Oides flava (Olivier)

Lee, Chi-Feng & Beenen, Ron, 2017, Revision of the Palaearctic and Oriental species of the genus Oides Weber, 1801 (Coleoptera: Chrysomelidae: Galerucinae), Zootaxa 4346 (1) : 54-57

Oides flava (Olivier)

( Figs 34D– 34I , 36 )

Adorium flavum Olivier, 1807: 611 (Bengale) .

Oides flava: Allard, 1891: 230 ( Laos: Luang Prabang); Weise, 1924: 3 (as synonym of O. concolor (Fabricius) , misidentification); Maulik, 1936: 118 ( India: United Provinces, Western Almora, Assam, Shillong); Gressitt & Kimoto, 1963: 478 ( China: Hainan; Vietnam); Kimoto & Takizawa, 1972: 217 ( Nepal); Wilcox, 1971: 9 (catalogue); Basu & Halder, 1987: 226 ( India: Orissa); Kimoto, 1989: 37 ( Laos); Kimoto, 1990: 219 (catalogue); Silfverberg, 1990: 201 ( Nepal); Samoderzhenkov, 1992: 108 ( Vietnam); Mohamedsaid, 1995: 5 ( Malaysia: Sabah); Basu, 1996: 679 ( India: West Bengal); Mohamedsaid, 1996: 3 ( Indonesia: Java); Mohamedsaid, 1997a: 169: ( Malaysia: Sarawak); Mohamedsaid, 1997b: 205 ( Malaysia: Sabah); Reid, 1998: 287 ( Indonesia: Java); Medvedev & Sprecher-Uebersax, 1999: 315 ( Nepal); Mohamedsaid, 1999: 16 ( Malaysia: Sarawak); Mohamedsaid & Holloway, 1999: 167 (Borneo); Mohamedsaid, 2000b: 353 (catalogue); Mohamedsaid, 2001: 165 ( Indonesia: Bali); Yang & Yao, 2002: 443 ( China: Hainan); Mohamedsaid, 2004: 65 (catalogue); Kimoto, 2005: 66 (catalogue); Takizawa & Kimoto, 2005: 103 ( Malaysia: Perak); Beenen, 2010: 491 (catalogue); Sprecher-Uebersax, 2011: 442 (catalogue); Yang et al., 2015: 96 (catalogue).

Adorium sumatrensis Blanchard, 1853: 337 ( Indonesia: Sumatra). New synonymy

Oides sumatrensis: Harold, 1876: 3556 (catalogue); Weise, 1924: 6 (catalogue); Laboissière, 1932: 147 (Indonesis Sumatra); Wilcox, 1971: 16 (catalogue); Kimoto, 1990: 221 (catalogue).

Adorium philippinensis Boheman, 1859: 175 ( Philippines: Mindanao). New synonymy

Oides philippinensis: Harold, 1876: 3556 (catalogue); Weise, 1924: 5 (catalogue); Wilcox, 1971: 13 (catalogue); Kimoto, 1990: 221 (catalogue).

Oides inornata Baly, 1879: 444 ( India: Assam); Weise, 1924: 4 (catalogue); Maulik, 1936: 118 (as synonym of O. flava ). Synonymy confirmed

Oides albicans Duvivier, 1884b: 237 ( Indonesia: Sumatra); Allard, 1891: 230 ( Laos: Luang Prabang); Weise, 1924: 2 (catalogue); Maulik, 1936: 118 (as synonym of O. flava ). Synonymy confirmed

Oides flavipes Duvivier, 1884b: 239 ( Indonesia: Timor); Weise, 1924: 4 (catalogue); Wilcox, 1971: 9 (catalogue); Kimoto, 1990: 219 (catalogue); Takizawa & Kimoto, 2005: 106 ( Indonesia: Timor); Mohamedsaid, 2009: 92 ( Indonesia: Lombok). New synonymy

