollis ). — Ganglbauer, 1895: 711 ( Geodromicus ; [Note: Misidentified as globulicollis according to Zanetti, 1987: 381]; characters; northern Europe; Pyrenees; Alps; Carpathians). — Luze, 1903: 113 ( Geodromicus ; [Note: Misidentified as globulicollis according to Zanetti, 1987: 381]; characters; Sweden; Norway; Lapland; Finland; England; Riesengebirge; Hungary; Tirol; Switzerland; Savoy; Caucasus; Armenia). — Reitter, 1909: 184 ( Geodromicus ; [Note: Misidentified as globulicollis according to Zanetti, 1987: 381]; characters; Germany). — Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of globulicollis ). — Portevin, 1929: 436 ( Geodromicus ; [Note: Misidentified as globulicollis according to Zanetti, 1987: 381]; characters; France). — Smetana, 1959c: 360 ( Geodromicus ; valid species; characters; notes; Czechoslovakia). — Horion, 1963: 136 ( Geodromicus ; France; Switzerland; Italy; Austria; Germany). — Lohse, 1964: 62 ( Geodromicus ; characters; central Europe). — Tóth, 1982: 104 ( Psephidonus ; cited as kunzeni; characters). — Bordoni, 1984a: 49 ( Geodromicus ; characters). — Zanetti, 1987: 381 ( Psephidonus ; subgenus Psephidonus ; characters; Italy). — Lucht, 1987: 90 ( Geodromicus ; distribution of nominate subspecies; Germany; Austria). — Bordoni, 1993: 157 ( Geodromicus ; notes). — Boháč, 1993: 41 ( Psephidonus ; Czech Republic). — Zanetti, 1995: 9 ( Geodromicus ; Italy). - DISTRIBUTION: Europe.

arvernus Sainte-Claire Deville, 1935: 84 ( Geodromicus ; variety of globulicollis ; Type locality: Mont-Dore). — Jarrige, 1946a: 111 ( Geodromicus ; subspecies of globulicollis ; France). — Jarrige, 1968b: 76 ( Geodromicus ; subspecies of kunzei ; France).

languidus Luze, 1903: 110 ( Geodromicus ; Type locality: Kaukasus: Hochgebirge östlich von Kasbek). — Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog). — Khnzorian, 1962: 108 ( Geodromicus ; characters). — Tikhomirova, 1973a: 140 ( Geodromicus ; checklist; Caucasus). — Bordoni, 1984a: 34 ( Geodromicus ; characters; Caucasus). - DISTRIBUTION: Georgia.

lapponicus Bordoni, 1984 a: 29 ( Geodromicus ; subgenus Geodromicus ; Type locality: Lapponia). - DISTRIBUTION: Finland.

laticollis Fagel, 1976: 385 ( Geodromicus ; Type locality: S.O. Anatolien: Ost Taurus bei Sürgü). — Bordoni, 1984a: 38 ( Geodromicus ; Turkey). - DISTRIBUTION: Turkey.

latiusculus Eppelsheim, 1878: 126 ( Geodromicus ; Type locality: Hochgebirge östlich vom Kasbek in den Gebieten der Chefsuren und Pschawen, 7000′; [Note: See lectotype designation by Bordoni, 1984a]). — Luze, 1903: 115 ( Geodromicus ; characters; Caucasus). — Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog). — Khnzorian, 1962: 110 ( Geodromicus ; characters). — Tikhomirova, 1973a: 141 ( Geodromicus ; checklist; Caucasus). — Bordoni, 1984a: 55 ( Geodromicus ; lectotype designation; characters; Caucasus). — Bordoni, 1993: 157 ( Geodromicus ; subgenus Brachydromicus ). - DISTRIBUTION: Georgia.

lebedevianus Roubal, 1929: 46 ( Geodromicus ; Type locality: Russisch-Asien: Talass-Tal).

— Tikhomirova, 1973a: 141 ( Geodromicus ; checklist; Central Asia).

- DISTRIBUTION: Russia.

ledouxi Tronquet, 1981: 73 ( Geodromicus ; Type locality: Afghanistan: vallée d’Ajar ).

- DISTRIBUTION: Afghanistan.

lestevoides Sharp, 1889: 470 ( Anthophagus ; Type locality: Hakodate).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

— Bernhauer and Schubert, 1910: 80 ( Anthophagus ; catalog).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog).

— Nakane, 1963a: 83 ( Psephidonus ; characters; habitus photograph; Japan).

— Watanabe, 1969: 624 ( Psephidonus ; characters; South Korea).

