Notiothereva brunnipes (Kröber), Krober

Webb, Donald W., 2005, Revision of the Neotropical Stiletto Fly Genus Notiothereva Metz & Irwin (Diptera: Therevidae: Therevinae), Zootaxa 1059, pp. 1-32 : 25-28

Notiothereva brunnipes (Kröber)

( Figs. 30–36 , 44).

brunnipes Kröber (1911:500 key, 509 orig. desc.) ( Psilocephala ); Metz & Irwin (Metz et al. 2003:244 listing, comb. change, trans. orig. desc.) ( Notiothereva ). Type locality Azapa, Chile. Holotype female in SMTD.

Diagnosis. Like Notiothereva argentina and N. bezzii in having the frontal setae absent lateral to antennal base; parafacial lacking setae; and the gena with an anterodorsal patch of short, dark setae. Like N. bezzii in having the halter knob yellow; fore­ and midcoxa with macrosetae dark reddish brown; and the posterior margin of male sternite 8 ( Fig. 31 ) slightly emarginate. It differs from N. bezzii in having the forefemur lacking anterodorsal macrosetae; cervical sclerite with setae on lateral fourth; and the female frons pruinose.

organs. Scale = 0.1 mm.

Description of male.

Morphometric variation (n=10).

Body length 5.3–6.8, 6.2 mm.

Head. Length 0.70–0.94, 0.85 mm. Frons black, pruinescence gray, dense; setae black, elongate, absent lateral to antennal base. Antenna dark brown, pruinescence gray on scape and pedicel, first flagellomere paler; antenna/head length 0.66–0.79, 0.70; scape length 0.0.16–0.18, 0.17 mm, width 0.10–0.13, 0.11 mm, length/width 1.2–1.7, 1.6, scape/pedicel length 1.6–2.3, 2.0, scape/flagellum width 0.6–0.8, 0.7, setae dark reddish brown, short; pedicel length 0.08–0.10, 0.08 mm, length/width 0.7–1.0, 0.8, setae dark reddish brown; flagellum length 0.30–0.36, 0.34 mm, width 0.12–0.17, 0.15 mm, length/width 2.0–2.6, 2.3, flagellum/scape length 1.9–2.1, 2.0. Parafacial pruinescence whitish gray; setae absent. Maxillary palpus yellowish brown, pruinescence white; length 0.32–0.42, 0.37 mm, length/width 5.3–7.0, 6.1; setae white, elongate. Genal setae white, elongate with small, anterodorsal patch of short, dark brown setae. Occipital setae white, elongate, restricted along dorsolateral margin; macrosetae dark brown extending in row at end of postocular macrosetae.

Thorax. Macrosetae 3 np, 1–2 sa, 1 pa, 2 dc, 2 sc. Mesonotum dark brown, pruinescence brownish gray, vittae dull brown separated by lines of gray pruinescence; setae pale brown, erect, elongate intermixed with whitish brown, appressed setae. Postpronotal lobe setae whitish brown intermixed with a few dark brown setae. Cervical sclerite setae white on lateral fourth. Pleuron pruinescence gray, dense; setae white, elongate on propleuron, anepisternum, katepisternum, laterotergite, metanepisternum, and scutellum. Wing. Membrane opaque with small dark brown spot at r­m, fork of R4+5, apical margin of cells br, bm, and dc; venation pale brown; length 4.8–5.7, 5.4 mm, length/width 2.7–3.0, 2.8; pterostigma brown; cell m 3 open widely. Halter knob yellow. Legs. Coxa pruinescence white, dense; setae white, elongate; apical macrosetae dark reddish brown. Femora dark reddish brown, pruinescence white; ventral setae white on fore­ and midfemora; ad 0:0:0, av 0–1:0–2:3–5, pv 0:0–1:2–3. Tibiae dark yellowish brown, apex dark reddish brown. Tarsomere 1 dark yellowish brown, apex dark reddish brown; 2–5 dark reddish brown.

Abdomen. Cuticle dark reddish brown, pruinescence gray, dense; dorsal setae white, elongate, appressed intermixed with pale brown, elongate, appressed setae medially; lateral setae white, elongate. Terminalia (MEI 083725). Reddish brown, pruinescence white. Tergite 8 ( Fig. 30 ) setae dark reddish brown. Sternite 8 ( Fig. 31 ) anterior margin truncate, sides rounded; posterior margin slightly emarginate; setae brown across posterior margin. Epandrium ( Fig. 30 ) setae dark brown, scattered on posterior third with occasional thicker setae. Cercus ( Fig. 30 ) ending above apex of hypoproct. Hypoproct ( Fig. 30 ) posterior margin truncate, ending below apex of cercus. Gonocoxite ( Figs. 31–32 ) rounded laterally, posterior margin rounded, setae yellow, macrosetae dark reddish brown, apical setae of inner gonocoxal process dark yellow. Aedeagus with dorsal apodeme ( Fig. 33 ) anteromedial projection short, pointed; ventral apodeme ( Fig. 34 ) elongate, slightly clavate anteriorly, not extending beyond anterior margin of dorsal apodeme; distiphallus ( Fig. 33 ) broad basally, tapered posteriorly, basiphallus lacking median carina; lateral view ( Fig. 35 ) broadly curved ventrally; ejaculatory apodeme ( Fig. 33 ) anterior half slightly expanded, anterior margin rounded.