Oides abdominalis Duvivier, 1884c: 133 ( Indonesia: Java); Weise, 1924: 1 (catalogue); Wilcox, 1971: 2 (catalogue); Kimoto, 1990: 218 (catalogue). New synonymy

Oides subtilissima Duvivier, 1884c: 136 ( Philippines: Luzon); Weise, 1924: 6 (catalogue); Wilcox, 1971: 16 (catalogue); Kimoto, 1990: 221 (catalogue). New synonymy

Oides flavida Duvivier, 1884c: 137 ( Philippines: Luzon); Weise, 1913 ( Philippines, Luzon); Weise, 1924: 4 (catalogue); Wilcox, 1971: 9 (catalogue); Kimoto, 1990: 219 (catalogue). New synonymy

Oides elliptica Duvivier, 1884c: 137 ( Philippines: Luzon); Weise, 1924: 3 (catalogue); Wilcox, 1971: 8 (catalogue); Kimoto, 1990: 218 (catalogue). New synonymy

Oides marcida Duvivier, 1884c: 138 ( Philippines: Luzon); Weise, 1924: 4 (catalogue); Wilcox, 1971: 12 (catalogue); Kimoto, 1990: 220 (catalogue). New synonymy

Oides rhabdoscelis Weise, 1922: 57 (East Malaysia: Sabah: Banguay island); Weise, 1924: 6 (catalogue); Wilcox, 1971: 15 (catalogue); Kimoto, 1990: 221 (catalogue); Mohamedsaid & Holloway, 1999: 168 (Borneo); Mohamedsaid, 2004: 66 (catalogue). New synonymy

Oides bakeri Laboissière, 1925: 609 ( Philippines: Mindanao); Wilcox, 1971: 4 (catalogue); Kimoto, 1990: 218 (catalogue). New synonymy

Oides ovalis Laboissière, 1925: 610 ( Philippines) ; Wilcox, 1971: 13 (catalogue); Kimoto, 1990: 220 (catalogue). New synonymy

Oides coomani Laboissière, 1927: 42 ( Vietnam) ; Wilcox, 1971: 7 (catalogue); Kimoto, 1989: 37 (as synonym of O. flava , synonymy confirmed).

Types. Adorium flavum . Holotype ♀ ( MNHN, by monotypy), labeled: “ Macé / Bengale [p, w, circle label] // A. flavum / Oliv. [h, w] // HOLOTYPE [p, r] // MUSÉUM PARIS [p] / Collection / GÉNERALE [h, w] // design. 1981 / A. VACHON—det. [p, w]”.

Adorium sumatrensis . Holotype ♀ (MNHN, by monotypy), labeled: “493 / 41 [h, w] // MUSEUM PARIS / SUMATRA / RADJA BASSA / JACQUINOT 1841 [p, w] // ♀ // HOLOTYPE [p, r] // VIDIT 0 2.1981 [h] / A. VACHON det [p, w] // Musuem Paris [p] / Sumatrensis [h, w]”.

Adorium philippinensis . Holotype ♂ (left elytron lost, NHRS, by monotypy), labeled: “male aedeagus and two broken antennae glued on the card) // Manilla [p, w] // Kinb. [p, w] // Type. [p, w] // Ad. Philippinensis / Bhn., [h, w] // vidit III-1975. [h] / A. VACHON det [p, w] // 89 [p] / 75 [h, r] // Typus [p, r] // NHRS-JLKB / 0 0 0 0 27150 [p, w]”.

Oides inornata . Lectotype ♀ (BMNH), here designated, labeled: “Kamrup [h, g] // Oides / inornata / Baly / Assam [h, b] // Type [p, w, circle label with red border] // SYN- / TYPE [p, w, circle label with blue border] // Baly Coll. [p, w] // Oides flava Olivier 1807 / = albicans Duvivier 1884 / = inornata Baly, 1877 [h] / A. VACHON—det. [p] / 2-1974 [p, w]”. Paralectotype. 1♂ (BMNH): “[male aedeagus glued on card] // Sibsagar [h, g] // Baly Coll. [p, w] // SYN- / TYPE [p, w, circle label with blue border] // Oides flava Olivier 1807 / = inornata Baly 1877 / ♂ dissected on February [h] / A. VACHON—det. [p] 1974 [h, w]”.

