Eunatalis hirta (Blackburn, 1890)

Gerstmeier, Roland & Seitner, Martin, 2013, Revision of the checkered beetle genus Eunatalis Schenkling, 1909 (Coleoptera: Cleridae: Clerinae), Zootaxa 3698 (1), pp. 1-77 : 20-24

Eunatalis hirta (Blackburn, 1890)

( Figures 21–22 , 55 F)

Natalis hirta Blackburn, 1890: 122 ; Blackburn 1899: 30.

Eunatalis longicollis (Blackburn, 1890) : 126; Blackburn 1899: 29 n. syn.

Specimens examined: Natalis hirta : Lectotype (designated here): Type; Australia, Blackburn Coll., B.M. 1910- 236; 3499T; Natalis hirta (♂, BMNH). Paralectotype (designated here): Israelite Bay; Natalis hirta Blackb , cotype; Natalis hirta, Blkb ,?W Australia; 25-035778 (SAMA).

Natalis longicollis : Lectotype (designated here): Type; 3445 Ad; Australia, Blackburn Coll., B.M. 1910-236; Natalis longicollis Blackb. (♂, BMNH). Paralectotype (designated here): Natalis longicollis , S. Australia, 9588 Cotype; S. Australia; 25-035800 (SAMA).

Comment on the type specimens: As Blackburn (1890) made no reference to type specimens in his descriptions of Natalis hirta and N. longicollis , all 'type' material for these nominal species-group taxa should be treated as syntypes (ICZN 1999: Article 73.2) and, as such, are available for lectotype designation (ICZN 1999: Article 74.1). Following Recommendation 74D (ICZN 1999) we have designated the BMNH syntypes as lectotypes because the majority of Blackburn's types are held in that institution (Lea 1912). Syntypes borrowed from SAMA were designated as paralectotypes.