— Yuh, Paik, Kwon, and Lee, 1985: 226 ( Psephidonus ; Korea).

— Watanabe, 1990: 274 ( Psephidonus ; characters; Japan).

— J. Li, 1993: 19 ( Psephidonus , cited as Pssephidonus; China).

- DISTRIBUTION: Japan, Korea, China.

lestevoides Reitter, 1900 , see: Hygrodromicus .

lewisi Cameron, 1930 , see: caliginosus Sharp, 1889 .

libanicus Coiffait, 1985: 210 ( Geodromicus ; Type locality: Liban: Les Cèdres , 2300 m).

- DISTRIBUTION: Lebanon.

lituratus Kraatz, 1857d: 928 ( Anthophagus ; Type locality: Tyrol).

— Redtenbacher, 1874: 266 ( Geodromicus ; characters).

— Ganglbauer, 1895: 711 ( Geodromicus ; characters; Tyrolian and Swiss Alps).

— Luze, 1903: 114 ( Geodromicus ; cited as aberration of globulicollis ; characters).

— Montandon, 1908: 78 ( Geodromicus ; Romania).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of globulicollis ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog).

— Horion, 1935: 170 ( Geodromicus ; characters; Austria; Switzerland).

— Lohse, 1964: 62 ( Geodromicus ; variety of kunzei ).

— Bordoni, 1984a: 28 ( Geodromicus ; characters; Switzerland).

— Lohse and Lucht, 1989: 127 ( Geodromicus ; valid species).

- DISTRIBUTION: Caucasus, Romania, Austria, Switzerland.

marginatus Fauvel, 1871 a: 135 [= 1872: 109] ( Geodromicus ; race of plagiatus ; Type locality: Alpes Suisses; Caucase).

— Ganglbauer, 1895: 711 ( Geodromicus ; synonym of lituratus ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of lituratus ).

lohsei Jarrige, 1963 , see: nigrita P. W. J. Müller, 1821 .

longicornis Coiffait, 1983: 278 ( Geodromicus ; Type locality: Cachemire: Himachal Pradesh, Mari , 3500 m).

- DISTRIBUTION: India.

longipalpis Coiffait, 1983: 279 ( Geodromicus ; Type locality: Cachemire: Gulmarg, 3000– 3500 m).

- DISTRIBUTION: India.

longipes Mannerheim, 1830: 56 ( Lesteva ; Type locality: Suecia: Wettern; Ostrogothia: Holkaberg ).

— Mannerheim, 1831: 470 ( Staphylinus ; Sweden).

— Stephens, 1834: 360 ( Lesteva ; cited as globulicollis ; [Note: Misidentification: This citatation probably refers to longipes ; see Pope, 1977: 22]; characters; Scotland).

— Stephens, 1839: 431 ( Lesteva ; cited as globulicollis ; [Note: Misidentification: This citatation probably refers to longipes ; see Pope, 1977: 22]; characters; Scotland).

— Erichson, 1840: 853 ( Anthophagus ; synonym of plagiatus ).

— Hardy, 1851: 55 ( Geodromus ; cited as globulicollis ; [Note: Misidentification: This citatation — Kraatz, 1857d: 927 ( Anthophagus ; synonym of globulicollis ).

— Fauvel, 1869: 493 ( Anthophagus ; synonym of globulicollis ).

— Fauvel, 1871a: 135 [= 1872: 109] ( Geodromicus ; synonym of globulicollis ).

— Mulsant and Rey, 1880: 60 ( Geodromicus ; synonym of globulicollis ).

— Fowler, 1888: 400 ( Geodromicus ; cited as globulicollis ; [Note: Misidentification: This citatation probably refers to longipes ; see Pope, 1977: 22]; characters; Britain).

— Ganglbauer, 1895: 711 ( Geodromicus ; synonym of globulicollis ).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; synonym of globulicollis ).

— Palm, 1948: 122 ( Geodromicus ; characters; Sweden; Norway; Finland).

— Larsson and Gígja, 1959: 72 ( Geodromicus ; cited as valid species; biological notes; Iceland). — Palm, 1961: 156 ( Geodromicus ; characters; Sweden).

— Steel, 1970a: 22 ( Geodromicus ; biological notes).

— Pope, 1977: 22 ( Geodromicus ; Britain).

— Bordoni, 1984a: 46 ( Geodromicus ; characters).

- DISTRIBUTION: Iceland, Sweden, Britain.

luzianus Bordoni, 1984 a: 35 ( Geodromicus ; subgenus Geodromicus ; Type locality: Armenia: Schawnabad).