Oides albicans . Lectotype ♂ (RMNH), here designated, labeled: “[male aedeagus glued on card] // Oides / albicans / type. Duviv. [h, w] // Dr. B. Hagen. / Tandjong Morawa. / Serdang / (N. O. Sumatra). [p, w] // HOLOTYPE [p, r] // A. VACHON—det. [p] IV.1977 [h, w] // Oides [p, r] // albicans 2 syntypes [p, w] // RMNH Leiden / ex Indo-Austr. / collection [p, w]”. Paralectotype. 1♀ (RMNH): “ Oides / albicans / type. Duviv. [h, w] // Dr. B. Hagen. / Tandjong Morawa. / Serdang / (N. O. Sumatra). [p, w] // ♀ [p, w] // ALLOTYPE [p, r] // A. VACHON—det. [p] IV.1977 [h, w] // RMNH Leiden / ex Indo-Austr. / collection [p, w]”.

Oides flavipes . Holotype ♀ (RMNH, by monotypy), labeled: “Huller / Timor [h, w, circle label] // Oides / flavipes , / type. Duviv. [h, w] // ♀ [p, w] // HOLOTYPE [p, r] // A. VACHON—det. [p] IV.1977 [h, w] // flavipes holotype [p, w] // RMNH Leiden / ex Indo-Austr. / collection [p, w]”.

Oides abdominalis . Holotype ♀ (IRSB, by monotypy), labeled: “ Oides / abdominalis / Java Duviv. [h. w] // Collect. Duvivier [p, w] [both cards glued on the following card] // Coll. R. I. Sc. N.B. / Java [p, y] // TYPE [p, pink label] // V. Laboissière [p] vid, [h] 1939 [p] / Oides / abdominalis Duv / Type [h, w] // cf. Ann. Soc. Ent. Belg. / C. R. p. CXXXIII, 28, 1884 [h, w] // A. VACHON—[p] vidit [h] v-1976 [p, w]”.

Oides subtilissima . Lectotype ♂ (IRSB), here designated, labeled: “(male aedeagus glued on card) // ♂ [p, w] / / Coll. R. I. Sc. N. B. / Philippines [p, y] // V. Laboissière det., 1939: / Oides [p] / subtilissima Duviv. [h, w] // Oides subtilissima Duv. / aedéotype [h] / A. VACHON—det. [p] V.1976 [h, w] // LECTOTYPE [p, r]”. Paralectotypes. 1♀ (IRSB): “ ♀ [p, w] // Leyte / Semper [p, w] // Coll. Chapuis [p, w] [both labels glued on the following label] // Coll. R. I. Sc. N. B. / Philippines [p, y] // Adorium / subtilissimum / Leyte Chp. [h, w] [glued on the following label] // A. Duvivier det. [p] 1884 [h, w] // Allotype [p, orange label] // cf. Ann. Soc. Ent. Belg. / C. R. p. CXXXVI, 28, 1884 [h, w] // V. Laboissière vid, 1939: [p] / Oides / subtilissima / Type Duv [h, w] // A. VACHON—det. [p] / V. 1976 [h, w]”; 1♀ (IRSB): “ ♀ [p, w] // Coll. R. I. Sc. N. B. / Philippines [p, y] // V. Laboissière vid, 1939: [p] / Oides / subtilissima / Duv [h, w] // A. VACHON –[p] vidit / V. 1976 [h, w]”.