Others: W. Australia; Natalis hirta Blackb, S. Schenkling det; Homotype, Eunatalis hirta Blackburn (BMNH). Ex Musaeo H.W. Bates 1892 (MNHN). East Hyden, 10-2-73 WA (ANIC). SEQ: 24°32'Sx 150°36'E, Mt. Scoria 200m, 29 Jun.–21 Oct. 2000, Cook, Wright, Vanderduys, vine scrub, FIT 9469 (QM). Qld: 22°28.7'Sx 147°52.3'E, Mt. Pollux, SW base, site 1, 12 Jan.–5 Mar. 2006, intercept, C. Burwell, 400m, brigalow, 13307 (QM). Eucla Basin, S.A. Eucla, 1600.2, nr the coast, 11.x.66, J. Lowry 42/40 (ANIC). Mullamullang area, Eucla Basin, S.A., surface, 29.viii.66 – 2.ix.66, J. Lowry, 42/33 (ANIC). W. Australia, Drummond Cove, 7m, N. of Geraldton, on coast, 3.v.1973; on sand dunes; N. McFarland Coll., B.M. 1973-399; DC, 3May73, uv.-NMcF. (BMNH). 25.23S, 135.25E, 13 km E. by N. of Andado HS., NT, 27.ix.72, J. Upton (ANIC). Greenough, (28.57S, 114.44E), Western Australia, 26–29 Aug. 1979, R.P. McMillan; Dune system; Reg.no. 83283 (WAMP). Australia; Geraldton; 25- 0 35795 (SAMA). C.QLD: 22°02'Sx 148°03'E, Moranbah, 6 km S 5646, 25 Jun.–20 Dec. 1997, G. Monteith & E. Kruck, Fit Intercept, Box flat, 5 ex. (QM). CQ: 22°03'Sx 148°04'E, Moranbah, 6 km S, 220m, 20 Dec. 1997 – 26 Apr. 1998, G. Monteith 5803, Flight Intercept, BoxFlat, 2 ex. (QM). SEQ: 24°38'Sx 149°01'E, Expedition Ra, Dawson Hwy, 27 Oct.–17 Dec. 1979, 420m, Cook & Monteith, FIT, Bottle Tree scrub 9821 Dalby Q, Mrs. F.H. Hobler; 25-035790 (SAMA). Dalby, N.W.A., W.D. Dodd; 25-035841 (SAMA). Boeyana, W.A., Miss Baisson; Col-65724 (NMV). QLD: 26°03'Sx 150°55'E, "Wonga-Hills", 10 km ENE, 11 Dec. 2001 – 4 Mar. 2002, Monteith & Cook, 320m, FIT, brigalow, 10405, 2 ex. (QM). Nyngan, NSW, 13-3-63 CW; 25-036039 (SAMA). QLD: 26°38'Sx 144°46'E, Cheeple, 26 km W, 4 May 2001, G.B. Monteith, 10 147, Pyrethrum, gidgee (QM). Albany; Natalis alternicostata Kuw. (MNHN). Cap York (NMPC). SEQ: 25°35'Sx 149°46'E, Taroon, 9 km N, Ooline Scrub, roadside BS 15, 15 June 1996, P. Lawless, 174 (QM). Helena & Aurora Ranges, 30°21.22'S 119°42.20'E, W. Aust. Site#1A, 26 Sept 1995, R.P. McMillan CALM Survey (mallee woodland), Reg.no. 83284 (WAMP). Mt. Gibson Stn, WA, 12 February 2000, S.R. Patterson, Reg.no. 83300 (WAMP). W.- Australia 9.11.87, 3 km s Kambalda West, leg. Gerstmeier/Baehr (RGCM). NSW, 60k N of Moree Careunga Nat. Reserve, 28 March 1990, leg. P.H. Colman, Hangay collection; K 324493 (AM). W.- Australia, Young River, 80 km w Esperance, 11./ 12.11.1987, leg. Gerstmeier/Baehr (RGCM). Etadunna Stn, S.A., K&B Oldfield, x.1972; Reg.no. 83278 (WAMP). Higgins, Winton, 1912 (QM).Clermont, Queensland, Dr.K.K. Spence; K 324343 (AM). Clermont, Queensland, 8, 28, Dr.K.K. Spence; K 324344 (AM). Minnie Downs, N.(?)Corner of S. Aust., L. Reese; 25-035998 (SAMA). Western Australia, Wilga, 18-12-1974, K.&E. Carnaby (ANIC). Adelaide, Blackburn; 25-035801 (SAMA). Ouldea, S.A., R.T. Maurice; 25-035781 (SAMA).Adelaide; Ex Musaeo, H.W. Bates, 1892 (MNHN).21°10'16S 148°29'56E, Eungella Nat. Pk., Broken River QLD, 31 Aug.–8 Sep. 1998, Neave, Edwards, Powell, Sutrisno & Hebbard (ANIC). QLD: 26°01'Sx 148°16'E, Mt. Hutton, 8,5 km SW, 15 Dec. 2001 – 6 Mar. 2002, Monteith & Cook, 550m, FIT, vine scrub, 10415 (QM).NSW, 4 km SE Mt. Harris, 7–8.ix.1982, B. Holloway; K 324482 (AM). Cunnamulla Dis., Q, Nov 32, N. Geary; K 324395 (AM). Cunnamulla, Q 12. Nov. 38, N. Geary; K 324374 + K 324375 (10.2.39) (AM). Cunnamulla, H. Hardcastle; 25-035787 (SAMA). Cunnamulla, Q., Oct 1941, N. Geary; K 324394 (AM).Lake Narran, N.S.W., 12.ix.1961, A. Holmes; K 324427 (AM). Swan R., J.S. Clark; A.H. Elston Collection; (BMNH), K 324386 (AM). Swan R, Lea; Bridgetown, W.A. Lea; hirta Blackb. ; Natalis hirta Blackb, W. Australia; 25-035789 (SAMA). Victoria, Barton (MNHN). Bernsborough Downs, via Winton Qld., 23 September 1973. attracted to light, R. Patterson F248; K 324467 (AM). 33.51S 123.00E, Thomas River, 23 km NW by W of Mt. Arid WA, 4–7.xi.1977, J.F. Lawrence (ANIC). SEQ: 25°41'Sx 151°25'E, Nipping Gully, site 3, 26 Jan.–2 Jun. 1999, 240m, Monteith & Thompson, open for, pitfall, 2707 (QM). Airlee Island, 21.20S 115.10E, Western Australia, 20July 1987, V. Long, AE 0 0 4, Reg.no. 83286 (WAMP). At light, Goolgow,N.S.W., 6-XI-89 (QM). QLD: 25°46'Sx 148°17'E, "Oak Wells", site 3, 15 Dec. 2001 – 7 Mar. 2002, Monteith & Cook, 600m, FIT, vine scrub, 10419 (QM). QLD: 25°46'Sx 148°17'E, "Oak Wells", site 3, 16 Dec. 2001, 10286, Monteith, Cook, Blosh & Wright, vine scrub (QM). W. Australien, Marloo Station, Wururga II 1935, A. Goerling S.G., 2 ex. (ZMB). Marloo Stn., Wururga, W.A., 1931–1941, A. Goerling, 6 ex. (ANIC). Alice Springs NT, 3 September 1992, light, R. Patterson, 2810 + 2799, 3 ex. (ANIC). SA: 18 km S Port Pirie, 7.v.1980, G.J.&A. Holloway; K 324473, K 324474, 2 ex. (AM).38-2324, Hopetoun; Reg.no. 83259 (WAMP). 34.03S 140.43E GPS, Calperum HS, S.A., 13–14 Dec. 1995, at light, K.R. Pullen; Calperum Station/Bookmark Biosphere Reserve, Invertebrate Survey (ANIC). S.Aust.

