Neoperla andreas, Sivec & Stark, 2011

Sivec, Ignac & Stark, Bill P., 2011, New Species Of Neoperla Needham And Phanoperla Banks (Plecoptera: Perlidae) From The Philippine Islands, Illiesia 7 (24), pp. 264-279 : 266-267

Neoperla andreas sp. n.

( Figs. 9-11 )

Material examined. Holotype ♂ and 1♂ paratypes, PHILIPPINE ISLANDS, Palawan, Salakot Falls Road, 300 m, 19 March 1996. Additional paratypes: Palawan, Estrella Falls , 3 March 1995, 1♂ (all specimens deposited in the PZ collection).

Adult habitus. General color pale brown patterned with yellow. Head pale brown over most of frons, but with a dark spot forward of M-line and a pair of brown spots forward of callosities ( Fig. 9 ). Pronotum with narrow median and sublateral dark bands. Legs, antennae and palpi pale; wings pale brown with darker venation. Ocelli large, about 1 diameter apart. Male. Forewing length 13 mm. Tergum 7 bearing a small posteromedian triangular lobe and a median sclerotized area ( Fig. 10 ). Tergum 8 with a small, median, recurved, tongue-shaped process. Tergum 9 with a small median patch of sensilla. Hemitergal lobes slender, conspicuously narrowed to apex, scarcely bent and directed mesad. Penis tube weakly sclerotized along ventral margin, bare, and subequal to sac in length ( Fig. 11 ). Sac curved dorsad near apex and armed over much of surface with variably sized spines, largest spines form a moderately wide penultimate band; subapex of sac bearing a narrow ring of fine spines; venter of sac bearing near midlength, a large, basally directed, weakly bilobed process, mostly bare but weakly armed on tips; dorsolateral of ventral lobe, a pair of spherical, spiny lobes are located.

Female. Unknown.

Larva. Unknown.

Etymology. The species name honors Dr. Andreas Zwick, who graciously forwarded this material for our study.

Diagnosis. Neoperla andreas is a member of the N. recta Banks, 1913 complex of the N. montivaga species group ( Sivec 1984; Zwick 1986). This complex currently includes four Philippine species, N. nishidai Sivec, 1984 , N. pseudorecta Sivec, 1984 , N. recta and N. zwicki Sivec, 1984 . The new species is the only one of this group in which the ventral penial lobe is bilobed and at least weakly armed.

Figures

Figs. 9-11. Neoperla andreas. 9. Head and pronotum. 10. Male terminalia. 11. Penis tube and everted sac, lateral aspect, and ventral aspect inset.