Oides flavida . Holotype ♂ (IRSB, by monotypy), labeled: “[male aedeagus glued on card] // ♂ [p, w] // Samar / Semper [p, w] // Coll. Chapuis [p, w] [both labels glued on the following label] // Coll. R. I. Sc. N. B. / Philippines [p, y] // Adorium / flavidum / Samar Chp. [h, w] [glued on the following label] // det. [p] Chapuis [h, w] // MUSÉUM PARIS / COLL H CLAVAREAU / 1932 [p, w] // ♂ holotype / Oides / flavida Duvivier 1884 [h] / A. VACHON—det. [p] / V. 1976 [h, w] // HOLOTYPE [p, r]”.

Oides elliptica . Lectotype ♂ (IRSB), here designated, labeled: “[male aedeagus glued on card] // ♂ [p, w] // Luzon / Semper [p, w] // Coll. Chapuis [p, w] [both labels glued on the following label] // Coll. R. I. Sc. N. B. / Philippines [p, y] // Adorium / ellipticum / Luzon Chp. [h, w] [glued on the following label] // det. [p] Chapuis [h, w] // Oides / elliptica [h, w] [glued on the following label] // A. Duvivier det. [p] 1884 [h, w] // MUSÉUM PARIS / COLL H CLAVAREAU / 1932 [p, w] // O. elliptica Duvivier 1884 / édéage prepare V-1973 [h] / A. VACHONdet. [p, w] // cf. Ann. Soc. Ent. Belg. / C. R. p. CXXXVII, 28, 1884 [h, w] // Holotype [p, r]”. Paratype: 1♀ (IRSB), labeled: “ ♀ [p, w] // Luzon / Semper [p, w] // Coll. Chapuis [p, w] [both labels glued on the following label] // Coll. R. I. Sc. N. B. / Philippines [p, y] // det. [p] Duvivier / Oides / elliptica Duv. [h, w] // Ex-Typis [p, red letters] / Duvivier 1884 [h, w] // cf. Ann. Soc. Ent. Belg. / C. R. p. CXXXVII, 28, 1884 [h, w] // V. Laboissiére [p] vid [h] 1939: [p] / Oides / elliptica Duv / Type [h, w] // TYPE [p] sec. Laboiss. 1939 [h, r] // A. VACHON—vidit V-1976 [p. w] // Allotype [p] / Vachon 1975 [h, r]”; 1♂ (MNHN): “[male aedeagus glued on the card] // ♂ [p, w] // Collect. / Duvivier [p, w] [this label glued on the following label] // Coll. R. I. Sc. N. B. / Philippines [p] Luzon / Semper [h, y] // det [p] Duvivier / Oides / elliptica Duv. [h, w] // Para- / type [p] sec. Laboiss. / 1939. [h, o] // V. Laboissiére det., 1939: [p] / Oides / elliptica Duv. / of typis [h, w]”.

Oides marcida . Lectotype ♀ (IRSB), here designated, labeled: “ ♀ [p, w] // Semper [p, w] // Coll. Chapuis [p, w] [both labels glued on the following label] // Coll. R. I. Sc. N. B. / Philippines [p, y] // Adorium / marcidum / Chp. [h, w] (glued on the following label) // det. [p] Chapuis [h, w] // TYPE [p, w] // MUSÉUM PARIS / COLL H CLAVAREAU / 1932 [p, w] // cf. Ann. Soc. Ent. Belg. / C. R. p. CXXXVIII, 28, 1 884 [h, w] // A. VACHON—[p] vidit V-1976 [h w]”. Paralectotypes. 1♀ (IRSB): “ Philippines / Semper [p, w] // Coll. Chapuis [p, w] [both labels glued on the following label] // Coll. R. I. Sc. N. B. / Philippine [p, y] // Oides (Adorium) / marcidum / Philip. Chp.