— Kashcheev, 1999c: 16 ( Geodromicus ; characters in key).

- DISTRIBUTION: Armenia.

macrothorax Kashcheev, 1999c: 15 ( Geodromicus ; Type locality: Kungei-Alatau, verkhnee techenie reki Chilik, urochishche Sarybastau).

- DISTRIBUTION: Kazakhstan.

maderi Bernhauer, 1943: 72 ( Geodromicus ; Type locality: China: Szechwan mer., Mts. Kinfushan, 2000 m, am Flusse Sung-Kanho).

- DISTRIBUTION: China.

major Motschulsky, 1860: 550 ( Geodromus ; Type locality: alpes du Caucase en Touchétie).

— Luze, 1903: 109 ( Geodromicus ; characters; Caucasus).

— Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog).

— Khnzorian, 1962: 108 ( Geodromicus ; characters).

— Fagel, 1963: 426 ( Geodromicus ; Turkey).

— Tikhomirova, 1973a: 140 ( Geodromicus ; checklist; Caucasus).

— Bordoni, 1984a: 32 ( Geodromicus ; characters; Caucasus).

— Iablokov-Khnzorian, 1989: 137, 138 ( Geodromicus ; characters).

— Kashcheev, 1999c: 16 ( Geodromicus ; characters in key).

- DISTRIBUTION: Caucasus, Turkey.

marginatus Fauvel, 1871 , see: lituratus Kraatz, 1857 .

meurguesae Tronquet, 1981 , see: parallelipennis Coiffait, 1979 .

miyamorii Watanabe, 1990: 287 ( Psephidonus ; Type locality: Japan: Hokkaido: Tonkeshi Riv., Iburi ).

- DISTRIBUTION: Japan.

montenegrinus Bernhauer, 1941: 210 ( Geodromicus ; Type locality: Montenegro: Ljuboten). — Zanetti, 1984: 78 ( Geodromicus ?; characters; notes; Bulgaria).

- DISTRIBUTION: Bulgaria, Yugoslavia.

nepalensis Scheerpeltz, 1976 a: 100 ( Geodromicus ; subgenus Geodromicus ; Type locality: Nepal: Khumbu Lobuche in etwa 4800 m).

- DISTRIBUTION: Nepal.

nepalicus Coiffait, 1977: 267 ( Geodromicus ; Type locality: Rive du tila Khola près Jumla, Népal occidental).

- DISTRIBUTION: Nepal.

nigrita P. W. J. Müller, 1821: 226 ( Anthophagus ; Type locality: Odenbach).

— Germar, 1827: 5 ( Anthophagus ; synonym of plagiatus ).

