Neuraphes (Pararaphes) tuyuhun, Jałoszyński, Paweł, 2015

Jałoszyński, Paweł, 2015, Three new species of Neuraphes in eastern Qinghai-Tibet Plateau (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 3972 (2), pp. 291-300 : 295

Neuraphes (Pararaphes) tuyuhun sp. n.

( Figs 2 , 8–10 , 17 )

Holotype: CHINA (Qinghai): ♂, two labels: " CHINA (Qinghai Prov.) / Daban Shan, 62 km NNW Honggu / 36º51'28''N / 102º37'07.6''E / 2240 m (creek valley, / Picea-Populus-Betula forest, / fresh downwashed loamy slope) / 26/ 29.VI.2011 D.W. Wrase [09D]" [white, printed], " NEURAPHES / ( PARARAPHES ) / tuyuhun m. / det. P. Jałoszyński '15 / HOLOTYPUS " [red, printed] ( PCMS). Paratypes (2 ♂♂): 1 ♂, same data as holotype, except 60 km NW Honggu, 36º49'10.7''N, 102º31'22.8''E, 2366–2400 m, dead wood and litter, 25.VI.2011, leg. M. Schülke (cPJ); 1 ♂, " CHINA: Qinghai Prov. [ CH 11-11a] / Lang Shi Dang Jing Qu (Park) 75 km / NW Honggu, 2925 m, 36º54'05.2''N, / 102º21'07.1''E, creek valley on over- / grown tree, Populus and Betula forest / with Rhododendron, litter sifted between / rocks, 28.VI.2011, leg. M. Schülke" [white, printed] ( PCMS). Paratypes with yellow PARATYPUS labels.

Diagnosis. Small species, 1.41–1.46 mm in length, bicolorous: head and pronotum dark brown, elytra and appendages lighter brown; head with barely noticeable pair of impressions; pronotum with median longitudinal carina of variable length, from about 1/4 to 1/2 of PL; each elytron with apical drop-shaped and elongate convexity indistinctly demarcated from surrounding surface and located closer to suture than to posterior elytral margin; ventral plate of aedeagus gradually broadening from base to apex.

Description. Body of male ( Fig. 2 ) strongly convex and elongate, with long appendages, BL 1.41–1.46 mm; head and pronotum dark brown, nearly black, elytra, legs except tarsi and antennae lighter brown, maxillary palps and tarsi yellowish; vestiture lighter than cuticle.

Head subtriangular, broadest at eyes, HL 0.21–0.23 mm, HW 0.26–0.28; tempora very short and strongly bent mesally; vertex convex; frons with posteromedian flattening bearing barely discernible traces of lateral pair of shallow impressions; supraantennal tubercles distinct; eyes large, strongly expanding laterally, coarsely faceted, in lateral view oval and vertically elongate. Punctures on vertex and frons fine and inconspicuous; setae sparse, long and erect. Antennae slender, AnL 0.68–0.69 mm, only antennomeres I, II and XI elongate, antennomeres III–VI about as long as broad and VII–IX distinctly transverse, antennomere XI slightly shorter than IX–X together, about 1.6 times as long as broad.

Pronotum in dorsal view subtrapezoidal, with lateral margins parallel from base to anterior third or indistinctly narrowing posteriorly from anterior third, PL 0.38 mm, PW 0.33–0.35 mm; anterior margin and sides in anterior third rounded, lateral margins in posterior half nearly straight, posterior corners slightly obtuse-angled and with acute tips, posterior margin arcuate; base with shallow transverse impression adjacent at each side to large subtriangular lateral impression and interrupted at middle by distinct median longitudinal carina equal to 1/4–1/2 of PL. Punctures on pronotal disc fine, sharply marked, very unevenly distributed, separated by spaces 0.5–5 times as wide as diameters of punctures. Setae on pronotal disc long, sparse and suberect.

Elytra suboval, broadest at or even slightly in front of anterior third, EL 0.83–0.88 mm, EW 0.58 mm, EI 1.43– 1.52; each elytron in anterior fourth with barely discernible adsutural flattening and apical drop-shaped protuberance which is distinctly elongate and located closer to elytral suture than to posterior elytral margin, protuberance has diffused margins and is indistinctly demarcated from surrounding cuticle; basal elytral foveae located much closer to scutellum than to humeri; humeral calli moderately distinct, developed as elongate protuberances. Punctures on elytra larger and denser than those on pronotum, but shallow and with diffused margins, separated by spaces subequal to or (especially in anterior half) slightly smaller than diameters of punctures; setae short, sparse and suberect. Hind wings well-developed, twice as long as elytra.

Legs long and slender; unmodified.

Aedeagus ( Figs 8–10 ) relatively stout, AeL 0.20 mm, in ventral view gradually broadening from base to subapical region; ventral plate narrow, with sharply demarcated lateral and apical margin and base indistinguishably fused with surrounding membranes, sides gradually narrowing from base to apical region; lateral plates in lateral view with subapical constriction demarcating round apical 'head', in ventral view apical parts of lateral plates bent laterally; parameres with subtriangular apices.

Female. Unknown.

Distribution. China, Qinghai ( Fig. 1 k).

Etymology. Tuyuhun is a name of a historical kingdom in the area where Neuraphes tuyuhun occurs.

Remarks. See remarks for N. sichuanus .

Figures

FIGURE 17. Distribution of nominal species of Neuraphes in mountain ranges that surround Qinghai-Tibet Plateau. Neuraphes aruensis Franz (a), N. jumlanus Franz (b), N. taksangensis Franz (c), N. himalayanus Franz (d), N. khumbuanus Jałoszyński (e), N. mephistopheles Jałoszyński (f), N. hengduanus Jałoszyński (g), N. tibetanus Jałoszyński (h), N. pseudojumlanus Jałoszyński (i), N. sichuanus sp. n. (j), N. tuyuhun sp. n. (k), N. miniqinghaiensis sp. n. (l), N. cf. qinghaiensis (m), N. qinghaiensis Jałoszyński (n).

FIGURES 1 – 4. Neuraphes, habitus of males: N. sichuanus sp. n. (1), N. tuyuhun sp. n. (2), N. miniqinghaiensis sp. n. (3), and N. cf. qinghaiensis (4).

FIGURES 5 – 10. Aedeagi in dorsal (5, 8), ventral (6, 9) and lateral (7, 10) views. Neuraphes sichuanus sp. n. (5 – 7), N. tuyuhun sp. n. (8 – 10).