[h, w] [glued on the following label] // A. Duvivier det. [p] 1884 [h, w] // Ex-Typis [p, red letters] / Duvivier, 1884 [h, w] // cf. Ann. Soc. Ent. Belg. / C. R. p. CXXXVIII, 28, 1884 [h, w] // V. Laboissière vid, 1939: [p] / Oides / marcida Duv / Type [h, w] // TYPE [p] / sec. Laboiss. 1939 [h, pink label] // A. VACHON—[p] vidit V-1976 [h w]”; 1♀ (IRSB): “ ♀ [p, w] // Semper [p, w] // Coll. Chapuis [p, w] [both labels glued on the following label] // Coll. R. I. Sc. N. B. / Philippines [p, y] // det. [p] Duvivier 1884 / Oides / marcida Duviv. [h, w] // Ex-Typis [p, red letters] / Duvivier, 1884 [h, w] // cf. Ann. Soc. Ent. Belg. / C. R. p. CXXXVIII, 28, 1884 [h, w] // V. Laboissière det., 1939: [p] / Oides / marcida Duv. / ex. typis [h, w] // Para- / type [p] sec. Laboiss. / 1939. [h, o] // A. VACHON—[p] vidit V-1976 [h w]”; 1♀ (IRSB): “ ♀ [p, w] // Collect. / Duvivier [p, w] [glued on the following label] // det. [p] Duvivier 1884 / Oides / marcida Duviv. [h, w] // Ex-Typis [p, red letters] / Duvivier, 1884 [h, w] // cf. Ann. Soc. Ent. Belg. / C. R. p. CXXXVIII, 28, 1 884 [h, w] // V. Laboissière det., 1939: / Oides [p] / marcida Duviv. [h, w] // Para- / type [p] sec. Laboiss. / 1939. [h, o] // A. VACHON—[p] vidit V-1976 [h w]”; 1♀ (MNHN): “Collect. / Duvivier [p, w] [glued on the following label] // Coll. R. I. Sc. N. B. / Philippines [p, y] // det. [p] Duvivier 1884 / Oides / marcida Duviv. [h, w] // Para- / type [p] sec. Laboiss. / 1939. [h, o] // V. Laboissière det., 1939: / OIDES [p] / marcida Duviv. [h, w]”.

Oides rhabodscelis . Lectotype (NHRS, sex undetermined), here designated, labeled: “ Banguey / Ins. Nördl. Borneo / W. Kedenburg [p, w] // rhabodscelis / m. [h, w, blue letters] // Type [p, w] // Typus [p, r] // NHRS-JLKB / 000025263”. Number of paralectotypes are uncertain.

Oides bakeri . Lectotype ♂ (ZMUH), here designated, labeled: “ Davao / Mindanao / Baker [p, w] // Oides / Bakeri m. [h] / V. Laboissière—Dét. [p, w] // TYPE [p, w, red letters] // Le Moult vend. / Via Reinbek / Eing. Nr. 1, 1957 [p, w]”. Paralectotype. 1 ex. (USNM, sex undetermined): “ Davao / Mindanao / Baker [p, w] // 8564 [h, w] // Oides / Bakeri m. [h] / V. Laboissière—Dét. [p, w] // TYPE [p, w, red letters] // BLNO / 0 0 2521 [p, b] // USNMENT / 0 0 871281 [p, w]”.

Oides ovalis . Lectotype ♀ (ZMUH), here designated, labeled: “Mt. Makiling / Luzon, Baker [p, w] // Oides / ovalis m / V. Laboissière—Dét [p, w] // TYPE [p, w, red letters] // Le Moult vend. / Via Reinbek / Eing. Nr. 1, 1957 [p, w] // 2427 [h, w]”. Paralecotype. 1 ex. (USNM, sex undetermined): “Mt. Makiling / Luzon, Baker [p, w] // Oides / ovalis m / V. Laboissière—Dét [p, w] // TYPE [p, w, red letters] // BLNO / 0 0 2676 [p, b] // USNMENT / 0 0 871525 [p, w]”. There should be one more type based on the original descriptions ( Laboissière 1925).

Oides coomani . Lectotype ♀ (ZMUH), here designated, labeled: “ Hoa Binh / Tonkin / R P De Cooman [h, w] / / Oides / Coomani m [h] / V. Laboissière—Dét [p, w] // TYPE [p, w, red letters] // Le Moult vend. / via Reinbek / Eing. Nr. 1, 1957 [p, w]”. Number of paralectotypes is uncertain.