— Curtis, 1829: 29 ( Lesteva ; cited as plagiatus ; catalog; Britain; [Note: Misidentification: Probably refers to nigrita ; plagiatus does not occur in Britain; see Pope, 1977: 22]). — Stephens, 1834: 362 ( Lesteva ; cited as plagiatus ; characters; England; [Note: Misidentification: Probably refers to nigrita ; plagiatus does not occur in Britain; see Pope, 1977: 22]). — Comolli, 1837: 15 ( Anthophagus ; Italy). — Stephens, 1839: 431 ( Lesteva ; cited as plagiatus ; characters; England; Scotland; [Note: Misidentification: Probably refers to nigrita ; plagiatus does not occur in Britain; see Pope, 1977: 22]). — Heer, 1839: 193 ( Geobius ; variety of plagiatus ; characters; Switzerland). — Erichson, 1840: 853 ( Anthophagus ; synonym of plagiatus ). — Hardy, 1851: 55 ( Geodromus ; synonym of plagiatus ). — Hardy, 1851: 55 ( Geodromus ; cited as plagiatus ; notes; England; [Note: Misidentification: Probably refers to nigritaus; plagiatus does not occur in Britain; see Pope, 1977: 22]). — Fauvel, 1865a: 315 ( Anthophagus ; characters; habitat; France). — Gemminger and Harold, 1868: 658 ( Anthophagus ; synonym of plagiatus ). — Fauvel, 1869: 493 ( Anthophagus ; synonym of plagiatus ). — Fauvel, 1871a: 134 [= 1872: 108] ( Geodromicus ; characters; cited as variety β of plagiatus ; France; Lapland; Scandinavia; Britain; Germany; Switzerland; Austria; Italy; Corsica). — Mulsant and Rey, 1880: 57 ( Geodromicus ; synonym of plagiatus ). — Fowler, 1888: 400 ( Geodromicus ; characters; Britain). — Ganglbauer, 1895: 711 ( Geodromicus ; synonym of plagiatus ). — Luze, 1903: 108 ( Geodromicus ; aberration of plagiatus ). — Bernhauer and Schubert, 1910: 77 ( Geodromicus ; aberration of plagiatus ). — Notman, 1920: 17 ( Geodromicus ; New York). — Scheerpeltz, 1933: 1066 ( Geodromicus ; aberration of plagiatus ). — Fagel, 1958b: 219 ( Geodromicus ; characters; western Europe; Alpes; Spain). — Lohse, 1958: 52 ( Geodromicus ; valid species; notes). — Smetana, 1959c: 357 ( Geodromicus ; characters; notes; Czechoslovakia). — Palm, 1961: 156 ( Geodromicus ; characters; Sweden). — Horion, 1963: 134 ( Geodromicus ; Sweden; Germany; Switzerland; France; Czechoslovakia; Austria). — Lohse, 1964: 62 ( Geodromicus ; characters; central Europe). — Steel, 1970a: 22 ( Geodromicus ; biological notes). — Pope, 1977: 22 ( Geodromicus ; Britain). — Hammond, 1980: 136 ( Geodromicus ; notes; Ireland). — Tóth, 1982: 103 ( Psephidonus ; characters). — Bordoni, 1984a: 26 ( Geodromicus ; characters; France). — Zanetti, 1984: 78 ( Geodromicus ; Bulgaria). — Segers, 1986: 22 ( Geodromicus ; checklist of species; Belgium). — Zanetti, 1987: 379 ( Psephidonus ; subgenus Psephidonus ; characters; Italy). — Lucht, 1987: 90 ( Geodromicus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Sweden). — Boháč, 1993: 41 ( Psephidonus ; Slovakia; Czech Republic). — Zanetti, 1995: 9 ( Geodromicus ; Italy). - DISTRIBUTION: Europe, USA.

lohsei Jarrige, 1963: 166 ( Geodromicus ; subspecies of nigrita ; Type locality: Forêt de Marialle, mousses de la Llipodère, 1700 m).

nipponensis Watanabe, 1990: 275 ( Psephidonus ; Type locality: Japan: Honshu: Gunma Pref.: near Nidoage, Asama Plateau). — Watanabe, 1996a: 9 ( Psephidonus ; cited as niponensis ; cave collection; Japan). - DISTRIBUTION: Japan.

niveus Bernhauer, 1941: 210 ( Geodromicus ; Type locality: Siera [sic] Nevada: Picachode Plakeria). - DISTRIBUTION: Spain.

nubilatus Casey, 1894: 408 ( Geodromicus ; Type locality: New Mexico: Las Vegas ).

— Fall and Cockerell, 1907: 168 ( Geodromicus ; New Mexico).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

- DISTRIBUTION: USA.

ohkurai Hayashi, 1992 a: 107 ( Psephidonus ; Type locality: Taiwan: Mt. Houfuan , Nantou Hsien).

- DISTRIBUTION: Taiwan.

yushanensis Hayashi, 1992 a: 110 ( Psephidonus ; subspecies of ohkurai ; Type locality: Taiwan: Mt. Yushan, Chiai Hsien).

ovalis Luze, 1904 , see: Hygrodromicus .

ovipennis LeConte, 1878: 452 ( Geodromicus ; Type locality: Leavenworth Valley, above Georgetown, Colo., 9000 to 10,000 feet).

— Fauvel, 1878d: 253 [= 1878e: 89] ( Geodromicus ; characters; Colorado).

— Casey, 1894: 408 ( Geodromicus ; notes).

— Ganglbauer, 1895: 711 ( Geodromicus ; synonym of plagiatus ).

— Fall and Cockerell, 1907: 168 ( Geodromicus ; New Mexico).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; synonym of plagiatus ).

— Fall, 1926: 145 ( Geodromicus ; notes Yukon Territory).

— Moore and Legner, 1975: 211 ( Psephidonus ; cited as valid species; checklist; USA).

— Downie and Arnett, 1996: 435 ( Psephidonus ; characters; USA).

- DISTRIBUTION: USA, Canada.

pakistanus Coiffait, 1984: 143 ( Geodromicus ; Type locality: Pakistan: Murree-Abbottabad , 2200–2500 m).

- DISTRIBUTION: Pakistan.

palpalis Coiffait, 1977: 271 ( Geodromicus ; Type locality: Dampelek près Jumla, Népal occidental).