1 km N White Bull, Yard Kalamurina Stn, 27°52'23"S 137°54'40"E, 5 Oct. 1999, at light, Warburton R. WHC exped.; 25-035911 (SAMA). Chowilla SA, 4/68 C. Watts; 25-036104 (SAMA). W.- Australia, 23.11.1987, Boyup Brook, Stadtpark, leg. Gerstmeier/Baehr (RGCM). W.- Australia, 12.11.1987, Oldfield River, 20 km w Munglinup, leg. Gerstmeier/Baehr (RGCM). Milmerran, SA, 3/51; J.G. Brooks, Bequest, 1976 (ANIC). W.A. Beverley; K 324353 (AM). W.A. Beverley; K 39149 ; N. porcata Fab., Id. by A.M. Lea; K 324353 (AM). 200 km N of Tennant, Creek township, Stuart Hwy, N.T., 4 April 1984, at light, K.&E. Carnaby (ANIC). S. Aust., Munyaroo CP, 12.9 km W Moonabbie, 33°23'14"S 137°15'25"E, 28th September 2002, under logs, SEG Munyaroo Survey; 25-035918 (SAMA). WA: Lake Moore, south BE11-SAP, wet pitfalls, 30°20'09" 117°29'48"E (GPS), 15 Sept 1998 - 25 Oct 1999, N.A. Guthrie, CALM Survey, Reg.no. 83309 (WAMP).S. Aust., Billiatt CP, 32 km S Alawoona, under bark, 34°59'29"S 140°28'22"E, 27 Mar 2000, JA Forrest; 25-035904 (SAMA). Natalis cribricollis Spin., Gayndah ; Pl. 34050; K 324351 (AM). 18mi. ESE of Penong, S.A., 2 Oct. 1968, Key, Upton, Balderson (ANIC). W.Aust., Faure Is., Shark Bay, 15 km SE Monkey Mia, 25°53'05"S 113°52'51"E, 27–31 May, 2000, pitfalls, JA Forrest, Triodia over sand, FAU 003; 25-035910 (SAMA). Burley Waterhole, 5mls S of Innaminka Station, Sth. Australia, II– 13.iii.61, R. Mackay; K 324426 (AM).Bogan R., N.S. Wales, J. Armstrong (ANIC). Bogan R., N.S. Wales (ANIC).Gawler (NMPC). Gawler (SDEI). Milly Soak, N of Cue, West. Aust., Em. Jan. 2007, P. Hutchinson; Reg.no. 83304 (WAMP).Dunmore, via Dalby Qld, 10.xi.1980, F.R. Wylie, M.DeBaar, Acc 1819/36; O-166586 + O-166587, 2 ex. (QDPC). Reid, ACT., 15 Apr 2007, B. Boyd (ANIC). 29.00S 151.05E, 18 km SSW Texas QLD, 24.11.83, D. Rentz, M. Harvey, Stop 66 (ANIC). Picton; Griffith Collection, Id. by A.M. Lea; N.porcata VAR.; 25- 0 35869 (SAMA).Redcliffs V., L.G. Chandler; K 324382 (AM). n. Jamvold, 25.V.1963, Mundarling, W. Australia; Reg.no. 83275 (WAMP). R.P. McMillan, Rottenest, on Acacia, Sept. 1939 ; Reg.no. 83273 (WAMP). 47–75, Perth; Reg.no. 83272 (WAMP).35-983, W.-Midland; Reg.no. 83258 (WAMP). Kenwick, W.A., leg. Demarz, 1964 (NHMB). S. Aust., nr Clifton Hills OS, 26°30'S 139°29'E, malaise trap, 18–20 Nov. 1993, JA Forrest, D Hirst; 25- 0 35907 (SAMA). 25.46S 133.18E, 9 km N of Kulgera, N.T., 1.x.72, J. Upton (ANIC). Qld: 25°14.5'Sx 148°37.3'E, Moolayember Creek NP, site 3, 14303, 27 Jun.–6 Sept. 2006, 410m, R. Raven, B. Baehr, A. Amey (QM). Milly Milly, 10-6-22; 1922-435; Reg.no. 83268 (WAMP). Australia, W.A., Kalamunda, I-16-1954, H. Demarz leg., 2 ex. (FMNH). Mallee District, Victoria (SDEI). Cumberland, N.S.W. (MNHN). Braemar S.F., via Kogan, S.E. Qld., 15–19 Oct 1979, G.B. Monteith & QM (QM). S. Aust., Kalumurrina, Water Hole, At light, 25 Nov. 1964, P. Aitken; 25-035919 (SAMA). Murray R., S.A., H.S. Cope; K 324340 (AM). 12.19S 133.19E, Nabarlek N.T., 19 Nov 1983, Melanie Webb (ANIC). Sea Lake, V., 5.1916, Goudie; Collection J.C. Goudie (NMV). Juranda Rockhole, 60mi S of Balladonia, W.A., 33.13S 123.27E, 21.xi.1969 Mallee at light, E. Britton (ANIC). (21.35S 117.04E), Millstream, WA, Deep Reach, 8.xi.70, at light, E.B. Britton (ANIC).Mullewa WA (SDEI). Cannuwaukaninna Dune, 16 km W of Etadunna Homestead, S. Aust., 5 Mar 1972, E.G. Matthews; 25-035914 (SAMA). Australien, 12/1969, Iron Baron, M. Buzga lgt. (JRCP).W. Australia, Coolgarlie, Queen Victoria Rock, 19.10.2001, Bily lgt. (JRCP). W. Australia, Peak Charles N.P., 8.xi.2001, Sv. Bily lgt. (JRCP). Austr., NT, 70 km SW of Mataranka, 22.12.0 8, 15°19'S 132°50'E, 190m, Sv. Bily leg. (JRCP).Austr., NT, 40 km W of Katherine, 28.11.0 8, 13°15'S, 130°44'E, 63m, Sv. Bily lgt. (JRCP).