Redescription. Length 8.4–11.0 mm, width 5.6–7.0 mm. General color ( Figs 34D–34F ) yellow; antennae yellow, antennomeres VII–XI black. Antennae filiform in males ( Fig. 36A ), antennomeres IV longest, VIII–X elongate, length ratios of antennomeres I–XI 1.0: 0.7: 1.3: 1.4: 1.3: 1.2: 1.2: 0.9: 0.8: 0.8: 1.1, length to width ratios of antennomeres I–XI 2.4: 2.2: 3.9: 3.9: 3.6: 3.3: 2.9: 2.6: 2.4: 2.4: 3.5; similar in females ( Fig. 36B ), length ratios of antennomeres I–XI 1.0: 0.7: 1.1: 1.4: 1.2: 1.0: 1.0: 0.9: 0.8: 0.8: 1.0, length to width ratios of antennomeres I–XI 2.3: 2.0: 3.2: 4.1: 3.3: 2.9: 2.8: 2.5: 2.3: 2.2: 3.1. Pronotum transverse, 2.0x wider than long, disc slightly convex, with dense, fine punctures, punctures reduced medially, baso- and apico-lateral angles rectangular, apical margin slightly concave. Elytra oblong, widest at middle, 1.3–1.4x longer than wide, disc without reticulate microsculpture, but with dense punctures; moderately convex, epipleurae near lateral margins, located 9/10 distance between suture and lateral margins. Penis ( Figs 36C–36E ) slender; 6.8x longer than wide, parallel-sided, apically tapering from apical 1/5, bifurcate at apical 1/10, lateral apices connected; tectum extremely elongate, apex a little longer equal to apex of penis, deeply bifurcate at middle, lateral processes elongate or apically broadened; slightly curved near base in lateral view, apices narrowly rounded; inner apical half of ventral surface membranous; endophallic sac without sclerites. Apical margin of abdominal ventrite V in female truncate. Gonocoxae reduced. Ventrite VIII ( Fig. 36L ) transverse, apical margin rounded, depressed medially, with long setae along apical margin, spiculum long and wide. Receptacle of spermatheca ( Fig. 36M ) as wide as pump, connected to pump, basally narrowed; pump strongly curved; proximal spermathecal duct extremely elongate, apical area covered by short teeth.

Variation. Adults are variable in color patterns and penis shapes. Some specimens from Java possess black abdomens and were described as O. abdominalis . One female from Timor (holotype of O. flavipes ) possesses black thoracic and abdominal ventrites, but males from the same area are yellowish brown. Specimens from Borneo and Banguay island possess darker antennomeres IV–XI, and those from Banguay island possess black outer margins of tibiae and were described as O. rhabodscelis . A number of specimens collected from Mindanao possess black heads, metasterna and abdomens, and black spots on the pronota ( Figs 34G– 34I ). Males from Borneo and Philippines possess swollen apico-lateral apices of the penis ( Figs 36F– 36I ) and some are more elongate than usual (9.4x longer than wide) ( Figs 36H– 36I ). The apico-lateral apices of the penis are angular or subapically angular (described as O. elliptica ) ( Figs 36J–36K ) in some males from the Philippines.

Diagnosis. Members of this species are similar to those of O. foveicollis Laboissière and O. femoralis Laboissière with the entirely yellow elytra but can be distinguished by the yellow scutellum and femora (black scutellum and femora in O. femoralis ) and relatively longer antennomere III (much shorter in O. foveicollis ).