- DISTRIBUTION: Nepal.

parallelipennis Coiffait, 1979d: 564 ( Geodromicus ; Type locality: vallée du Parum, 2400–2800 m).

— Tronquet, 1981: 69 ( Geodromicus ; Afghanistan).

- DISTRIBUTION: Afghanistan.

meurguesae Tronquet, 1981: 75 ( Geodromicus ; subspecies of parallelipennis ; Type locality: Afghanistan: Kahr-Shah-Foladi).

paramerus Shavrin, 2000: 87 ( Geodromicus ; Type locality: Chitinskaia Obl., Krasno- Chikoiskii r-n, pos. Malaia Kudara ).

- DISTRIBUTION: Russia.

pedemontanus Fagel, 1958b: 221 ( Geodromicus ; Type locality: Alpes maritimes piémontaises: Limone, torrente Vermenagna, 1000–1100 m).

— Bordoni, 1984a: 26 ( Geodromicus ; characters; notes; France; Italy).

— Zanetti, 1987: 377 ( Psephidonus ; characters; Italy).

— Zanetti, 1995: 9 ( Geodromicus ; Italy).

- DISTRIBUTION: France, Italy.

peraffinis Cameron, 1945: 65 ( Geodromicus ; Type locality: Kashmir: Lianmarg, altitude 10,000 feet).

- DISTRIBUTION: India.

perplexus Cameron, 1941: 144 ( Geodromicus ; Type locality: Kashmir: Lobzang).

- DISTRIBUTION: India.

plagiatus Fabricius, 1798: 180 ( Staphylinus ; Type locality: Europa borealis).

— Paykull, 1800: 387 ( Staphylinus ; characters; Sweden).

— Fabricius, 1801: 597 ( Staphylinus ; characters).

— Gravenhorst, 1802: 123, 189 ( Anthophagus ; characters).

— Gravenhorst, 1806: 222 ( Anthophagus ; characters; Sweden).

— Gyllenhal, 1810: 195 ( Anthophagus ; characters; Sweden).

— Gyllenhal, 1827: 461 ( Anthophagus ).

— C. Sahlberg, 1827: 276 ( Anthophagus ; characters; Finland).

— Germar, 1827: 5 ( Anthophagus ; characters).

— Zetterstedt, 1828: 44 ( Anthophagus ; characters; Lapland).

— Mannerheim, 1830: 56 ( Lesteva ; Finland; Sweden).

— Mannerheim, 1831: 470 ( Lesteva ; Finland; Sweden).

— Runde, 1835: 24 ( Lesteva ; characters; Germany).

— Lacordaire, 1835: 482 ( Lesteva ; characters; France).

— Zetterstedt, 1838: 47 ( Anthophagus ; characters; Lapland; Sweden).

— Heer, 1839: 193 (characters).

— Erichson, 1840: 852 ( Anthophagus ; characters; Europe).

— Hochhuth, 1849: 202 ( Anthophagus ; characters; Iberia).

— Fairmaire and Laboulbène, 1856: 629 ( Geodromus ; characters; France).

— Redtenbacher, 1857: 244 ( Geodromicus ; characters; Austria).

— Kraatz, 1857d: 925 ( Anthophagus ; characters; Germany).

— Thomson, 1859: 48 (type species of Geodromicus ).

— Thomson, 1861: 181 ( Geodromicus ; characters; Scandinavia).

— Hochhuth, 1862: 102 ( Anthophagus ; Russia).

— Fauvel, 1871a: 133 [= 1872: 107] ( Geodromicus ; characters; France; Lapland; Scandinavia; Germany; Switzerland; Austria).

— Redtenbacher, 1874: 266 ( Geodromicus ; characters; Austria).

— Seidlitz, 1875: 241 ( Anthophagus ; characters; Baltic region).

— J. Sahlberg, 1876: 208 ( Geodromicus ; Finland).

— Mulsant and Rey, 1880: 56 ( Geodromicus ; characters; France).

— Ganglbauer, 1895: 711 ( Geodromicus ; characters; north and middle Europe; Siberia; North America ).

— Luze, 1903: 108 ( Geodromicus ; characters; north and central Europe; Siberia; Caucasus;

North America ).

— Sainte-Claire Deville, 1906: 72 ( Geodromicus ; Corsica).

— Sainte-Claire Deville, 1907: 50 ( Geodromicus ; characters; France).

— Reitter, 1909: 183 ( Geodromicus ; characters; Germany).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

— Johansen, 1914: 587 ( Geodromicus ; characters; Denmark).

— Portevin, 1929: 436 ( Geodromicus ; characters; France).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog).