Diagnosis: As indicated by Blackburn (1890), the 2nd, 4th and 6th elytral interstices are charactistically ridged in E. hirta ; as is the 8th interstice being conspicuously ridged behind the middle. The phallic tip has an open Z-shaped structure and less than one-fifth of the spicular fork is fused.

Length: (171 specimens measured) Total length: 10–29 mm.

Head: Dark brown to black, with dense pilosity; densely punctated with small punctures and isolated large punctures; epistomal suture more or less conspicuous; interocular space about 1.5 to double eye width.

Thorax: Pronotal length to width ratio 1.12:1; pronotum light brown to dark brown, with dense pilosity, with dense and small punctation and isolated large punctation; inconspicuously wrinkled; widest anterior to middle, slightly convergent towards apex; grooves of sub-basal collar separated by a conspicuous, wide bar. Scutellum light brown to dark brown, with weak punctation and isolated pilosity.

Elytra: Elytral length to width ratio 2.64:1; elytra light brown to dark brown, with dense greyish pilosity; elytral base slightly broader than pronotum, broader behind middle; punctations in basal half as wide as interstices, becoming only slightly smaller behind middle; elytral punctation with three nodules; punctations of tenth stria distinct just before apex; punctations of ninth stria vestigial; interstices 2 slightly, interstices 4 and 6 just shortly behind middle and interstices 8 behind middle conspicuously ridged.

Lower surface: Light brown to dark brown, with dense pilosity; procoxal cavities slightly opened to closed posteriorly; mesoventrite and metaventrite with dense punctation; posterior part of mesepisternum smooth.

Legs: Light brown to dark brown, with dense pilosity and dense punctation; mesotibiae conspicuously bent.

Abdomen: Tegmen closed ventrally; phallus tip with an open Z-shaped structure; spicular fork fused less than one-fifth.

Distribution: Australia.

Seasonal occurrence: Collected the whole year.

Remarks: E. longicollis was synonymized with E. hirta because there were no differences in the aedeagi, pygidia and 6th ventrites. Within the species there is a conspicuous variability regarding the pronotum, distinctness of the elytral punctation in the apical half, and of the pilosity. As Blackburn described in 1890, the ridged interstices 2, 4 and 6 are particular.

Figures

FIGURE 55. (A) Eunatalis auricoma n. sp., (B) Eunatalis debilis, (C) Eunatalis fasciata, (D) Eunatalis flavopilosa n. sp., (E) Eunatalis foveata n. sp., (F) Eunatalis hirta.

FIGURE 21. E. hirta: (A) Phallus lateral, (B) Phallus ventral, (C) Tegmen ventral, (D) Spicular fork, (E) Pygidium, (F) Ventrite VI.

FIGURE 22. Distribution of Eunatalis hirta.