Other specimens examined. INDIA. Goa: 8 exs. ( JBCB), 30 km S of Margao (= Madgaon), Palolem env., 15°00’47”N 74°01’58”E, 0–20m, 12–14.VIII.2002, leg. P. Šípek & M. Fikáček; Himachal Pradesch: 1 ex. ( JBCB), Dharamsala, 12.IX.1994, leg. P. Hanáček; Karnataka: 1 ex. ( RBCN), 2 exs. ( RSCW), Jog Falls (Shimoga), 26.IX.1991, leg. Schuh; Maharashtra: 5 exs. ( JBCB), 4 km S of Lonavala, Bhushi dam env., 500m, 24–28.IX.2005, leg. J. Bezděk; 1 ex. ( JBCB), same but with “leg. F. & L. Kantner”; 1 ex. ( JBCB), 70 km S of Pune, Wai env., 3– 6.X.2005, leg. F. & L. Kantner; 4 exs. ( JBCB), same locality, 2–7.X.2005, leg. J. Bezděk; 1 ex. ( JBCB), Pune, VIII.1984, leg. I. Pokorný; INDONESIA. Bali: 1♀ ( TCHU), 20.I.1998, leg. Detani; 1♂ ( TCHU), same but with “ 1.II.1998 ”; 1♂ ( RMNH), Jembrana, 1919, leg. Brescher; 2♂♂, 1♀ ( TCHU), Yeh Sumbul Negan, 5–8.XII.1999, leg. H. Takizawa; 1♀ ( TCHU), same but with “ 24.IX.1998 ”; Java: 1♀ ( RMNH), Banjoewangi, 1911, leg. M. Gillavry; 1 ex. ( RMMH), Djampang Tengah, Priangan, 1600–2400m, 22–26.V.1939, leg. J.M.A. van Groenendael; 1 ex. ( RMNH), Djampang Tengah, Tjimerang, 1800m, 28.IV.1939, leg. J.M.A. van Groenendael; 1 ex. ( RMNH), same but with “ 24.IV.1939 ”; 1 ex. ( RMNH), Djampang Tengah, Soekaboemi, 1800–2200m, 8.V.1939, leg. J.M.A. van Groenendael; 2 exs. ( RMNH), Malang, leg. A. Haller; 1♂ ( RMNH), Noesa Kambangan, I.1914, leg. Drescher; 1♀ ( RMNH), same but with “ VI.1916 ”; 2♀♀ ( RMNH), Preanger, 1899, leg. Regentschappen; 1♀ ( RMNH), Toegoe, leg. J. D. Pasteur; 3 exs. ( NHMB), Trawas E slope, Gn Penaggungan, 1000m, 6–9.V.2001, leg. Bolm; Kalimantan: 1♀ ( MNHUB), Mahakam; Sumatra: 1♂ ( MCSN), Liangagas, leg. Dohrn; 1♂ ( MNHUB), Semarang, leg. Drescher; 1♂, 2♀♀ (1♂, 1♀: MCSN; 1♀: MNHUB), Soekaranda, I.1894, leg. Dohrn; 1♂ ( RMNH), Tandjong, Karang, Ketaton, leg. J. v. d. Vecht; 1 ex. ( RMNH), Tandjong, Morawa, Serdang, leg. B. Hagen; 1♂ ( TCHU), Watu Ulo, Jember, IV.2005, leg. Detani; Timor: 1♂ ( RMNH), Muller; LAOS. Khammouan: 1 ex. ( JBCB), Ban Khoun Ngeun, 18°07’N 104°29’E, 200m, 19–31.V.2001, leg. C. L. Peša; EAST MALAYSIA. Sabah: 7 exs. (6: FREY; 1: MNHUB), Banguey island; 1♂ ( MNHUB), Kinabalu; 1♂ ( JBCB), Lembah Danum, 27–31.X.1991, leg. Salleh et al.; Sarawak: 1 ex. Kuala Bok, 1.III.1969, leg. T. Kunou & Y. Arita; MYANMAR. 1♀ ( MCSN), Carin Cheba, 1889, leg. L. Fea; 1♀ ( MCSN), same but with “ V.–XII.1888 ”; PHILIPPINES. Luzon: 1♂, 2♀♀ ( NMNS), Aurora Prov., Dipaculao near Brgy Calaocan, 8km NW of Dinadiawan, 20.VII.2012, leg. M. L. Jeng; 3♂♂, 1♀ ( NMNS), Cagayan Prov., Sta. Ana Nangaramoan, 22. IV.2012, leg. M. L. Jeng & H. Cahilo; 2♀♀ ( MNHUB), Mt. Banahao, 20–23.IV.1914, leg. Boettcher; 1♀ ( MNHUB), same but with “ 26.IV.1914 ”; 1♀ ( MNHUB), same but with “ 20.VI.1914 ”; 1♂, 2♀♀ ( MNHUB), Jagor; 1♂ ( MNHUB), Limay, 27.VII.1914, leg. G. Boettcher; 1♂, 2♀♀ ( MNHUB), Milate, 26.VIII.1912; 1♂ ( SEHU), Los Banos, 5.VI.1982, leg. A. Tanaka; 1♂ ( MNHUB), same locality, 9.II.1914, leg. G. Boettcher; 1♂ ( MNHUB), same but with “ 24.II.1914 ”; 1♂ ( SEHU), same but with “ 4.VI.1983 ”; 2♀♀ ( SEHU), same but with “ 10.III.1984 ”; 5♂♂, 4♀♀ ( MNHUB), Polillo island, VIII.1915; 1♂ ( MNHUB), same locality, VIII.1915, leg. G. Boettcher; 1♀ ( NMNS), Quirino Prov., Maddela near Brgy Dismungal, 21.VII.2012, leg. M.-L. Jeng; Catandunes: 1♀ ( NMNS), Virax, San Vicente, 4.V.2013, leg. M. L. Jeng; Hasbate: 2♂♂ ( MNHUB), Aroroy, 21.VIII.1917; Mindanao: 2♀♀ ( SEHU), Mt. Apo, 27.VIII.1975, leg. Y. Komiya; 1♀ ( SEHU), Guianga, 16. VIII.1 975, leg. Y. Komiya; 1♀ ( SMNS), Cotabato Prov., Manobo Tasaday Forest Reserve, Mt. Tasaday, 3.II.–10.III.1991, leg. P. Lays; 13 exs. ( NHMB), 30km W of Maramag, 28– 30.XII.1990, leg. Bolm; 1 ex. ( NHMB), Portulin, 3.I.1991, leg. Bolm; 1♂ ( MNHUB), Surigao, 16.XI.1915; 1♂ ( MNHUB), same but with “ 4.VIII.1915 ”; THAILAND. Phangnga: 1♂ ( TCHU), Si Phangnga N. P., 21.IV.2000, leg. S. Tsuyuki.