— Palm, 1948: 121 ( Geodromicus ; characters; Sweden; Denmark; Norway; Finland).

— Fagel, 1958b: 217 ( Geodromicus ; characters; notes; Scandinavia; Alpes).

— Smetana, 1959c: 358 ( Geodromicus ; characters; notes; Czechoslovakia).

— Palm, 1961: 155 ( Geodromicus ; characters; Sweden).

— Khnzorian, 1962: 109 ( Geodromicus ; characters).

— Horion, 1963: 133 ( Geodromicus ; Caucasus; Siberia; Germany; Austria; Hungary; Norway; Sweden; Czech Republic; Slovakia; Italy; Britain; Belgium; France; Switzerland; Italy; Corsica; Sardinia; Bosnia Hercegovina; Romania; Denmark; Holland).

— Lohse, 1964: 62 ( Geodromicus ; characters; central Europe).

— Tóth, 1982: 101 ( Psephidonus ; characters).

— Frank, 1982: 8 ( Geodromicus ; list of parasites).

— Bordoni, 1984a: 24 ( Geodromicus ; characters; notes).

— Zanetti, 1987: 377 ( Psephidonus ; subgenus Psephidonus ; characters; Italy).

— Lucht, 1987: 90 ( Geodromicus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Denmark; Sweden).

— Boháč, 1993: 41 ( Psephidonus ; Slovakia; Czech Republic).

— Zanetti, 1995: 9 ( Geodromicus ; Italy).

— Ryabukhin, 1999: 29 ( Geodromicus ; catalog; habitat; distribution; northeastern Russia).

globulicollis Mannerheim, 1830: 56 ( Lesteva ; Type locality: Finlandia orientalis: Saima). — Mannerheim, 1831: 470 ( Lesteva ; Finland). — Zetterstedt, 1838: 49 ( Anthophagus ; characters; habitat; Lapland; Finland). — Erichson, 1840: 853 ( Anthophagus ; synonym of plagiatus ). — Erichson, 1840: 852, 853 ( Anthophagus ; variety a, e; [Note: Bernhauer and Schubert, 1910: 76 cited plagiatus variety a and e Erichson as synonyms of globulicollis but neither varietal name is available]; characters). — Thomson, 1861: 181 ( Geodromicus ; characters; Scandinavia). — Hochhuth, 1862: 102 ( Anthophagus ; Russia). — Fauvel, 1865a: 315 ( Anthophagus ; habitat; France). — Fauvel, 1865a: 316 ( Anthophagus ; [Note: Misidentification: Bernhauer and Schubert, 1910: 76 cited lituratus Fauvel as a synonym of globulicollis ; Fauvel attributed it to Kraatz]; habitat; France). — Fauvel, 1871a: 135 [= 1872: 109] ( Geodromicus ; variety of plagiatus ; [Note: Bernhauer and Schubert, 1910: 76 cited plagiatus variety Ε as a synonym of globulicollis ; there is no available name]; characters; France; Lapland; Scandinavia; Britain; Switzerland; Germany; Austria; Caucasus). — Redtenbacher, 1874: 266 ( Geodromicus ; characters). — J. Sahlberg, 1876: 208 ( Geodromicus ; Finland). — Bernhauer and Schubert, 1910: 76 ( Geodromicus ; catalog). — Scheerpeltz, 1933: 1065 ( Geodromicus ; catalog). — Smetana, 1959c: 359 ( Geodromicus ; characters; notes; Czechoslovakia). — Khnzorian, 1962: 110 ( Geodromicus ; characters). — Lohse, 1964: 62 ( Geodromicus ; synonym of plagiatus ). — Boháč, 1993: 41 ( Psephidonus ; cited as valid species; Czech Republic?).

verticalis Say, 1831: 49 [= 1834: 463] ( Anthophagus ; Type locality: Pennsylvania). — Erichson, 1840: 854 ( Anthophagus ; characters; Pennsylvania). — LeConte, 1850: 220 ( Anthophagus ; Lake Superior). — LeConte, 1866: 375 ( Anthophagus ; notes; Lake Superior; Pennsylvania). — Fauvel, 1878d: 254 [= 1878e: 90] ( Geodromicus ; synonym of nigrita ). — Ganglbauer, 1895: 711 ( Geodromicus ; synonym of plagiatus ). — Bernhauer and Schubert, 1910: 77 ( Geodromicus ; synonym of plagiatus ).