Redescription of female.

Morphometric variation (n=4).

Body length 5.7–7.2, 6.6 mm.

Head. Length 0.70–0.92, 0.83 mm. Frons pruinescence gray, dense with whitish gray pruinescence ventrolaterally and a band on brownish gray pruinescence ventrally across frons; setae black, short, absent lateral to antennal base. Antenna/head length 0.65–0.80, 0.73; scape length 0.14–0.19, 0.17 mm, width 0.12–0.15, 0.13 mm, length/width 1.2–1.3, 1.3, scape/pedicel length 1.8–2.3, 2.0, scape/flagellum width 0.67–0.88, 0.82, setae dark reddish brown, short; pedicel length 0.08–0.09, 0.08 mm, length/width 0.6–0.8, 0.7; flagellum length 0.32–0.38, 0.35 mm, width 0.14–0.18, 0.16 mm, length/width 2.0–2.3, 2.2, flagellum/scape 1.9–2.6, 2.1 times. Maxillary palpus length 0.34–0.40, 0.38 mm, length/width 3.8–6.7, 5.6.

Thorax. Macrosetae 3 np, 1–2 sa, 1 pa, 2 dc, 2 sc. Mesonotum dull brown with lines of gray pruinescence separating vittae; setae brown, erect, elongate intermixed with appressed, yellowish brown setae. Wing. Length 5.2–6.1, 5.7 mm, length/width 2.8–3.0, 2.9. Legs. Anterodorsal macrosetae 0:0:0, av 0:1–2:3–7, pv 0:0–2:2–4.

Abdomen. Cuticle dark brown, pruinescence whitish gray, tergites 1–6 with posterior margin orange brown, becoming broader on posterior tergites; dorsal setae dark yellow, appressed intermixed with appressed, dark reddish brown setae medially on dark reddish brown areas of tergites; lateral setae dark reddish brown intermixed with white, appressed setae. Terminalia (MEI 083741). Furca ( Fig. 36 ) quadrate, sides rounded; length 0.44 mm, width 0.34 mm; anterolateral projection short.

Immature stages. Unknown.

Distribution. Notiothereva brunnipes is known from Peru ( Fig. 44).

Habitats and Temporal Phenology. Notiothereva brunnipes has been collected in October and November and January through April.

Specimens Examined. Type specimen. The holotype female of Psilocephala brunnipes Kröber is labeled " Chile, Arica, 7. X. " (SMTD). Other specimens. CHILE. TARAPACA. Azapa [­18.52, ­70.18], 21.III.1967, Campos, L. E., 9 males MEI 0 28464 (USNM), 0 28465 (BMNH), 083728–31, 083733–4 (MEI), 083743(SMTD), 3 females 0 28466 (USNM), 0 28467 (BMNH), 0 83739 (MEI); Azapa­Grande, – XI.1964, collector?, 1 male MEI 0 83727 (MEI); – II.1965, collector?, 1 male MEI 0 83726, 1 female 0 83738 (INHS); – I.1966, collector?, 1 female MEI 0 83740 (MEI); – II.1966, collector?, 1 male MEI 0 83737 (MEI); – III.1966, collector?, 1 male MEI 83732 (MEI); – IV.1966, collector?, 1 male MEI 0 83725 (MEI); mouth of Lluta River [­18.4833, ­70.3333], 1.X.1966, MEI, 2 males MEI 083735–6 (MEI); Tierià Mar, – XI.1965, R. C., 1 female MEI 0 83741 (MEI).

Figures

FIGURES 30 – 43. Figures 30 – 36. Notiothereva brunnipes. 30. Tergite 8, epandrium, cerci, hypoproct, dorsal view. 31. Sternite 8, hypandrium, gonocoxite, ventral view. 32. Gonocoxite, hypandrium, dorsal view. 33. Aedeagus, dorsal view. 34. Aedeagus, ventral view. 35. Aedeagus, left lateral view. 36. Female internal reproductive organs. Figures 37 – 43. N. simulata. 37. Tergite 8, epandrium, cerci, hypoproct, dorsal view. 38. Sternite 8, hypandrium, gonocoxite, ventral view. 39. Gonocoxite, hypandrium, dorsal view. 40. Aedeagus, dorsal view. 41. Aedeagus, ventral view. 42. Aedeagus, left lateral view. 43. Female internal reproductive