Distribution. China (?), India, Indonesia (Bali, Java, Kalimantan, Sumatra), Laos, East Malaysia, Myanmar, Nepal (?), Philippines, Thailand ( Fig. 37 ). Records from China and Nepal need confirmation because no specimens were available for study.

Figures

FIGURE 34. Habitus of Oides femoralis Laboissière and O. flava (Olivier). 34A. O. femoralis, dorsal view; 34B. Same, ventral view; 34C. Same, lateral view; 34D. O. flava, dorsal view; 34E. Same, ventral view; 34F. Same, ventral view; 34G. Color variation, from Philipines, dorsal view; 34H. Same, ventral view; 34I. Same, lateral view.

FIGURE 36. Diagnostic characters of Oides flava (Olivier). 356. Antenna, male; 35B. Antenna, female; 36C. Penis, from India, dorsal view; 36D. Same, lateral view; 36E. Same, ventral view; 35F. Penis, from Borneo, dorsal; 35G. Same, lateral view; 36H. Penis, from Philippines, dorsal view; 36I. Same, lateral view; 36J. Apex of penis, from Philippine, lateral view; 36K. Apex of penis, from Philippines, lateral view; 36L.Abdominal ventrite VIII; 36M.Spermatheca.

FIGURE 37. Distribution map of Oides flava.