canaliculatus Stephens, 1834: 360 ( Lesteva ; Type locality: Scotland; Cumberland; near Edinburgh). — Stephens, 1839: 431 ( Lesteva ; characters; England, Scotland). — G. Waterhouse, 1858: 31 ( Geodromicus ; synonym of globulicollis ).

anthracinus Brisout, 1863: 40 ( Geodromicus ; Type locality: Mont-Louis). — Gemminger and Harold, 1868: 658 ( Anthophagus ; synonym of plagiatus ). — Fauvel, 1871a: 134 [= 1872: 108] ( Geodromicus ; synonym of plagiatus variety β). — Ganglbauer, 1895: 711 ( Geodromicus ; synonym of plagiatus ). — Bernhauer and Schubert, 1910: 77 ( Geodromicus ; synonym of plagiatus ).

subnigritus Lohse, 1958: 53 ( Geodromicus ; aberration of plagiatus ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; no locality cited). — Palm, 1961: 156 ( Geodromicus ; aberration of plagiatus ; characters; Sweden). — Zanetti, 1987: 378 ( Psephidonus ; synonym of plagiatus ).

ponticus Fagel, 1976: 388 ( Geodromicus ; Type locality: N. E. Anatolie: Bicik, Giresum, 1600 m). — Bordoni, 1984a: 37 ( Geodromicus ; characters; Turkey). - DISTRIBUTION: Turkey.

puncticollis Weise, 1875: 364 ( Anthophagus ; subgenus Geodromicus ; Type locality: Luhy). — Eppelsheim, 1887b: 432 ( Geodromicus ; synonym of globulicollis ). — Ganglbauer, 1895: 711 ( Geodromicus ; characters; Europe). — Luze, 1903: 111 ( Geodromicus ; characters; Hungary; Siebenbürgen; Balkans).

— Smetana, 1959c: 362 ( Geodromicus ; characters; notes; Ukrainian Carpathians).

— Tóth, 1982: 104 ( Psephidonus ; characters).

— Bordoni, 1984a: 54 ( Geodromicus ; characters; Ukraine).

— Zerche, 1988a: 362 (types listed).

- DISTRIBUTION: Ukraine, Romania.

cordicollis Petri, 1891: 18 ( Geodromicus ; Type locality: Schässburg).

— Ganglbauer, 1895: 711 ( Geodromicus ; synonym of puncticollis ).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; synonym of puncticollis ).

punctulatus Luze, 1904 , see: Paratrichodromeus .

pusillus Coiffait, 1983: 279 ( Geodromicus ; Type locality: Cachemire: Rohtang Pass, 4000 m). - DISTRIBUTION: India.

pusillus Watanabe, 1990 , see: hermani Watanabe, 1991 .

pyrenaeus Jarrige, 1963: 166 ( Geodromicus ; Type locality: Gave d’Holçarté, Larrau, 600 m, Basses Pyrénées).

— Bordoni, 1984a: 27 ( Geodromicus ; characters; France).

- DISTRIBUTION: France.

recticollis Cameron, 1924: 172 , 173 ( Geodromicus ; Type locality: Chakrata District; Sainj Khud, 6500 feet).

— Cameron, 1930: 163 ( Geodromicus ; characters; India).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog).

- DISTRIBUTION: India.

reitteri Scheerpeltz, 1933 , see: Hygrodromicus .

rivularis Khnzorian, 1962: 107 ( Geodromicus ; Type locality: ArmSSR: Megri, u berega Araksa).

— Tikhomirova, 1973a: 141 ( Geodromicus ; checklist; Armenia).

— Iablokov-Khnzorian, 1989: 137, 138 ( Geodromicus ; characters).

- DISTRIBUTION: Armenia.

robusticornis Bernhauer, 1941: 209 ( Geodromicus ; Type locality: Bulgarien: Rila Musala ).

- DISTRIBUTION: Bulgaria.

rousi Bordoni, 1984 a: 39 ( Geodromicus ; subgenus Geodromicus ; Type locality: Caucaso).

— Iablokov-Khnzorian, 1989: 137, 138 ( Geodromicus ; characters).

- DISTRIBUTION: Caucasus.

ruficornis Cameron, 1941: 143 ( Geodromicus ; Type locality: Kashmir: Gulmarg, Lianmarg, altitude 8000– 10,000 feet).

- DISTRIBUTION: India.

rufipennis Van Dyke, 1924: 16 ( Geodromicus ; Type locality: Brooks Falls, Naknek Lake, Alaska Peninsula ).

- DISTRIBUTION: USA.

salangensis Tronquet, 1981: 76 ( Geodromicus ; Type locality: Afghanistan: versant nord du col de Salang, 3000–3200 m).

- DISTRIBUTION: Afghanistan.

schoenmanni Bordoni, 1984 a: 31 ( Geodromicus ; subgenus Geodromicus ; Type locality: Caucaso).

— Iablokov-Khnzorian, 1989: 137, 138 ( Geodromicus ; characters).

- DISTRIBUTION: Caucasus.

schubertorum Scheerpeltz, 1937b: 197 ( Geodromicus ; cited as schuberti [Note: Incorrect original spelling; named for two people named Schubert]; Type locality: Bulgarien: Massiv des Jumruktschal, Zentral-Balkan, 1400–1800 m; Rosalito-polje, Schipka-Balkan, ca. 1900 m).

— Zanetti, 1984: 78 ( Geodromicus ; cited as schuberti ; characters; Bulgaria).

schultzi Hatch, 1957: 70 ( Geodromicus ; Type locality: Washington: Kittitas Co.: Swauk Cr. Also cited from Idaho and British Columbia ).

- DISTRIBUTION: Canada, USA.

sibiricus Bernhauer, 1915c: 69 ( Geodromicus ; Type locality: Ostsibirien: Chitaizki Sterena).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog).

— Tikhomirova, 1973a: 141 ( Geodromicus ; checklist; eastern Siberia).

— Gusarov, 1992a: 775 [= 1993a: 62] ( Geodromicus ; synonymic notes).

— Kashcheev, 1999c: 16 ( Geodromicus ; characters in key).

- DISTRIBUTION: Russia.

kirschenblati Tikhomirova, 1973: 155 ( Geodromicus ; Type locality: Suputinskii zapovednik).

— Coiffait, 1974a: 198 ( Geodromicus ; Siberia).

— Gusarov, 1992a: 775 [= 1993a: 63] ( Geodromicus ; synonym of sibiricus ).

similis Cameron, 1930: 455 ( Geodromicus ; Type locality: Kashmir: Gulmarg, Ferozepur Nala, alt. 6500 feet).

— Cameron, 1934: 18 ( Geodromicus ; Indian Tibet).

— Cameron, 1935b: 288 ( Geodromicus ; India).

— Coiffait, 1982: 43 ( Geodromicus ; India).

- DISTRIBUTION: India.

sinuatocollis Bernhauer, 1938b: 50 ( Geodromicus ; Type locality: Mandschurei: Tcheuly-Nord: Nanye-li).

- DISTRIBUTION: China.

sinuatus Sharp, 1889: 470 ( Anthophagus ; Type locality: Sapporo).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; catalog).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog).

— Watanabe, 1990: 282 ( Psephidonus ; characters; Japan).

— J. Li, 1993: 19 ( Psephidonus ; China).

- DISTRIBUTION: Japan, China.

sprius Watanabe, 1990: 272 ( Psephidonus ; Type locality: Japan: Honshu: Kanagawa Pref.: Kojiri, Hakone ).

- DISTRIBUTION: Japan.

stiricidii † Scudder, 1895: 43 ( Geodromicus ; fossil, interglacial clays; Type locality: Scarboro , Ontario) .

- DISTRIBUTION: Canada.

striatus Khnzorian, 1962: 105 ( Geodromicus ; Type locality: ArmSSR: Gekhard, Kotaikskii r-n).

— Tikhomirova, 1973a: 141 ( Geodromicus ; checklist; Armenia).

- DISTRIBUTION: Armenia.

strictus Fauvel, 1889 a: 126 ( Geodromicus ; named for species misidentified as nigrita by Fauvel; Type locality: New York; Mont Washington).

— Fauvel, 1878d: 254 [= 1878e: 90] ( Olophrum ; [Note: Misidentification: Bernhauer and Schubert, 1910: 77 cited nigrita Fauvel as a synonym of strictus ; Fauvel did not describe the species as new, he attributed it to Müller]; characters; Michigan; Lake Superior; New York; New Hampshire; Pennsylvania).

— Casey, 1894: 408 ( Geodromicus ; New York and Massachusetts to Michigan).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

— Blatchley, 1910: 475 ( Geodromicus ; cited as stictus ; characters; Indiana).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog).

— Dearborn and Donahue, 1994: 14 ( Geodromicus ; Maine).

— Downie and Arnett, 1996: 435 ( Psephidonus ; characters; USA).

- DISTRIBUTION: USA.

suavis Coiffait, 1979d: 566 ( Geodromicus ; Type locality: Nouristan: haute vallée de l’Agôr

Khwar, 3200–3900 m).